360 lines
44 KiB
XML
360 lines
44 KiB
XML
<document id="4FF10C67C64A502A103A38AF6C6AE8DC" ID-DOI="10.11646/zootaxa.5374.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10152641" ID-ZooBank="75A1CC8A-D4EB-4457-86D0-01A89891C535" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700265469172" checkinUser="plazi" docAuthor="Volynkin, Anton V., Titov, Sergey V., Matov, Alexey Yu., Tóth, Balázs, Saldaitis, Aidas, Rakhimov, Ruslan D. & Egorov, Petr V." docDate="2023" docId="FA5487A0FFD5A621FF344BE3FEC53441" docLanguage="en" docName="zootaxa.5374.3.5.pdf" docOrigin="Zootaxa 5374 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5374.3.5/52296" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isochlora hreblayi Volynkin, Toth, Titov & Saldaitis 2023, sp. n." docType="treatment" docVersion="2" lastPageNumber="413" masterDocId="066DFFD8FFD7A625FFA34F4CFFC13050" masterDocTitle="On the taxonomy of the genus Isochlora Staudinger with descriptions of two new species from Mongolia and Qinghai, China (Lepidoptera: Noctuidae: Noctuinae)" masterLastPageNumber="423" masterPageNumber="409" pageNumber="411" updateTime="1700265878127" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6181C0DA0DF18F353287CACE21E7E37A">On the taxonomy of the genus Isochlora Staudinger with descriptions of two new species from Mongolia and Qinghai, China (Lepidoptera: Noctuidae: Noctuinae)</mods:title>
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<mods:namePart id="A6FB9AD266AEB176AD78A24229D8756F">Volynkin, Anton V.</mods:namePart>
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<mods:namePart id="27E17B07CFC312A795B78D7646DBDF9F">Titov, Sergey V.</mods:namePart>
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<mods:affiliation id="0DD65F589FF193EE1D3A1E9741BD3BB9">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. & Department of Biology and Ecology; the Research Centre for Environmental “ Monitoring ”, Toraighyrov University, Lomova str. 64, KZ- 140008, Pavlodar, Kazakhstan. & Scientific Center “ Bioarchaeology ”, Institute of Archaeological Research, Margulan University, Olzhabay Batyr str. 60, KZ- 140002, Pavlodar, Kazakhstan. & sandibicumar @ gmail. com; https: // orcid. org / 0000 - 0003 - 2196 - 280 X</mods:affiliation>
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<mods:namePart id="D8CA23294C675184446B532B60871981">Matov, Alexey Yu.</mods:namePart>
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<mods:affiliation id="4F369C4018DF668B2F166E5D19333814">Zoological Institute, Russian Academy of Sciences, Universitetskaya Embankment 1, St. - Petersburg, RF- 199034 Russia. alexey. matov @ zin. ru; https: // orcid. org / 0000 - 0002 - 6066 - 6440</mods:affiliation>
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<mods:nameIdentifier id="DEF78C5C0AD18D8B174A81700D526DF7" type="email">alexey.matov@zin.ru</mods:nameIdentifier>
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<mods:namePart id="ED3F8B8A2B3131871E9A7A8EAB9E3F43">Tóth, Balázs</mods:namePart>
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<mods:affiliation id="66C9C819E4F3B9D4CDE668DD47133C5F">Department of Zoology, Hungarian Natural History Museum, Baross utca, 13, H- 1088 Budapest, Hungary. toth. balazs @ nhmus. hu; https: // orcid. org / 0000 - 0002 - 6089 - 1218</mods:affiliation>
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<mods:namePart id="BEC367B514055684EB62C518A957BAF5">Saldaitis, Aidas</mods:namePart>
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<mods:namePart id="210A38F034DF66E221CB9CBFC1EC0270">Rakhimov, Ruslan D.</mods:namePart>
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<mods:affiliation id="D98ADE1E1A1EDAB13F19DE88DA87EC23">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. & frontosa 88 @ mail. ru; https: // orcid. org / 0000 - 0001 - 5650 - 7239</mods:affiliation>
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<mods:namePart id="159834EEBA13FC4E816EAE7F093829FA">Egorov, Petr V.</mods:namePart>
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<mods:affiliation id="558761D016258D4C4255C524DC4D9D1F">Institute of Zoology, Entomology laboratory, Al-Farabi Avenue, 93, KZ- 050060, Almaty, Kazakhstan. & ent. helios _ 2908 @ mail. ru; https: // orcid. org / 0000 - 0001 - 7686 - 6845</mods:affiliation>
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<mods:date id="CE41CC2C1569F1634EE3C47CD3BA9C79">2023</mods:date>
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<taxonomicName id="B5FD4D35FFD5A627FF344BE3FCDD3499" authority="Volynkin, Toth, Titov & Saldaitis" authorityName="Volynkin, Toth, Titov & Saldaitis" authorityYear="2023" box="[151,796,1198,1226]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="hreblayi" status="sp. nov.">
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<emphasis id="4089EAA4FFD5A627FF344BE3FEB4349A" bold="true" box="[151,373,1199,1226]" italics="true" pageId="2" pageNumber="411">Isochlora hreblayi</emphasis>
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Volynkin, Tóth, Titov & Saldaitis
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,
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(
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,
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<figureCitation id="EAC62A33FFD5A627FF574B9FFED034BD" box="[244,273,1235,1261]" captionStart="FIGURES 20–23" captionStartId="9.[151,264,1938,1962]" captionTargetBox="[156,1431,187,1902]" captionTargetId="figure-18@9.[151,1435,181,1915]" captionTargetPageId="9" captionText="FIGURES 20–23. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 20 and 21 in MfN; 22 in MH/ HNHM; 23 in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152649" httpUri="https://zenodo.org/record/10152649/files/figure.png" pageId="2" pageNumber="411">22</figureCitation>
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)
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<paragraph id="724236B6FFD5A627FF344A57FEA5352D" blockId="2.[151,1437,1307,1981]" pageId="2" pageNumber="411">
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<emphasis id="4089EAA4FFD5A627FF344A57FEFF3565" bold="true" box="[151,318,1307,1333]" pageId="2" pageNumber="411">Type material</emphasis>
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.
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<materialsCitation id="C2953CEBFFD5A627FEEE4A57FEA5352D" collectionCode="MH, HNHM" collectorName="Alph. & Kobdo. & A. Meinhard." pageId="2" pageNumber="411" specimenCount="1" typeStatus="holotype">
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<emphasis id="4089EAA4FFD5A627FEEE4A57FE7B3565" bold="true" box="[333,442,1307,1333]" pageId="2" pageNumber="411">
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<typeStatus id="AD468814FFD5A627FEEE4A57FE7B3565" box="[333,442,1307,1333]" pageId="2" pageNumber="411" type="holotype">Holotype</typeStatus>
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</emphasis>
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(
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<figureCitation id="EAC62A33FFD5A627FE6A4A57FDD03565" box="[457,529,1307,1333]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[163,1423,182,1902]" captionTargetId="figure-18@3.[158,1430,179,1918]" captionTargetPageId="3" captionText="FIGURES 1–8. Isochlora spp.: adults. Depositories of the specimens: 1 and 2 in MfN; 3 in MH/HNHM; 4–6 and 8 in ZISP; 7 in CAV." figureDoi="http://doi.org/10.5281/zenodo.10152643" httpUri="https://zenodo.org/record/10152643/files/figure.png" pageId="2" pageNumber="411">Figs 3</figureCitation>
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,
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<figureCitation id="EAC62A33FFD5A627FDBD4A57FDFF3565" box="[542,574,1307,1333]" captionStart="FIGURES 20–23" captionStartId="9.[151,264,1938,1962]" captionTargetBox="[156,1431,187,1902]" captionTargetId="figure-18@9.[151,1435,181,1915]" captionTargetPageId="9" captionText="FIGURES 20–23. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 20 and 21 in MfN; 22 in MH/ HNHM; 23 in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152649" httpUri="https://zenodo.org/record/10152649/files/figure.png" pageId="2" pageNumber="411">22</figureCitation>
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): male,
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<taxonomicName id="B5FD4D35FFD5A627FD3B4A57FCCA3565" authorityName="Alpheraky" authorityYear="1897" box="[664,779,1307,1333]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="arctomys">“arctomys</taxonomicName>
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|
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<collectorName id="DF085360FFD5A627FCBD4A57FC9E3565" box="[798,863,1307,1333]" pageId="2" pageNumber="411">Alph.</collectorName>
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? |
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<collectorName id="DF085360FFD5A627FCDD4A57FC103565" box="[894,977,1307,1333]" pageId="2" pageNumber="411">Kobdo.</collectorName>
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| 6.VII.09. |
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<collectorName id="DF085360FFD5A627FBFD4A57FB393565" box="[1118,1272,1307,1333]" pageId="2" pageNumber="411">A. Meinhard.</collectorName>
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” (upper side) | “16812.” (under side) / “E[x] coll. L. | Krulikovsky | coll. L. | Sheljuzhko” / pink label “gen.prep. Hreblay | N: 10686 ³” (
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<collectionCode id="14ECAE73FFD5A627FF644A2FFF37352D" box="[199,246,1379,1405]" country="Switzerland" name="Naturhistorisches Museum, Basel" pageId="2" pageNumber="411">MH</collectionCode>
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/
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<collectionCode id="14ECAE73FFD5A627FF5C4A2FFE98352D" box="[255,345,1379,1405]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="2" pageNumber="411">HNHM</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="724236B6FFD5A627FF644ACBFB1037ED" blockId="2.[151,1437,1307,1981]" pageId="2" pageNumber="411">
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<emphasis id="4089EAA4FFD5A627FF644ACBFE2435F1" bold="true" box="[199,485,1415,1441]" pageId="2" pageNumber="411">Note on the type locality</emphasis>
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||
. The label of the
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<typeStatus id="AD468814FFD5A627FD064ACBFCC635F1" box="[677,775,1415,1441]" pageId="2" pageNumber="411" type="holotype">holotype</typeStatus>
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contains the name of A.A. Meinhard (engineer and naturalist at Tomsk University), who never visited
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<collectingCountry id="0AEA7626FFD5A627FD274AE7FD303595" box="[644,753,1451,1477]" name="Mongolia" pageId="2" pageNumber="411">Mongolia</collectingCountry>
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but processed
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<taxonomicName id="B5FD4D35FFD5A627FC3A4AE7FBE13595" authorityName="Linnaeus" authorityYear="1758" box="[921,1056,1451,1477]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
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materials collected by
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<collectionCode id="14ECAE73FFD5A627FA874AE0FAF93596" box="[1316,1336,1452,1478]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
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.
|
||
<collectionCode id="14ECAE73FFD5A627FA9F4AE0FA913596" box="[1340,1360,1452,1478]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
|
||
. Sapozhnikov in a series of expeditions to the upper reaches of the Kobdo River (an old name of Khovd-Gol River) basin in western
|
||
<collectingCountry id="0AEA7626FFD5A627FEB14ABFFEBE365D" box="[274,383,1523,1549]" name="Mongolia" pageId="2" pageNumber="411">Mongolia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="166C4B47FFD5A627FE2E4ABFFD8B365D" author="Meinhard, A. A." box="[397,586,1523,1549]" pageId="2" pageNumber="411" pagination="1 - 17" refId="ref8280" refString="Meinhard, A. A. (1910 a) A list of Lepidoptera collection of scientific expedition in south-eastern part of Russian Altai and Mongolia (Kobdo) in 1905 - 1906 by Prof. V. V. Sapozhnikov. Izvestiya of the Imperial Tomsk University, 37, 1 - 17. [in Russian]" type="journal article" year="1910">
|
||
Meinhard 1910
|
||
<bibRefCitation id="166C4B47FFD5A627FD9E4ABFFD8B365D" author="Meinhard, A. A." box="[573,586,1523,1549]" pageId="2" pageNumber="411" pagination="19 - 27" refId="ref8339" refString="Meinhard, A. A. (1910 b) A list of Lepidoptera collection of expedition in Mongolia (Kobdo) in 1908 by Prof. V. V. Sapozhnikov. Izvestiya of the Imperial Tomsk University, 37, 19 - 27. [in Russian]" type="journal article" year="1910">a</bibRefCitation>
|
||
</bibRefCitation>
|
||
, 1910b;
|
||
<bibRefCitation id="166C4B47FFD5A627FD094ABFFC96365D" author="Nekrylov, S. A." box="[682,855,1523,1549]" pageId="2" pageNumber="411" refId="ref8389" refString="Nekrylov, S. A. (2011) Tomsk University - the first scientific center in the Asian part of Russia's history (mid- 1870 s - 1919). Vol. 2. Tomsk University publishing house, Tomsk, 598 pp. [in Russian]" type="book" year="2011">Nekrylov 2011</bibRefCitation>
|
||
). It is possible to assume that the specimen was collected by
|
||
<collectionCode id="14ECAE73FFD5A627FEA04954FED63662" box="[259,279,1560,1586]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
|
||
.
|
||
<collectionCode id="14ECAE73FFD5A627FEB84954FEEE3662" box="[283,303,1560,1586]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
|
||
. Sapozhnikov, and Meinhard’s name on the label belongs to the tentative identification as [
|
||
<taxonomicName id="B5FD4D35FFD5A627FA8B495BFA523661" authorityName="Staudinger" authorityYear="1882" box="[1320,1427,1559,1585]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4089EAA4FFD5A627FA8B495BFA523661" box="[1320,1427,1559,1585]" italics="true" pageId="2" pageNumber="411">Isochlora</emphasis>
|
||
</taxonomicName>
|
||
]
|
||
<taxonomicName id="B5FD4D35FFD5A627FF344977FE923605" authority="Alph." authorityName="Alpheraky" authorityYear="1897" box="[151,339,1595,1621]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="arctomys">
|
||
“
|
||
<emphasis id="4089EAA4FFD5A627FF074970FECB3605" box="[164,266,1596,1621]" italics="true" pageId="2" pageNumber="411">arctomys</emphasis>
|
||
Alph.
|
||
</taxonomicName>
|
||
?”. However, the date on the label is also questionable. The year is handwritten illegibly and may be [19]04 or [19]09, but the first
|
||
<collectionCode id="14ECAE73FFD5A627FE44492CFE3A362A" box="[487,507,1632,1658]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
|
||
.
|
||
<collectionCode id="14ECAE73FFD5A627FE5C492CFDD2362A" box="[511,531,1632,1658]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
|
||
. Sapozhnikov expedition was in 1905 whereas on
|
||
<date id="06431076FFD5A627FBEB4913FB103629" box="[1096,1233,1631,1657]" pageId="2" pageNumber="411" value="1909-07-06">July 6, 1909</date>
|
||
he was in the
|
||
<collectingCountry id="0AEA7626FFD5A627FAD1492CFF3836CD" name="Kazakhstan" pageId="2" pageNumber="411">Kazakhstan</collectingCountry>
|
||
part of Altai Mountains, on his way to
|
||
<collectingCountry id="0AEA7626FFD5A627FD1649CFFCE336CD" box="[693,802,1667,1693]" name="Mongolia" pageId="2" pageNumber="411">Mongolia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="166C4B47FFD5A627FC9349CFFBC636CD" author="Sapozhnikov, V. V." box="[816,1031,1667,1693]" pageId="2" pageNumber="411" refId="ref8463" refString="Sapozhnikov, V. V. (1911) Mongolian Altai at the sources of the Irtysh and Kobdo. Travels 1905 - 1909. Siberian Printing Partnership, Tomsk, 408 p. [in Russian]" type="book" year="1911">Sapozhnikov 1911</bibRefCitation>
|
||
). According to the diaries published after
|
||
<collectionCode id="14ECAE73FFD5A627FF7249E4FF243692" box="[209,229,1704,1730]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
|
||
.
|
||
<collectionCode id="14ECAE73FFD5A627FF4A49E4FF3C3692" box="[233,253,1704,1730]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="411" type="Museum">V</collectionCode>
|
||
. Sapozhnikov expeditions (
|
||
<bibRefCitation id="166C4B47FFD5A627FD9349EBFCC43691" author="Sapozhnikov, V. V." box="[560,773,1703,1729]" pageId="2" pageNumber="411" refId="ref8463" refString="Sapozhnikov, V. V. (1911) Mongolian Altai at the sources of the Irtysh and Kobdo. Travels 1905 - 1909. Siberian Printing Partnership, Tomsk, 408 p. [in Russian]" type="book" year="1911">Sapozhnikov 1911</bibRefCitation>
|
||
), the only year suitable for the date and geography is 1905. On
|
||
<date id="06431076FFD5A627FF624987FE8A36B6" box="[193,331,1739,1766]" pageId="2" pageNumber="411" value="1905-07-06">July 6, 1905</date>
|
||
the expedition was working exactly in the highlands of Mongolian
|
||
<collectingRegion id="B039F854FFD5A627FBE24987FBB836B5" box="[1089,1145,1739,1765]" country="Russia" name="Altay" pageId="2" pageNumber="411">Altai</collectingRegion>
|
||
near glaciers in the upper stream of Tsagan-Gol River, which is a tributary of the river Khovd-Gol (=Kobdo) (
|
||
<figureCitation id="EAC62A33FFD5A627FB9D49A3FB503759" box="[1086,1169,1775,1801]" captionStart="FIGURE 33" captionStartId="12.[151,250,967,991]" captionTargetBox="[151,1436,181,943]" captionTargetId="figure-23@12.[151,1436,181,943]" captionTargetPageId="12" captionText="FIGURE 33. Distribution map of Isochlora spp. in the Tien Shan Massif and Mongolian Altai." figureDoi="http://doi.org/10.5281/zenodo.10152657" httpUri="https://zenodo.org/record/10152657/files/figure.png" pageId="2" pageNumber="411">Fig. 33</figureCitation>
|
||
). As the closest species,
|
||
<taxonomicName id="B5FD4D35FFD5A627FF344858FED5377D" baseAuthorityName="Alpheraky" baseAuthorityYear="1882" box="[151,276,1812,1837]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD5A627FF344858FED5377D" box="[151,276,1812,1837]" italics="true" pageId="2" pageNumber="411">I. intricans</emphasis>
|
||
</taxonomicName>
|
||
, as well as most other species in the genus
|
||
<taxonomicName id="B5FD4D35FFD5A627FCAE485FFCB9377D" authorityName="Staudinger" authorityYear="1882" box="[781,888,1811,1837]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4089EAA4FFD5A627FCAE485FFCB9377D" box="[781,888,1811,1837]" italics="true" pageId="2" pageNumber="411">Isochlora</emphasis>
|
||
</taxonomicName>
|
||
, inhabits the highest mountain biotopes, the assumed collection locality of
|
||
<taxonomicName id="B5FD4D35FFD5A627FE714874FD853701" authority="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov, 2023" authorityName="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov" authorityYear="2023" box="[466,580,1847,1873]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="species" species="hreblayi" status="sp. n.">
|
||
<emphasis id="4089EAA4FFD5A627FE714874FD853701" box="[466,580,1847,1873]" italics="true" pageId="2" pageNumber="411">I. hreblayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4089EAA4FFD5A627FDE9487BFD493701" bold="true" box="[586,648,1847,1873]" pageId="2" pageNumber="411">
|
||
<taxonomicNameLabel id="5BBA57DFFFD5A627FDE9487BFD493701" box="[586,648,1847,1873]" pageId="2" pageNumber="411" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
matches the genus habitat preferences well. Additionally, in the collection of the
|
||
<collectingRegion id="B039F854FFD5A627FEB34817FE5B3725" box="[272,410,1883,1909]" country="Russia" name="Tomsk" pageId="2" pageNumber="411">Tomsk State</collectingRegion>
|
||
University (
|
||
<collectingRegion id="B039F854FFD5A627FD864817FDB73725" box="[549,630,1883,1909]" country="Russia" name="Tomsk" pageId="2" pageNumber="411">Tomsk</collectingRegion>
|
||
,
|
||
<collectingCountry id="0AEA7626FFD5A627FD234817FD0E3725" box="[640,719,1883,1909]" name="Russia" pageId="2" pageNumber="411">Russia</collectingCountry>
|
||
), there is a specimen of the fritillary
|
||
<taxonomicName id="B5FD4D35FFD5A627FBC44817FE0537C9" authority="P. Gorbunov & Korshunov" authorityName="P. Gorbunov & Korshunov" class="Insecta" family="Nymphalidae" genus="Clossiana" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="subSpecies" species="tritonia" subSpecies="matveevi">
|
||
<emphasis id="4089EAA4FFD5A627FBC44817FA5C3725" box="[1127,1437,1883,1909]" italics="true" pageId="2" pageNumber="411">Clossiana tritonia matveevi</emphasis>
|
||
P. Gorbunov & Korshunov
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="B5FD4D35FFD5A627FE704833FDAF37C9" box="[467,622,1919,1945]" class="Insecta" family="Nymphalidae" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="411" phylum="Arthropoda" rank="family">Nymphalidae</taxonomicName>
|
||
) with a label “
|
||
<collectingCountry id="0AEA7626FFD5A627FCB24833FC4237C9" box="[785,899,1919,1945]" name="Mongolia" pageId="2" pageNumber="411">Mongolia</collectingCountry>
|
||
, Kobdo, Tsagan-Kol [Tsagan-Gol],
|
||
<date id="06431076FFD5A627FAB64833FA5337CA" box="[1301,1426,1919,1946]" pageId="2" pageNumber="411" value="1905-07-06">6.VII.1905</date>
|
||
” (
|
||
<bibRefCitation id="166C4B47FFD5A627FF3C48EFFE1F37ED" author="Dubatolov, V. V. & Kosterin, O. E." box="[159,478,1955,1981]" pageId="2" pageNumber="411" pagination="184 - 188" refId="ref7646" refString="Dubatolov, V. V. & Kosterin, O. E. (2010) What is Matveev's Fritillary (Clossiana matveevi P. Gorbunov et Korshunov in Korshunov, Gorbunov, 1995)? Amurian zoological journal, 2 (2), 184 - 188. [in Russian] https: // doi. org / 10.33910 / 1999 - 4079 - 2010 - 2 - 2 - 184 - 188" type="journal article" year="2010">Dubatolov & Kosterin 2010</bibRefCitation>
|
||
), which also confirms that the expedition collected insects that day.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="2682663EFFD4A626FF3448DBFF293784" ID-DOI="http://doi.org/10.5281/zenodo.10152643" ID-Zenodo-Dep="10152643" httpUri="https://zenodo.org/record/10152643/files/figure.png" pageId="3" pageNumber="412" startId="3.[151,264,1943,1967]" targetBox="[163,1423,182,1902]" targetPageId="3" targetType="figure">
|
||
<paragraph id="724236B6FFD4A626FF3448DBFF293784" blockId="3.[151,1437,1943,2004]" pageId="3" pageNumber="412">
|
||
<emphasis id="4089EAA4FFD4A626FF3448DBFEF537FF" bold="true" box="[151,308,1943,1967]" pageId="3" pageNumber="412">FIGURES 1–8</emphasis>
|
||
.
|
||
<taxonomicName id="B5FD4D35FFD4A626FEE248D4FE1737FF" box="[321,470,1944,1968]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="412" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="4089EAA4FFD4A626FEE248D4FE6337E0" box="[321,418,1944,1968]" italics="true" pageId="3" pageNumber="412">Isochlora</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
: adults. Depositories of the specimens: 1 and 2 in MfN; 3 in MH/HNHM; 4–6 and 8 in ZISP; 7 in CAV.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3AE7653DFFD3A621FF644FDAFB2531D9" pageId="4" pageNumber="413" type="diagnosis">
|
||
<paragraph id="724236B6FFD3A621FF644FDAFB2531D9" blockId="4.[151,1437,150,1041]" pageId="4" pageNumber="413">
|
||
<emphasis id="4089EAA4FFD3A621FF644FDAFEFB30E0" bold="true" box="[199,314,150,176]" pageId="4" pageNumber="413">Diagnosis</emphasis>
|
||
. The new species (
|
||
<figureCitation id="EAC62A33FFD3A621FDB34FDBFD9330E1" box="[528,594,151,177]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[163,1423,182,1902]" captionTargetId="figure-18@3.[158,1430,179,1918]" captionTargetPageId="3" captionText="FIGURES 1–8. Isochlora spp.: adults. Depositories of the specimens: 1 and 2 in MfN; 3 in MH/HNHM; 4–6 and 8 in ZISP; 7 in CAV." figureDoi="http://doi.org/10.5281/zenodo.10152643" httpUri="https://zenodo.org/record/10152643/files/figure.png" pageId="4" pageNumber="413">Fig. 3</figureCitation>
|
||
) is externally very similar to
|
||
<taxonomicName id="B5FD4D35FFD3A621FC3F4FDBFBD630E0" baseAuthorityName="Alpheraky" baseAuthorityYear="1882" box="[924,1047,151,176]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD3A621FC3F4FDBFBD630E0" box="[924,1047,151,176]" italics="true" pageId="4" pageNumber="413">I. intricans</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="EAC62A33FFD3A621FB864FDBFB4830E1" box="[1061,1161,151,177]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[163,1423,182,1902]" captionTargetId="figure-18@3.[158,1430,179,1918]" captionTargetPageId="3" captionText="FIGURES 1–8. Isochlora spp.: adults. Depositories of the specimens: 1 and 2 in MfN; 3 in MH/HNHM; 4–6 and 8 in ZISP; 7 in CAV." figureDoi="http://doi.org/10.5281/zenodo.10152643" httpUri="https://zenodo.org/record/10152643/files/figure.png" pageId="4" pageNumber="413">Figs 4–8</figureCitation>
|
||
) but differs in the somewhat more distinct forewing markings. Compared to
|
||
<taxonomicName id="B5FD4D35FFD3A621FD494FF7FCA43084" baseAuthorityName="Alpheraky" baseAuthorityYear="1882" box="[746,869,187,212]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD3A621FD494FF7FCA43084" box="[746,869,187,212]" italics="true" pageId="4" pageNumber="413">I. intricans</emphasis>
|
||
</taxonomicName>
|
||
, the male genital capsule of
|
||
<taxonomicName id="B5FD4D35FFD3A621FB054FF7FAD83085" authority="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov, 2023" authorityName="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov" authorityYear="2023" box="[1190,1305,187,213]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="hreblayi" status="sp. n.">
|
||
<emphasis id="4089EAA4FFD3A621FB054FF7FAD83085" box="[1190,1305,187,213]" italics="true" pageId="4" pageNumber="413">I. hreblayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4089EAA4FFD3A621FA834FF7FAA13085" bold="true" box="[1312,1376,187,213]" pageId="4" pageNumber="413">
|
||
<taxonomicNameLabel id="5BBA57DFFFD3A621FA834FF7FAA13085" box="[1312,1376,187,213]" pageId="4" pageNumber="413" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="EAC62A33FFD3A621FACC4FF7FF7530A9" captionStart="FIGURES 20–23" captionStartId="9.[151,264,1938,1962]" captionTargetBox="[156,1431,187,1902]" captionTargetId="figure-18@9.[151,1435,181,1915]" captionTargetPageId="9" captionText="FIGURES 20–23. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 20 and 21 in MfN; 22 in MH/ HNHM; 23 in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152649" httpUri="https://zenodo.org/record/10152649/files/figure.png" pageId="4" pageNumber="413">Fig. 22</figureCitation>
|
||
) has a somewhat shorter uncus, a slightly shorter penicular lobe, and a reduced harpe whereas in
|
||
<taxonomicName id="B5FD4D35FFD3A621FB5B4F93FAB230A8" baseAuthorityName="Alpheraky" baseAuthorityYear="1882" box="[1272,1395,223,248]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD3A621FB5B4F93FAB230A8" box="[1272,1395,223,248]" italics="true" pageId="4" pageNumber="413">I. intricans</emphasis>
|
||
</taxonomicName>
|
||
the harpe is present although very short (
|
||
<figureCitation id="EAC62A33FFD3A621FD904E4FFD46314D" box="[563,647,259,285]" captionStart="FIGURES 20–23" captionStartId="9.[151,264,1938,1962]" captionTargetBox="[156,1431,187,1902]" captionTargetId="figure-18@9.[151,1435,181,1915]" captionTargetPageId="9" captionText="FIGURES 20–23. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 20 and 21 in MfN; 22 in MH/ HNHM; 23 in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152649" httpUri="https://zenodo.org/record/10152649/files/figure.png" pageId="4" pageNumber="413">Fig. 23</figureCitation>
|
||
). Additionally, in
|
||
<taxonomicName id="B5FD4D35FFD3A621FCEE4E4FFC01314D" authority="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov, 2023" authorityName="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov" authorityYear="2023" box="[845,960,259,285]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="hreblayi" status="sp. n.">
|
||
<emphasis id="4089EAA4FFD3A621FCEE4E4FFC01314D" box="[845,960,259,285]" italics="true" pageId="4" pageNumber="413">I. hreblayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4089EAA4FFD3A621FC654E4FFBC4314D" bold="true" box="[966,1029,259,285]" pageId="4" pageNumber="413">
|
||
<taxonomicNameLabel id="5BBA57DFFFD3A621FC654E4FFBC4314D" box="[966,1029,259,285]" pageId="4" pageNumber="413" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, the distal setose cluster of the valva is longer, sparser and consists of shorter setae than in
|
||
<taxonomicName id="B5FD4D35FFD3A621FD564E6BFCB13110" baseAuthorityName="Alpheraky" baseAuthorityYear="1882" box="[757,880,295,320]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD3A621FD564E6BFCB13110" box="[757,880,295,320]" italics="true" pageId="4" pageNumber="413">I. intricans</emphasis>
|
||
</taxonomicName>
|
||
. The phallus of the new species is shorter than in
|
||
<taxonomicName id="B5FD4D35FFD3A621FF344E07FED23134" baseAuthorityName="Alpheraky" baseAuthorityYear="1882" box="[151,275,331,356]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD3A621FF344E07FED23134" box="[151,275,331,356]" italics="true" pageId="4" pageNumber="413">I. intricans</emphasis>
|
||
</taxonomicName>
|
||
. The vesica of
|
||
<taxonomicName id="B5FD4D35FFD3A621FE624E07FDF43135" authority="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov, 2023" authorityName="Volynkin & Titov & Matov & Tóth & Saldaitis & Rakhimov & Egorov" authorityYear="2023" box="[449,565,331,357]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="hreblayi" status="sp. n.">
|
||
<emphasis id="4089EAA4FFD3A621FE624E07FDF43135" box="[449,565,331,357]" italics="true" pageId="4" pageNumber="413">I. hreblayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="4089EAA4FFD3A621FD9E4E07FDBC3135" bold="true" box="[573,637,331,357]" pageId="4" pageNumber="413">
|
||
<taxonomicNameLabel id="5BBA57DFFFD3A621FD9E4E07FDBC3135" box="[573,637,331,357]" pageId="4" pageNumber="413" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="B5FD4D35FFD3A621FCB04E07FC4E3134" baseAuthorityName="Alpheraky" baseAuthorityYear="1882" box="[787,911,331,356]" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD3A621FCB04E07FC4E3134" box="[787,911,331,356]" italics="true" pageId="4" pageNumber="413">I. intricans</emphasis>
|
||
</taxonomicName>
|
||
in the lack of the proximal row of cornuti, the broader medial section, and the smaller distal cluster of scobination consisting of weaker spinules.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AE7653DFFD3A621FF644EDEFE5033D1" pageId="4" pageNumber="413" type="description">
|
||
<paragraph id="724236B6FFD3A621FF644EDEFB38330D" blockId="4.[151,1437,150,1041]" pageId="4" pageNumber="413">
|
||
<emphasis id="4089EAA4FFD3A621FF644EDEFE9031FC" bold="true" box="[199,337,402,428]" pageId="4" pageNumber="413">Description</emphasis>
|
||
.
|
||
<emphasis id="4089EAA4FFD3A621FEFD4EDEFE6331FC" bold="true" box="[350,418,402,428]" pageId="4" pageNumber="413">Adult</emphasis>
|
||
(
|
||
<figureCitation id="EAC62A33FFD3A621FE134EDFFE3231FD" box="[432,499,403,429]" captionStart="FIGURES 1–8" captionStartId="3.[151,264,1943,1967]" captionTargetBox="[163,1423,182,1902]" captionTargetId="figure-18@3.[158,1430,179,1918]" captionTargetPageId="3" captionText="FIGURES 1–8. Isochlora spp.: adults. Depositories of the specimens: 1 and 2 in MfN; 3 in MH/HNHM; 4–6 and 8 in ZISP; 7 in CAV." figureDoi="http://doi.org/10.5281/zenodo.10152643" httpUri="https://zenodo.org/record/10152643/files/figure.png" pageId="4" pageNumber="413">Fig. 3</figureCitation>
|
||
). Forewing length 15.5 mm in male
|
||
<typeStatus id="AD468814FFD3A621FC204EDFFC2831FD" box="[899,1001,403,429]" pageId="4" pageNumber="413" type="holotype">holotype</typeStatus>
|
||
. Male antenna bipectinate. Head ochreous brown. Thorax pale rust. Forewing ground colour ochreous, markings pale rust. Antemedial area with diffuse elliptical rust spot subbasally and medially. Antemedial line thin and irregularly zigzagged. Medial area rust posteriorly and in cell and pale ochreous along costal margin. Orbicular stigma indistinct. Reniform stigma unicolorous dark brown with diffuse margins. Postmedial line curved outwards between vein A1 and costal margin, dentate on veins. Subterminal line diffuse, irregularly zigzagged, forming elongate trapezoid diffuse dash at costal margin. Terminal line thin, continuous, pale rust. Cilia ochreous with admixture of rust. Hindwing brown with orange rust suffusion along costal margin. Cilia ochreous.
|
||
<emphasis id="4089EAA4FFD3A621FD3A4DC3FC8232F8" bold="true" box="[665,835,654,681]" pageId="4" pageNumber="413">Male genitalia</emphasis>
|
||
(
|
||
<figureCitation id="EAC62A33FFD3A621FCF34DC3FC6232F9" box="[848,931,655,681]" captionStart="FIGURES 20–23" captionStartId="9.[151,264,1938,1962]" captionTargetBox="[156,1431,187,1902]" captionTargetId="figure-18@9.[151,1435,181,1915]" captionTargetPageId="9" captionText="FIGURES 20–23. Isochlora spp.: male genitalia. Depositories of the specimens dissected: 20 and 21 in MfN; 22 in MH/ HNHM; 23 in ZISP." figureDoi="http://doi.org/10.5281/zenodo.10152649" httpUri="https://zenodo.org/record/10152649/files/figure.png" pageId="4" pageNumber="413">Fig. 22</figureCitation>
|
||
). Uncus medially dilated and distally tapered, weakly setose. Tegumen with semicircular penicular lobe. Vinculum slightly longer than tegumen, V-shaped. Valva lobular with almost parallel margins, slightly upcurved and apically rounded, its distal third with cluster of weak setae. Sacculus short (ca. 1/3 of valva length), elliptical. Clasper thin, distally dilated, harpe reduced. Juxta shieldlike. Anellus weakly sclerotised. Phallus cylindrical, slightly downcurved medially, with rounded coecum. Vesica recurved ventrally, dilated medially and tapered distally, with cluster of minute spinules subapically.
|
||
</paragraph>
|
||
<paragraph id="724236B6FFD3A621FF644C2BFE5033D1" blockId="4.[151,1437,150,1041]" box="[199,401,871,897]" pageId="4" pageNumber="413">Female unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AE7653DFFD3A621FF644CC6FC8333F5" box="[199,834,906,933]" pageId="4" pageNumber="413" type="distribution">
|
||
<paragraph id="724236B6FFD3A621FF644CC6FC8333F5" blockId="4.[151,1437,150,1041]" box="[199,834,906,933]" pageId="4" pageNumber="413">
|
||
<emphasis id="4089EAA4FFD3A621FF644CC6FE9833F4" bold="true" box="[199,345,906,932]" pageId="4" pageNumber="413">Distribution</emphasis>
|
||
. Western Mongolia (Mongolian
|
||
<collectingRegion id="B039F854FFD3A621FD644CC7FD3E33F5" box="[711,767,907,933]" country="Russia" name="Altay" pageId="4" pageNumber="413">Altai</collectingRegion>
|
||
Mts).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3AE7653DFFD3A621FF644CE3FEC53441" pageId="4" pageNumber="413" type="etymology">
|
||
<paragraph id="724236B6FFD3A621FF644CE3FEC53441" blockId="4.[151,1437,150,1041]" pageId="4" pageNumber="413">
|
||
<emphasis id="4089EAA4FFD3A621FF644CE3FE893399" bold="true" box="[199,328,943,969]" pageId="4" pageNumber="413">Etymology</emphasis>
|
||
. The new species is named after the late Dr Márton Hreblay (1963–2000), a renowned lepidopterist, researcher of Eurasiatic
|
||
<taxonomicName id="B5FD4D35FFD3A621FE0A4C9FFDDE33BD" box="[425,543,979,1005]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="family">Noctuidae</taxonomicName>
|
||
, who first illustrated the
|
||
<typeStatus id="AD468814FFD3A621FC9A4C9FFC5A33BD" box="[825,923,979,1005]" pageId="4" pageNumber="413" type="holotype">holotype</typeStatus>
|
||
of the new species as
|
||
<taxonomicName id="B5FD4D35FFD3A621FB3A4C9FFF3E3441" authority="(Hreblay et al. 1998)" baseAuthorityName="Hreblay" baseAuthorityYear="1998" class="Insecta" family="Noctuidae" genus="Isochlora" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="413" phylum="Arthropoda" rank="species" species="intricans">
|
||
<emphasis id="4089EAA4FFD3A621FB3A4C9FFAD733BC" box="[1177,1302,979,1004]" italics="true" pageId="4" pageNumber="413">I. intricans</emphasis>
|
||
(
|
||
<bibRefCitation id="166C4B47FFD3A621FA874C9FFF363441" author="Hreblay, M. & Ronkay, L. & Plante, J." pageId="4" pageNumber="413" pagination="69 - 184" refId="ref7926" refString="Hreblay, M., Ronkay, L. & Plante, J. (1998) Contribution to the Noctuidae (Lepidoptera) fauna of Tibet and adjacent regions (II.) A systematic survey of the Tibetan Noctuidae fauna based on the material of the Schafer-expedition (1938 - 1939) and recent expeditions (1993 - 1997) (Lepidoptera: Noctuidae). Esperiana, 6, 69 - 184." type="journal article" year="1998">
|
||
Hreblay
|
||
<emphasis id="4089EAA4FFD3A621FA2B4C9FFF753441" italics="true" pageId="4" pageNumber="413">et al.</emphasis>
|
||
1998
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |