291 lines
24 KiB
XML
291 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.756.24397" ID-GBIF-Dataset="4f06fa64-12e5-416c-8261-d8e1b330e8cb" ID-PMC="PMC5956031" ID-Pensoft-Pub="1313-2970-756-1" ID-PubMed="29773959" ID-ZBK="83CE3672A4E14990A54C5D712D09974E" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-756-1" ModsDocOrigin="ZooKeys 756" ModsDocTitle="Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia" checkinTime="1525927788324" checkinUser="pensoft" docAuthor="Rix, Michael G., Huey, Joel A., Cooper, Steven J. B., Austin, Andrew D. & Harvey, Mark S." docDate="2018" docId="E5B74E6840F550EA62699E2CA6E72A2D" docLanguage="en" docName="ZooKeys 756: 1-121" docOrigin="ZooKeys 756" docSource="http://dx.doi.org/10.3897/zookeys.756.24397" docTitle="Idiosoma arenaceum Rix & Harvey, sp. n." docType="treatment" docUuid="2D4DE886-0589-4BFF-9BA4-D59BB03AFD74" docUuidSource="ZooBank" docVersion="5" lastPageNumber="23" masterDocId="F86B8B0DFFFA8657FFF5FFC4FFCDFFAD" masterDocTitle="Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia" masterLastPageNumber="121" masterPageNumber="1" pageNumber="20" updateTime="1668165771019" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Conservation systematics of the shield-backed trapdoor spiders of the nigrum-group (Mygalomorphae, Idiopidae, Idiosoma): integrative taxonomy reveals a diverse and threatened fauna from south-western Australia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rix, Michael G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huey, Joel A.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Cooper, Steven J. B.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Austin, Andrew D.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harvey, Mark S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>756</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>121</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.756.24397</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.756.24397</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-756-1</mods:identifier>
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<mods:identifier type="ZBK">83CE3672A4E14990A54C5D712D09974E</mods:identifier>
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<mods:identifier type="ZooBank">83CE3672A4E14990A54C5D712D09974E</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="143929125" LSID="urn:lsid:zoobank.org:act:2D4DE886-0589-4BFF-9BA4-D59BB03AFD74" httpUri="http://treatment.plazi.org/id/E5B74E6840F550EA62699E2CA6E72A2D" lastPageId="22" lastPageNumber="23" pageId="19" pageNumber="20">
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<subSubSection pageId="19" pageNumber="20" type="nomenclature">
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<paragraph pageId="19" pageNumber="20">
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<taxonomicName LSID="http://zoobank.org/2D4DE886-0589-4BFF-9BA4-D59BB03AFD74" authority="Rix & Harvey" class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma arenaceum" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="arenaceum">Idiosoma arenaceum Rix & Harvey</taxonomicName>
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<taxonomicNameLabel pageId="19" pageNumber="20">sp. n.</taxonomicNameLabel>
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Figs 11, 12, 19, 20, 25, 57-66, 67-69, 70-78, 374
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="reference_group">
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<paragraph pageId="20" pageNumber="21">
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<taxonomicName class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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<pageBreakToken pageId="20" pageNumber="21" start="start">Idiosoma</pageBreakToken>
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</taxonomicName>
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<normalizedToken originalValue="‘nigrum’">'nigrum'</normalizedToken>
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Main, 1957b: 440 (in part; cited specimens from Canna, N. of Galena and Woolaga Creek).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="type material">
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<paragraph pageId="20" pageNumber="21">Type material.</paragraph>
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<paragraph pageId="20" pageNumber="21">
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Holotype male. Zuytdorp, site ZU1 (IBRA_GES), Western Australia, Australia,
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-27.261667">27°15'42"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="114.025">114°01'30"E</geoCoordinate>
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, wet pitfall trap, 19
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<normalizedToken originalValue="May–">May-</normalizedToken>
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17 August 1995, N. Hall, WAM-CALM Carnarvon Survey (WAM T139527).
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</paragraph>
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<paragraph pageId="20" pageNumber="21">Paratype. 1 ♂, same data as holotype (WAM T41787).</paragraph>
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</subSubSection>
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<subSubSection pageId="20" pageNumber="21" type="materials_examined">
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<paragraph pageId="20" pageNumber="21">Other material examined.</paragraph>
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<paragraph pageId="20" pageNumber="21">
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AUSTRALIA: Western Australia: 2 ♂, Zuytdorp, site ZU3 (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-27.259445">27°15'34"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="114.0675">114°04'03"E</geoCoordinate>
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, wet pitfall trap, 18
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<normalizedToken originalValue="May–">May-</normalizedToken>
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16 August 1995, N. Hall, WAM-CALM Carnarvon Survey (WAM T41364DNA_Voucher_NCB_017); 1 ♂, Zuytdorp Nature Reserve, site ZU2 (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-27.261389">27°15'41"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="114.03001">114°01'48"E</geoCoordinate>
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, wet pitfall trap, 18
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<normalizedToken originalValue="May–">May-</normalizedToken>
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17 August 1995, N. Hall, WAM-CALM Carnarvon Survey (WAM T41788); 1 ♀, creek just N. of Canna (Irwin River system) (IBRA_AVW),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.9">28°54'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="115.86667">115°52'E</geoCoordinate>
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, 5 October 1953, B.Y. Main (WAM T144780); 1 ♀, 2.3 miles S. of Ebano Creek in Woolaga Creek (IBRA_AVW),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-29.2">29°12'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="115.65">115°39'E</geoCoordinate>
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, 29 May 1954, B.Y. Main (WAM T144783); 1 juvenile, same data (WAM T144784); 1 juvenile, Galena, 2 miles N. of Murchison River on North-west Coastal Highway (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-27.8">27°48'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.7">114°42'E</geoCoordinate>
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, 9 July 1954, B.Y. Main (WAM T144788); 1 ♂, Geraldton (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.783333">28°47'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.61667">114°37'E</geoCoordinate>
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, 1943, E.G. Osborne (AMS KS17388); 1 ♀, same data (AMS KS6393); 1 ♀, same locality data except
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.766666">28°46'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.61667">114°37'E</geoCoordinate>
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, 16 February 1983, A. Mollan (WAM T27120); 1 ♂, Geraldton, Minnenooka (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.816668">28°49'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.9">114°54'E</geoCoordinate>
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, 19 August 1971, J. Bryo (WAM T27119); 1 ♀, Kalbarri National Park, George Grey Drive, 18 km (by road) S. of Kalbarri (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="15" value="-27.856112">27°51'22"S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="114.15056">114°09'02"E</geoCoordinate>
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, dug from burrow, 17 August 2016, M.S. Harvey, M.E. Blosfelds (WAM T141118DNA_Voucher_NCB_006); 1 ♀, 27 km S. of Kalbarri (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-27.95">27°57'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.166664">114°10'E</geoCoordinate>
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, 23 September 1989, D. Mead-Hunter (WAM T21810); 1 ♀, 18 miles S. of Mullewa on Mingenew Road (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.8">28°48'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="115.51667">115°31'E</geoCoordinate>
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, 15 July 1955, B.Y. Main (WAM T144835); 1 ♀, same data (WAM T144836); 1 ♀, N. side of Murchison River (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-27.833334">27°50'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.683334">114°41'E</geoCoordinate>
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, 9 July 1954, B.Y. Main (WAM T144817); 1 ♀, same data (WAM T144789); 1 ♂, Northampton (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.35">28°21'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.63333">114°38'E</geoCoordinate>
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, hand collected under sheet of iron, 1 January 1982, Dr Allan (WAM T27123); 1 ♀, Sunset Beach (IBRA_GES),
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<geoCoordinate direction="south" orientation="latitude" precision="925" value="-28.716667">28°43'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="925" value="114.61667">114°37'E</geoCoordinate>
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, 9 September 1997, T. Burrows (WAM T44355).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="etymology">
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<paragraph pageId="21" pageNumber="22">
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<pageBreakToken pageId="21" pageNumber="22" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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The specific epithet is derived from the Latin arenaceus (adjective: 'of
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<normalizedToken originalValue="sand’">sand'</normalizedToken>
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,
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<normalizedToken originalValue="‘sandy’">'sandy'</normalizedToken>
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; see Brown 1956), in reference to the near-coastal sandy habitats occupied by this species in the Geraldton Sandplains bioregion.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="diagnosis">
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<paragraph pageId="21" pageNumber="22">Diagnosis.</paragraph>
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<paragraph pageId="21" pageNumber="22">
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<taxonomicName class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma arenaceum" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arenaceum">Idiosoma arenaceum</taxonomicName>
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is one of seven highly autapomorphic species in the polyphyletic 'sigillate
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<normalizedToken originalValue="complex’">complex'</normalizedToken>
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(Fig. 25); members of this complex can be distinguished from all other species in the nigrum-group from south-western Australia (i.e.,
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<taxonomicName lsidName="I. formosum" pageId="21" pageNumber="22" rank="species" species="formosum">I. formosum</taxonomicName>
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,
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<taxonomicName lsidName="I. gardneri" pageId="21" pageNumber="22" rank="species" species="gardneri">I. gardneri</taxonomicName>
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,
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<taxonomicName lsidName="I. gutharuka" pageId="21" pageNumber="22" rank="species" species="gutharuka">I. gutharuka</taxonomicName>
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,
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<taxonomicName lsidName="I. incomptum" pageId="21" pageNumber="22" rank="species" species="incomptum">I. incomptum</taxonomicName>
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,
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<taxonomicName lsidName="I. intermedium" pageId="21" pageNumber="22" rank="species" species="intermedium">I. intermedium</taxonomicName>
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,
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<taxonomicName lsidName="I. jarrah" pageId="21" pageNumber="22" rank="species" species="jarrah">I. jarrah</taxonomicName>
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,
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<taxonomicName lsidName="I. mcclementsorum" pageId="21" pageNumber="22" rank="species" species="mcclementsorum">I. mcclementsorum</taxonomicName>
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,
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<taxonomicName lsidName="I. mcnamarai" pageId="21" pageNumber="22" rank="species" species="mcnamarai">I. mcnamarai</taxonomicName>
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and
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<taxonomicName lsidName="I. sigillatum" pageId="21" pageNumber="22" rank="species" species="sigillatum">I. sigillatum</taxonomicName>
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) by the presence of well-defined lateral sclerotic strips on the male abdomen (e.g., Figs 32, 63, 256), and by the very heavily sclerotised, leathery,
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<normalizedToken originalValue="‘shield-like’">'shield-like'</normalizedToken>
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morphology of the female abdomen (e.g., Figs 1-3, 9-12, 52, 74, 96). Males and females of
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<taxonomicName lsidName="I. arenaceum" pageId="21" pageNumber="22" rank="species" species="arenaceum">I. arenaceum</taxonomicName>
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can be further distinguished from those of all other known 'sigillate
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<normalizedToken originalValue="complex’">complex'</normalizedToken>
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species (i.e.,
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<taxonomicName lsidName="I. clypeatum" pageId="21" pageNumber="22" rank="species" species="clypeatum">I. clypeatum</taxonomicName>
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,
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<taxonomicName lsidName="I. dandaragan" pageId="21" pageNumber="22" rank="species" species="dandaragan">I. dandaragan</taxonomicName>
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,
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<taxonomicName lsidName="I. kopejtkaorum" pageId="21" pageNumber="22" rank="species" species="kopejtkaorum">I. kopejtkaorum</taxonomicName>
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,
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<taxonomicName lsidName="I. kwongan" pageId="21" pageNumber="22" rank="species" species="kwongan">I. kwongan</taxonomicName>
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,
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<taxonomicName lsidName="I. nigrum" pageId="21" pageNumber="22" rank="species" species="nigrum">I. nigrum</taxonomicName>
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and
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<taxonomicName lsidName="I. schoknechtorum" pageId="21" pageNumber="22" rank="species" species="schoknechtorum">I. schoknechtorum</taxonomicName>
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) by the shape of the SP4 sclerites, which are longitudinally elongate-oval (Figs 63, 74, Key panes 7.1, 21.1) [NB. females of
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<taxonomicName lsidName="I. kwongan" pageId="21" pageNumber="22" rank="species" species="kwongan">I. kwongan</taxonomicName>
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are unknown].
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</paragraph>
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<paragraph pageId="21" pageNumber="22">
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This species can also be distinguished from
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<taxonomicName lsidName="I. corrugatum" pageId="21" pageNumber="22" rank="species" species="corrugatum">I. corrugatum</taxonomicName>
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(from the Eyre Peninsula of South Australia) by the shape of the prolateral clasping spurs on the male tibia I, which are oriented longitudinally (Fig. 65; cf. Fig. 109), and by the shape of the female eye group, which is broadly trapezoidal (Fig. 73; cf. Fig. 117).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="21" pageNumber="22" type="description">
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<paragraph pageId="21" pageNumber="22">Description (male holotype).</paragraph>
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<paragraph pageId="21" pageNumber="22">
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Total length 19.7. Carapace 8.0 long, 5.6 wide. Abdomen 8.4 long, 6.4 wide. Carapace (Fig. 57) dark chocolate-brown, with darker ocular region; lateral margins with uniformly-spaced fringe of porrect black setae; fovea procurved. Eye group (Fig. 60) trapezoidal (anterior eye row strongly procurved), 0.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide,
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<normalizedToken originalValue="PLE–PLE/ALE–ALE">PLE-PLE/ALE-ALE</normalizedToken>
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ratio 2.0; ALE almost contiguous; AME separated by less than their own diameter; PME separated by 2.8
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<normalizedToken originalValue="×">x</normalizedToken>
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their own diameter; PME and PLE separated by slightly more than diameter of PME, PME positioned slightly posterior to level of PLE. Maxillae with field of small cuspules confined to inner corner; labium without cuspules. Abdomen (Figs 58, 63) broadly oval, light beige-brown in dorsal view with lateral sclerotic strips, dorso-lateral corrugations and scattered dorsal sclerotic spots. Dorsal surface of abdomen (Fig. 58) more heavily setose anteriorly, with assortment of stiff, porrect black setae, each with slightly raised, dark brown sclerotic base. Posterior abdomen strongly sigillate (Figs 58, 63); SP2 sclerites small oval spots; SP3 sclerites very large and circular; SP4 sclerites longitudinally elongate-oval; SP5 obscured. Legs (Figs 64-66) variable shades of dark brown, with light scopulae on tarsi
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<normalizedToken originalValue="I–II">I-II</normalizedToken>
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; distal tibia I with pair of large prolateral clasping spurs oriented longitudinally. Leg I: femur 7.2; patella 3.5; tibia 4.7; metatarsus 5.0; tarsus 3.4; total 23.8. Leg I
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<normalizedToken originalValue="femur–tarsus">femur-tarsus</normalizedToken>
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/carapace length ratio 3.0. Pedipalpal tibia (Figs 67-69) 2.5
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<normalizedToken originalValue="×">x</normalizedToken>
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longer than wide; RTA burr-like, with conical basal protuberance and field of retroventral spinules; digital process porrect, unmodified. Cymbium (Figs 67-69) setose, with field of spinules disto-dorsally. Embolus (Figs 67-69) broadly twisted and sharply tapering distally, with prominent longitudinal flange and triangular (sub-distal) embolic apophysis.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22" type="description">
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<paragraph pageId="21" pageNumber="22">Description (female AMS KS6393).</paragraph>
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<paragraph lastPageId="22" lastPageNumber="23" pageId="21" pageNumber="22">
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Total length 24.8. Carapace 10.9 long, 8.1 wide. Abdomen 11.6 long, 10.8 wide. Carapace (Fig. 70) dark tan and chocolate-
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<pageBreakToken pageId="22" pageNumber="23" start="start">brown</pageBreakToken>
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, with darker ocular region; fovea procurved. Eye group (Fig. 73) trapezoidal (anterior eye row strongly procurved), 0.6
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide,
|
||
<normalizedToken originalValue="PLE–PLE/ALE–ALE">PLE-PLE/ALE-ALE</normalizedToken>
|
||
ratio 2.6; ALE almost contiguous; AME separated by approximately their own diameter; PME separated by 5.0
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
their own diameter; PME and PLE separated by more than diameter of PME, PME positioned in line with level of PLE. Maxillae with field of cuspules confined to inner corner (Fig. 75); labium without cuspules. Abdomen (Figs 71, 74) dark maroon-brown, corrugate and highly sclerotised, with leathery appearance typical of those species in the 'sigillate
|
||
<normalizedToken originalValue="complex’">complex'</normalizedToken>
|
||
(see Fig. 25). Posterior face of abdomen (Fig. 74, Key pane 21.1) with truncate
|
||
<normalizedToken originalValue="‘shield-like’">'shield-like'</normalizedToken>
|
||
morphology; SP3 sclerites very large and circular; SP4 sclerites longitudinally elongate-oval; SP5 obscured by thickened cuticle. Legs (Figs 76-77) variable shades of dark tan; scopulae present on tarsi and metatarsi
|
||
<normalizedToken originalValue="I–II">I-II</normalizedToken>
|
||
; tibia I with one stout pro-distal macroseta and row of five longer retroventral macrosetae; metatarsus I with six stout macrosetae (at least one broken off at base); tarsus I with distal cluster of short macrosetae. Leg I: femur 6.8; patella 4.3; tibia 3.9; metatarsus 3.3; tarsus 2.3; total 20.7. Leg I
|
||
<normalizedToken originalValue="femur–tarsus">femur-tarsus</normalizedToken>
|
||
/carapace length ratio 1.9. Pedipalp dark tan, spinose on tibia and tarsus, with thick tarsal scopula. Genitalia (Fig. 78) with pair of short, subtriangular spermathecae, each bearing dense field of glandular vesicles distally and more sparsely distributed glandular field sub-distally.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="22" pageNumber="23" type="distribution">
|
||
<paragraph pageId="22" pageNumber="23">Distribution and remarks.</paragraph>
|
||
<paragraph pageId="22" pageNumber="23">
|
||
<taxonomicName class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma arenaceum" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="arenaceum">Idiosoma arenaceum</taxonomicName>
|
||
(formerly known by WAM identification code
|
||
<normalizedToken originalValue="‘MYG478’">'MYG478'</normalizedToken>
|
||
) (Figs 11-12) has a moderately widespread distribution in the Geraldton Sandplains and far northern Wheatbelt bioregions of south-western Western Australia, from near Yandanooka, Canna, and Geraldton north to Zuytdorp (Fig. 374). It is closely related to the three other 'sigillate
|
||
<normalizedToken originalValue="complex’">complex'</normalizedToken>
|
||
species in the northern clypeatum-clade:
|
||
<taxonomicName lsidName="I. clypeatum" pageId="22" pageNumber="23" rank="species" species="clypeatum">I. clypeatum</taxonomicName>
|
||
,
|
||
<taxonomicName lsidName="I. kopejtkaorum" pageId="22" pageNumber="23" rank="species" species="kopejtkaorum">I. kopejtkaorum</taxonomicName>
|
||
, and
|
||
<taxonomicName lsidName="I. kwongan" pageId="22" pageNumber="23" rank="species" species="kwongan">I. kwongan</taxonomicName>
|
||
(Fig. 25). Burrows are adorned with a
|
||
<normalizedToken originalValue="‘moustache-like’">'moustache-like'</normalizedToken>
|
||
arrangement of twig-lines, sometimes under
|
||
<taxonomicName class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Casuarina" order="Fagales" pageId="22" pageNumber="23" phylum="Tracheophyta" rank="genus">Casuarina</taxonomicName>
|
||
(Figs 19, 20), and male specimens have been collected wandering in search of females in late autumn and winter, with an outlying record from January.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="22" pageNumber="23" type="conservation assessment">
|
||
<paragraph pageId="22" pageNumber="23">Conservation assessment.</paragraph>
|
||
<paragraph pageId="22" pageNumber="23">
|
||
<taxonomicName class="Arachnida" family="Idiopidae" genus="Idiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Idiosoma arenaceum" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="arenaceum">Idiosoma arenaceum</taxonomicName>
|
||
has a known extent of occurrence (EOO) of nearly 12,000 km2 [11,939 km2; with coastline as western margin], although this value is possibly an underestimate due to fairly limited survey effort. The area of occupancy within that range is similarly difficult to estimate, although is likely to be small as a proportion of EOO due to the scale of land clearing throughout most of its range. We do not currently consider this species to be of conservation concern, although further close assessment under both Criteria A and B is warranted in the future.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |