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154 lines
16 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.77.936" ID-GBIF-Dataset="41b91a41-397c-4bf0-a178-2aff5bb42c34" ID-PMC="PMC3088053" ID-Pensoft-Pub="1313-2970-77-31" ID-PubMed="21594150" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-77-31" ModsDocOrigin="ZooKeys 77" ModsDocTitle="Review of the European Greenomyia Brunetti (Diptera, Mycetophilidae) with new descriptions of females" checkinTime="1451250825072" checkinUser="pensoft" docAuthor="Kurina, Olavi, Hedmark, Kjell, Karstroem, Mats & Kjaerandsen, Jostein" docDate="2011" docId="FAC2405A80F1D6197F71874FA372AB22" docLanguage="en" docName="ZooKeys 77: 31-50" docOrigin="ZooKeys 77" docSource="http://dx.doi.org/10.3897/zookeys.77.936" docTitle="Greenomyia mongolica Lastovka et Matile 1974" docType="treatment" docVersion="4" lastPageNumber="42" masterDocId="D37EFFC2FFEAFFB1FFA2FFC2FFA1FFC6" masterDocTitle="Review of the European Greenomyia Brunetti (Diptera, Mycetophilidae) with new descriptions of females" masterLastPageNumber="50" masterPageNumber="31" pageNumber="41" updateTime="1668166018994" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the European Greenomyia Brunetti (Diptera, Mycetophilidae) with new descriptions of females</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kurina, Olavi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hedmark, Kjell</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Karstroem, Mats</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kjaerandsen, Jostein</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2011</mods:date>
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<mods:detail type="volume">
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<mods:number>77</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>31</mods:start>
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<mods:end>50</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.77.936</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.77.936</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-77-31</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="159361149" LSID="urn:lsid:plazi:treatment:FAC2405A80F1D6197F71874FA372AB22" httpUri="http://treatment.plazi.org/id/FAC2405A80F1D6197F71874FA372AB22" lastPageId="11" lastPageNumber="42" pageId="10" pageNumber="41">
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<subSubSection pageId="10" pageNumber="41" type="nomenclature">
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<paragraph pageId="10" pageNumber="41">
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<taxonomicName authority="Lastovka et Matile, 1974" authorityName="Lastovka et Matile" authorityYear="1974" class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia mongolica" order="Diptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="mongolica">
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<pageBreakToken pageId="10" pageNumber="41" start="start">Greenomyia</pageBreakToken>
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mongolica
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<normalizedToken originalValue="Laštovka">Lastovka</normalizedToken>
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et Matile, 1974
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</taxonomicName>
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Figures 371115192327
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="41" type="reference_group">
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<paragraph pageId="10" pageNumber="41">
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia theresae" order="Diptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="theresae">Greenomyia theresae</taxonomicName>
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Matile, 2002, syn. n.
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</paragraph>
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<paragraph pageId="10" pageNumber="41">
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia theresae" order="Diptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="theresae">Greenomyia theresae</taxonomicName>
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<bibRefCitation author="Kurina, O" editor="Ziegler J (Ed) Diptera, Stelviana" journalOrPublisher="Supplements" pageId="15" pageNumber="46" pagination="245 - 293" title="Sciaroidea excl. Sciaridae." volume="16" volumeTitle="A dipterological perspective on a changing alpine landscape. Volume 1. Studia Dipterologica." year="2008 a">Kurina 2008a</bibRefCitation>
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: 255, 270.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="41" type="type material studied">
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<paragraph pageId="10" pageNumber="41">Type material studied:</paragraph>
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<paragraph pageId="10" pageNumber="41">
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Paratype ♂ of
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia mongolica" order="Diptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="mongolica">Greenomyia mongolica</taxonomicName>
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: MONGOLIA. Central aimak, Tosgoni ovoo, 5-10 km N von Ulan-Baator, 1500-1700 m a.s.l., Exp. Dr. Z. Kaszab 1967 nr. 926, 19-24 Jul 1967 (Z. Kaszab leg.) [MNHN, JKJ-SPM-011843, on pin]
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</paragraph>
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<paragraph pageId="10" pageNumber="41">
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Holotype ♂ of
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia theresae" order="Diptera" pageId="10" pageNumber="41" phylum="Arthropoda" rank="species" species="theresae">Greenomyia theresae</taxonomicName>
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: ITALY. Aosta, Champlong, Dessus, 1000 m a.s.l., "courant sur Feuille de
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<normalizedToken originalValue="Frêne">Frene</normalizedToken>
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- 2m -", 26 Aug 1997 (L. Matile leg.) [MNHN, JKJ-SPM-011844, on pin].
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</paragraph>
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<paragraph pageId="10" pageNumber="41">
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Other material studied: SWEDEN. 1♂,
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<normalizedToken originalValue="SÖ">SOe</normalizedToken>
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, Stockholm,
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<normalizedToken originalValue="Skarpnäck">Skarpnaeck</normalizedToken>
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, Skarpa by, 13.VII.-4.X.2003 (B. Viklund leg.) [MZLU, in alcohol]. GERMANY. 1♂, D. Oberpfals, NM Main-Donau-Kanal (Proj. Warncke), 12.IX.-5.X.1988 (S. Blank leg.) [No. 30132 in ZSM, in alcohol]. RUSSIA. 2♂♂ 3♀♀, Nikolsk-Ussur, 29.VII.1926 (Kuznetzov leg.) [ZIN, on pins]. ESTONIA. 1♀,
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<normalizedToken originalValue="Kääriku">Kaeaeriku</normalizedToken>
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, 5.X.1985 (H. Remm leg.) [IZBE, on pin]; 1♀, Luunja, 20.X.1996, on the house wall (O. Kurina leg.) [IZBE, on pin]; 6♂♂ 3♀♀, Karilatsi near Tartu, bait traps, 19.-28.VIII.2005 and 04.-25.IX.2005 (T. Tammaru leg.) [IZBE, on pins]. HUNGARY. 38♂♂ 3♀♀, 10 km S Eger, 47°49'11"N, 020°21'37"E, 20 Aug 1989 (R. Danielsson leg.) [MZLU, on pins]. ITALY. 1♂, Aosta valley, Verrayes, Promellian, 1200 m.a.s.l., sweeping, 17.VI.2007 (V. Soon leg.) [IZBE, on pin]; 1♀, Siena, 6.V.2007 (A. Selin leg.) [Coll. Selin, on pin]; 3♂♂, Trentino-Alto Adige, Prov. Bolzano, Parco Nationale dello Stelvia, Sulden Valley near Schmelz southwest of Prad, 46°36'42.1"N, 010°34'35.6"E, 940 m.a.s.l., 5.IX.-14.X.2005 (J. Ziegler and C. Lange leg.) [1♂ in ZMHB, 2♂♂ in IZBE, mounted from alcohol]. GREECE. 1♂ 2♀♀, Central Macedonia, Kerkini lakes area, village Vironia, Beabies site, 41°19'15.4"N, 023°13'39.6"E, 1150 m.a.s.l., Malaise trap, 30.VI.-6.VII.2008 (G. Ramel leg.) [IZBE, mounted from alcohol]; 6♀♀, Central Macedonia, Kerkini lakes area, village Vironia, Ramna site, 41°17'42.5"N, 023°11'33.1"E, 750 m.a.s.l., Malaise trap, 23.-29.VI.2008 (G. Ramel leg.) [IZBE, mounted from alcohol].
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="42" pageId="10" pageNumber="41" type="diagnostic characters">
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<paragraph pageId="10" pageNumber="41">Diagnostic characters.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="42" pageId="10" pageNumber="41">
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Female. Thorax dark brown to blackish. Abdomen entirely blackish brown or first three segments slightly paler laterally. cx1 entirely yellow or darkened in basal half, cx2 and cx3 entirely dark brown to black. Fore trochanter yellow basally, brown apically. Mid and hind trochanters brown. f1 and f2 yellow, f3 yellow with brown apical fifth. Tibiae yellow, apically slightly darkened, with dense brown setae. Scape, pedicel and flagellomeres brown. Mouthparts pale yellow. Apical palpal segment 1.8-2.2 (n=5) times as long as penultimate segment. Wing tip shaded on about apical third, with darkened area along fore margin. All veins reach wing margin, M2 sometimes basally obsolete or very weak, A1 ending close to, sometimes fused into base of CuA2. Terminalia brown. Cercus distinctly two-segmented, apical segment small, ovate. Gonapophysis IX membranous, widely protruding apically, not visible
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<pageBreakToken pageId="11" pageNumber="42" start="start">in</pageBreakToken>
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lateral view. Tergite VIII larger than tergite IX. Sternite VIII apically with shallow medial incision, moderately emarginated basally.
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</paragraph>
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<paragraph pageId="11" pageNumber="42">Male. Coloration and other non-terminal characters including palpi similar to female. Apical palpal segment is 1.7-2.1 (n=5) times as long as penultimate segment.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="42" type="remarks">
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<paragraph pageId="11" pageNumber="42">Remarks.</paragraph>
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<paragraph pageId="11" pageNumber="42">
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This species was originally described by
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<bibRefCitation author="Lastovka, P" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="15" pageNumber="46" pagination="93 - 135" title="Mycetophilidae (Diptera) de Mongolie" volume="20" year="1974">
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<normalizedToken originalValue="Laštovka">Lastovka</normalizedToken>
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and Matile (1974)
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</bibRefCitation>
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based on Mongolian material and subsequently widely recorded in Europe.
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<bibRefCitation author="Chandler, PJ" editor="de Jong, H" journalOrPublisher="Schweizerische Entomologische Gesellschaft" pageId="14" pageNumber="45" title="Fauna Europaea: Diptera, Nematocera. Fauna Europaea, version 1.2" url="http://www.faunaeur.org" year="2005">Chandler (2005)</bibRefCitation>
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did not include
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia mongolica" order="Diptera" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="mongolica">Greenomyia mongolica</taxonomicName>
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in the European list and assigned all records to
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia theresae" order="Diptera" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="theresae">Greenomyia theresae</taxonomicName>
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, a species described from northern Italy by
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<bibRefCitation author="Matile, L" journalOrPublisher="Annales de la Societe Entomologique de France (Nouvelle serie)" pageId="15" pageNumber="46" pagination="13 - 16" title="Notes sur le genre Greenomyia Brunetti et description d'une espece nouvelle des Alpes et des Apennins (Diptera: Mycetophilidae)." volume="38" year="2002">Matile (2002)</bibRefCitation>
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. Careful comparisons of type material of both species at MNHN in Paris (independently undertaken by two of the authors, OK and JK; the holotype of
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia mongolica" order="Diptera" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="mongolica">Greenomyia mongolica</taxonomicName>
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deposited in the Hungarian Natural History Museum was not available for the study) did not indicate any substantial differences in their male terminalia. The minor diagnostic characters as indicated in the original description and illustrations by
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<bibRefCitation author="Matile, L" journalOrPublisher="Annales de la Societe Entomologique de France (Nouvelle serie)" pageId="15" pageNumber="46" pagination="13 - 16" title="Notes sur le genre Greenomyia Brunetti et description d'une espece nouvelle des Alpes et des Apennins (Diptera: Mycetophilidae)." volume="38" year="2002">Matile (2002)</bibRefCitation>
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are liable to different angles of views only. Consequently we have come to the conclusion that
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia theresae" order="Diptera" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="theresae">Greenomyia theresae</taxonomicName>
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at present state of knowledge must be treated as a junior synonym of
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<taxonomicName class="Insecta" family="Mycetophilidae" genus="Greenomyia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Greenomyia mongolica" order="Diptera" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="mongolica">Greenomyia mongolica</taxonomicName>
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and that all published records in Europe should rather be associated with the latter. In addition to the studied type material, we also compared the terminalia of female specimens from the Russian Far East, Estonia and northern Greece without finding any reliable differences. Moreover,
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<bibRefCitation author="Papp, L" journalOrPublisher="Folia Entomologica Hungarica" pageId="15" pageNumber="46" pagination="219 - 231" title="Pediciidae, Bolitophilidae, Keroplatidae, Mycetophilidae and Dixidae: genera and species new to Hungary (Diptera)." volume="61" year="2000">Papp (2000)</bibRefCitation>
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confirmed conspecificity when he compared central European material from Hungary with the Mongolian type material. Male terminalia are figured by
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<bibRefCitation author="Lastovka, P" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="15" pageNumber="46" pagination="93 - 135" title="Mycetophilidae (Diptera) de Mongolie" volume="20" year="1974">
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<normalizedToken originalValue="Laštovka">Lastovka</normalizedToken>
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and Matile (1974)
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</bibRefCitation>
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and subsequently by
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<bibRefCitation author="Matile, L" journalOrPublisher="Annales de la Societe Entomologique de France (Nouvelle serie)" pageId="15" pageNumber="46" pagination="13 - 16" title="Notes sur le genre Greenomyia Brunetti et description d'une espece nouvelle des Alpes et des Apennins (Diptera: Mycetophilidae)." volume="38" year="2002">Matile (2002)</bibRefCitation>
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, while female terminalia have previously been figured by
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<bibRefCitation author="Zaitzev, AI" journalOrPublisher="Vestnik Zoologii" pageId="16" pageNumber="47" pagination="25 - 32" title="Greenomyia and Neoclastobasis. Fungus gnats (Diptera, Mycetophilidae) of the USSR" volume="1982" year="1982">Zaitzev (1982)</bibRefCitation>
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and
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<bibRefCitation author="Kurina, O" journalOrPublisher="An international Journal of Dipterological Research" pageId="15" pageNumber="46" pagination="69 - 71" title="Greenomyia mongolica Lastovka et Matile, 1974 (Diptera, Mycetophilidae) found in Estonia." volume="8" year="1997">Kurina (1997)</bibRefCitation>
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. Our association of males and females are based on multiple simultaneous findings in trap samples (see above) that agrees with previous descriptions of the female. In the Pre-Balkan mountain range in Bulgaria, the species has been collected in xerotermic oak forest (
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<bibRefCitation author="Bechev, D" journalOrPublisher="Travaux Scientifiques d'Universite de Plovdiv, Animalia" pageId="14" pageNumber="45" pagination="51 - 66" title="Seasonal activity and vertical distribution of fungus gnats (Diptera: Sciaroidea, excluding Sciaridae) in the western and the central parts of Stara Planina ridge (Bulgaria)." volume="36" year="2000">Bechev 2000</bibRefCitation>
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). The species was quite common in samples taken in a bait trap, operated on the basis of a mixture of fermenting sugar and red wine, in southern Estonia (see also
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<bibRefCitation author="Kurina, O" journalOrPublisher="Sahlbergia" pageId="15" pageNumber="46" pagination="18 - 22" title="A review of Estonian wood gnats (Diptera: Anisopodidae)." volume="11" year="2006">Kurina 2006</bibRefCitation>
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). The above-mentioned specimens from Greece are the first records from the country.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |