treatments-xml/data/E2/63/D9/E263D96D7DC95618A258F0EE34C194EE.xml

<document id="62D7003938D14FF1D2CDC9F58B4DFE4E" ID-DOI="10.3897/evolsyst.8.126393" ID-ZooBank="973DECB5-340A-41CE-9AEE-B968664489B2" ID-publisher-id="126393" URI-arpha="D96A320E-35D2-5E32-8DA0-4988C27D745A" URI-zoobank="http://zoobank.org/973DECB5-340A-41CE-9AEE-B968664489B2" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" article-type="research-article" checkinTime="1726056890737" checkinUser="pensoft" docAuthor="Costa, Wilson J. E. M., Feltrin, Caio R. M., Mattos, José Leonardo O. &amp; Katz, Axel M." docDate="2024" docId="E263D96D7DC95618A258F0EE34C194EE" docLanguage="en" docName="Evolutionary Systematics 8(2): 199-218" docOrigin="Evolutionary Systematics 8 (2)" docSource="https://evolsyst.pensoft.net/article/126393/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Cambeva atrobrunnea Costa, Feltrin &amp; Katz 2024, sp. nov." docType="treatment" docUuid="A2FF0368-0311-4739-9CB2-415343090897" docUuidSource="ZooBank" docVersion="3" dtd-version="3.0" lastPageNumber="218" masterDocId="D96A320E35D25E328DA04988C27D745A" masterDocTitle="Top mountain areas of subtropical southern Brazil sheltering four new small-ranged catfishes (Siluriformes, Trichomycteridae): relationships and taxonomy" masterLastPageNumber="218" masterPageNumber="199" pageNumber="199" updateTime="1726068310652" updateUser="admin">
<mods:mods id="9C6D09DB44D049F60C1C2C23127239B9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="217C69251E1D9FFA8F5B486F26185FA6">
<mods:title id="CDF298FC24971B7E596446E8FA1011D5">Top mountain areas of subtropical southern Brazil sheltering four new small-ranged catfishes (Siluriformes, Trichomycteridae): relationships and taxonomy</mods:title>
</mods:titleInfo>
<mods:name id="7C694B715A5C9F86830C92930CC54783" type="personal">
<mods:role id="DD47278285B018DDF3E2E798266085D6">
<mods:roleTerm id="3CDED7AF7052CBB65EE38B21260AB6FD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F604B3625FB01A413EDF5C70BDDBAA44">Costa, Wilson J. E. M.</mods:namePart>
<mods:nameIdentifier id="A124D2511DAC06F1BE98081BFE50A277" type="ORCID">0000-0002-0428-638X</mods:nameIdentifier>
<mods:affiliation id="69DF466F6DDC848536D927701A4CDB41">Laboratory of Systematics and Evolution of Teleost Fishes, Institute of Biology, Federal University of Rio de Janeiro, Caixa Postal 68049, CEP 21941 - 971, Rio de Janeiro, Brazil</mods:affiliation>
</mods:name>
<mods:name id="57958E0B97891D1F33017F781D059978" type="personal">
<mods:role id="BF8D0D28C0CB98E9BAD9E41363642D75">
<mods:roleTerm id="8987824733821420F6987D268AAE49EE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="69F5F8144915638220764CDC2E8D34F0">Feltrin, Caio R. M.</mods:namePart>
<mods:affiliation id="B71AAAB6A634BD701D34DF6140BDA3EC">Laboratory of Systematics and Evolution of Teleost Fishes, Institute of Biology, Federal University of Rio de Janeiro, Caixa Postal 68049, CEP 21941 - 971, Rio de Janeiro, Brazil</mods:affiliation>
</mods:name>
<mods:name id="08D8B4BCA4015D530375FDB5FD0386FE" type="personal">
<mods:role id="64ED1CE440593DFF4FD5165D7C2CA2D7">
<mods:roleTerm id="53D60B2CB338172B820ED027B1329BF2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DCB480F1D356814465A1E4442BDA0109">Mattos, José Leonardo O.</mods:namePart>
<mods:affiliation id="10D2ED14C880822456D6EE605452F5E1">Laboratory of Systematics and Evolution of Teleost Fishes, Institute of Biology, Federal University of Rio de Janeiro, Caixa Postal 68049, CEP 21941 - 971, Rio de Janeiro, Brazil</mods:affiliation>
</mods:name>
<mods:name id="81B25A562659CC7B6AFF5A8CF3B05AEB" type="personal">
<mods:role id="B6C3C3766E73AAE78152FA13C552B156">
<mods:roleTerm id="7A1C4DBA24F6E71DE7D5119C7D868680">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9DE6C7E81742FC956E20E42529FAE4F6">Katz, Axel M.</mods:namePart>
<mods:nameIdentifier id="AE56D0E530468D8E75F0D7C410CF2461" type="ORCID">0000-0002-2933-7163</mods:nameIdentifier>
<mods:affiliation id="DE4F76703A756074CFD652333B5E7518">Laboratory of Systematics and Evolution of Teleost Fishes, Institute of Biology, Federal University of Rio de Janeiro, Caixa Postal 68049, CEP 21941 - 971, Rio de Janeiro, Brazil</mods:affiliation>
</mods:name>
<mods:typeOfResource id="5F1FF1BF3B9122F3F8865491AA04AF34">text</mods:typeOfResource>
<mods:relatedItem id="3DE664EBAAB8CF3D9978AEECC6874056" type="host">
<mods:titleInfo id="D39B21699D73D74C05EB5D6D0ADC8E90">
<mods:title id="CEC20D102CA12933CB934240FA71CA09">Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part id="E0413061BE8CB6B85458F8967C3FF25C">
<mods:date id="1EEA5604978369D90506EE595C3BC7C5">2024</mods:date>
<mods:detail id="47ADE8C60EB456EC3AE34B683AFE7B9C" type="pubDate">
<mods:number id="1A7BEA7F093F8C7108646E7A7AA836BB">2024-09-11</mods:number>
</mods:detail>
<mods:detail id="365AF3A7E9A917E69799F0FEC85AF7B5" type="volume">
<mods:number id="0131A88BF620B61AA6FD349EB9DEDE1B">8</mods:number>
</mods:detail>
<mods:detail id="5A3DF66754512CCDABB77623973EB323" type="issue">
<mods:number id="1053B7D21DB17AB3C28E42FCD06C6367">2</mods:number>
</mods:detail>
<mods:extent id="5741153CBB73E209BF0BF799CF4828A6" unit="page">
<mods:start id="C2041381AD9F516956949E56CE9155B1">199</mods:start>
<mods:end id="99AA47B61CF8D5889B791FB51E9911E2">218</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C2FC94C31BA08DFCBA3F361BC8CBD4DC">journal article</mods:classification>
<mods:identifier id="C7632C6844B4164631A0004DD3591E6E" type="DOI">10.3897/evolsyst.8.126393</mods:identifier>
<mods:identifier id="FF9ACAC97F5273B78C0E5F637EA2B9C5" type="ZooBank">973DECB5-340A-41CE-9AEE-B968664489B2</mods:identifier>
</mods:mods>
<treatment id="E263D96D7DC95618A258F0EE34C194EE" ID-arpha="E263D96D-7DC9-5618-A258-F0EE34C194EE" ID-zoobank="https://zoobank.org/A2FF0368-0311-4739-9CB2-415343090897" LSID="urn:lsid:zoobank.org:act:A2FF0368-0311-4739-9CB2-415343090897" LSID-ZBK="urn:lsid:zoobank.org:act:A2FF0368-0311-4739-9CB2-415343090897" httpUri="http://treatment.plazi.org/id/E263D96D7DC95618A258F0EE34C194EE">
<subSubSection id="FA614D9DC0AB1BE4D6D6892FDB35B163" type="nomenclature">
<paragraph id="2C37C06EC3A27576E12B27FB0A79CCC4">
<taxonomicName id="5799CFFB0698D481BABC0D486E33F4F8" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea" status="sp. nov.">
<emphasis id="1F526969732FC328DE323A13FD7F9EDD" italics="true">Cambeva atrobrunnea</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
<taxonomicNameLabel id="5D59D924AFC45818EB059F3606A160FC" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="F6E4449DA3ACAEC1AA7425A98D096FAF" type="description">
<paragraph id="AC5E54B82D5669DB7B6D0789478EC120">
<figureCitation id="1FCF7F9FE6B9EC822C5DAA72B129914A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Osteological structures of: A – C. Cambeva galactica Costa, Feltrin &amp; Katz, sp. nov.; D – F. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov.; G – I. Cambeva luteoreticulata Costa, Feltrin &amp; Katz, sp. nov., and J – L. Cambeva rotundipinna Costa, Feltrin &amp; Katz, sp. nov. A, D, G, J. Mesethmoidal region and adjacent structures, left and middle portions, dorsal view; B, E, H, K. Left jaw suspensorium and opercular series, lateral view. C, F, I, L. Parurohyal, ventral view. Abbreviation of structure indicated by arrow: aes, anteromedial expansion of sesamoid supraorbital. Larger stippling represents cartilaginous areas." httpUri="https://binary.pensoft.net/fig/1131386">Figs 4 D – F</figureCitation>
, 
<figureCitation id="915A4EBD9B106C1B6EBFD63EE4BB0524" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov., holotype, UFRJ 14066, 70.1 mm SL. A. Lateral view; B. Dorsal view; C. Ventral view." httpUri="https://binary.pensoft.net/fig/1131388">6</figureCitation>
, 
<figureCitation id="0BFD4AC6A5228DEED6BED558F3E2082E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov., paratype, UFRJ 13821, 33.4 mm SL. A. Lateral view; B. Dorsal view; C. Ventral view." httpUri="https://binary.pensoft.net/fig/1131389">7</figureCitation>
, 
<tableCitation id="B2F1EFE905091A4C440C038D8E76DCB7" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometric data of Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov." httpUri="http://table.plazi.org/id/D596199D62DCD4AF7E683F5341BA35E7" tableUuid="D596199D62DCD4AF7E683F5341BA35E7">Table 4</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EUAAI" type="type material">
<paragraph id="CBE6C9131BF072A00E7604240F3719B1">
<heading id="FB511FF1AED9DC487F1A3523A74B8193" reason="title">Type material.</heading>
</paragraph>
<paragraph id="7D75E130CEE8E55E01C0EFE0419C34EC">
<materialsCitation id="CB39A46AF6039F55603D5D5E789EB41E" collectingDate="2023-06-29" collectionCode="R" collectorName="C. R. M. Feltrin" country="Brazil" county="Timbo Grande Municipality" elevation="970" latitude="-26.578056" location="stream tributary of Rio Timbo, Rio Iguacu drainage, Rio Parana basin" longLatPrecision="20" longitude="-50.674446" municipality="Timbo Grande Municipality" specimenCode="UFRJ 14066" specimenCount="1" stateProvince="Santa Catarina State" typeStatus="holotype">
<emphasis id="49BDE8FD8EDE39CAA1CFD027C2BF1C41" bold="true">
<emphasis id="6301577505C2F745E5524BFEBF192AE2" bold="true" italics="true">
<typeStatus id="769614D569BA3DDCAE91F05EE94B5EC4" type="holotype">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="AC7B63B2C09C2537C5166075B0E2085E" name="Brazil">Brazil</collectingCountry>
• 
<quantity id="4AD3339A2BF1B65C9425F94D5172F365" metricMagnitude="-2" metricUnit="m" metricValue="7.01" unit="mm" value="70.1">70.1 mm</quantity>
SL; 
<collectingRegion id="6A67ED2587B8D5AA6AE51CD524DBFA62" country="Brazil" name="Santa Catarina">Santa Catarina State</collectingRegion>
: 
<collectingMunicipality id="6737EB75215B3C1C19161EB04A876155">Timbó Grande Municipality</collectingMunicipality>
: 
<location id="F4EFA154D7042A876086CFD8B9D3A542" LSID="urn:lsid:plazi:treatment:E263D96D7DC95618A258F0EE34C194EE:F4EFA154D7042A876086CFD8B9D3A542" country="Brazil" county="Timbo Grande Municipality" latitude="-26.578056" longLatPrecision="20" longitude="-50.674446" municipality="Timbo Grande Municipality" name="stream tributary of Rio Timbo, Rio Iguacu drainage, Rio Parana basin" stateProvince="Santa Catarina State">stream tributary of Rio Timbó, Rio Iguaçu drainage, Rio Paraná basin</location>
; 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0ECBAI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[-50.674444,-26.578056]}">
<geoCoordinate id="51C4C7A74257C7459B169EBB5A22693B" degrees="26" direction="south" minutes="34" orientation="latitude" precision="15" seconds="41" value="-26.578056">26 ° 34 ' 41 &quot; S</geoCoordinate>
, 
<geoCoordinate id="80A76A405B5A6B44BDA5A512429F919A" degrees="50" direction="west" minutes="40" orientation="longitude" precision="15" seconds="28" value="-50.674446">50 ° 40 ' 28 &quot; W</geoCoordinate>
</named-content>
; about 
<elevation id="A9A32B581E8C7184CCEF81095482CE94" metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">
<quantity id="C3E7E8F1F7DEDAB0A6D9D0C4D22E1D59" metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">970 m</quantity>
asl
</elevation>
; 
<date id="29F4FEC39AF147991711C4CC871F137D" value="2023-06-29">
<collectingDate id="405BC82BFEDF2ECA3AF5E617812FB877" value="2023-06-29">29 Jun. 2023</collectingDate>
</date>
; 
<collectorName id="75B6B5F50FF07B473B2541A95B17B626">C. R. M. Feltrin</collectorName>
, leg; 
<specimenCode id="0B40240D89CF6EAAA2BD1CEE0B981D89">
<named-content id="EA3E8C4B5998F0B3034D36FC49CED565" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-rural-do-rio-de-janeiro" xlink_title="Instituto de Biologia, Universidade Federal do Rio de Janeiro">UFRJ</named-content>
14066
</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="099B10ED61DF9AFC4C6AC3BC2919E12A">
<materialsCitation id="352C08CDB241025DE54AF722B3A5B1B3" collectingDate="2023-06-29" collectionCode="R" collectorName="C. R. M. Feltrin" country="Brazil" county="Timbo Grande Municipality" elevation="970" latitude="-26.578056" location="Brazil" longLatPrecision="20" longitude="-50.674446" municipality="Rio Timbo" specimenCode="UFRJ 13821" specimenCount="7" stateProvince="Santa Catarina" typeStatus="paratype">
<emphasis id="4AAC8E3DA0FF387C3E27FACFAFD3EBC1" bold="true">
<emphasis id="2E62354B31A7BF21B48CA84E272A5CB8" bold="true" italics="true">
<typeStatus id="BA14C8C5A78EA282905B8D5B3D5F6F3D" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="CE9C27FEE113699591F9BECA1DA50FA0" name="Brazil">Brazil</collectingCountry>
• 
<specimenCount id="7C958E5DD7D792B57028A13679F473C1" count="7" type="generic">7 ex.</specimenCount>
, 
<quantity id="5803E018F134D2CFD20DF1061EC11217" metricMagnitude="-2" metricUnit="m" metricValue="4.834999999999999" metricValueMax="7.01" metricValueMin="2.66" unit="mm" value="48.349999999999994" valueMax="70.1" valueMin="26.6">26.6–70.1 mm</quantity>
SL; same data as holotype; 
<specimenCode id="60114FB964AF32C2C4BE5E787D99AA82">
<named-content id="DBAFBC14B43B8A0339AF3E2E626E4C9E" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-rural-do-rio-de-janeiro" xlink_title="Instituto de Biologia, Universidade Federal do Rio de Janeiro">UFRJ</named-content>
13821
</specimenCode>
</materialsCitation>
. • 
<materialsCitation id="077FB680D3E49B95199EC457E9BAF738" collectingDate="2023-06-29" collectionCode="R" collectorName="C. R. M. Feltrin" country="Brazil" county="Timbo Grande Municipality" elevation="970" latitude="-26.578056" location="Rio Iguacu" longLatPrecision="20" longitude="-50.674446" municipality="Rio Timbo" specimenCode="UFRJ 14067" specimenCount="2" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="387B5C7392351CDF89C6C122D46D279D" count="2" type="generic">2 ex.</specimenCount>
(C &amp; S), 
<quantity id="801A9293FAEF2F09561D206EC76FD4C8" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" metricValueMax="4.42" metricValueMin="2.7399999999999998" unit="mm" value="35.8" valueMax="44.2" valueMin="27.4">27.4–44.2 mm</quantity>
SL; same data as holotype; 
<specimenCode id="D60C65C6DAFFA83B7EB4E41F88903EB2">
<named-content id="12B94441EE77B3E9443A63FE64569700" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-rural-do-rio-de-janeiro" xlink_title="Instituto de Biologia, Universidade Federal do Rio de Janeiro">UFRJ</named-content>
14067
</specimenCode>
</materialsCitation>
. • 
<materialsCitation id="4FC98D5372F3D8C8C0449F1A69BDBD0A" collectingDate="2023-03-31" collectionCode="R" collectorName="C. R. M. Feltrin" country="Brazil" county="Timbo Grande Municipality" elevation="970" latitude="-26.578056" location="Rio Iguacu" longLatPrecision="20" longitude="-50.674446" municipality="Rio Timbo" specimenCode="UFRJ 13553" specimenCount="2" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="E1FCF4BC0B6D0D2621A5FBE68B11B9FB" count="2" type="generic">2 ex.</specimenCount>
, 
<quantity id="20FF110D7C1C4EA73719DCF1C65FDFAD" metricMagnitude="-2" metricUnit="m" metricValue="3.8349999999999995" metricValueMax="4.68" metricValueMin="2.9899999999999998" unit="mm" value="38.349999999999994" valueMax="46.8" valueMin="29.9">29.9–46.8 mm</quantity>
SL. same locality and collector as holotype; 
<date id="12656554CE49111FB3981090527D5389" value="2023-03-31">
<collectingDate id="66CE5E187E5B0346DD105463688F5E33" value="2023-03-31">31 Mar. 2023</collectingDate>
</date>
; 
<specimenCode id="7FDE0CC74B472657FB26F8934BAB8177">
<named-content id="24FDA8A7324B016EF9A8CE0BC991EEC6" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-rural-do-rio-de-janeiro" xlink_title="Instituto de Biologia, Universidade Federal do Rio de Janeiro">UFRJ</named-content>
13553
</specimenCode>
</materialsCitation>
. • 
<materialsCitation id="8613A7FDEA8B4AAE395C67FFE2D1588D" collectingDate="2023-03-31" collectionCode="R" collectorName="C. R. M. Feltrin" country="Brazil" county="Timbo Grande Municipality" elevation="970" latitude="-26.578056" location="Rio Iguacu" longLatPrecision="20" longitude="-50.674446" municipality="Rio Timbo" specimenCode="UFRJ 13554" specimenCount="3" stateProvince="Santa Catarina" typeStatus="paratype">
<specimenCount id="10828171FEAA55ECC321F5654213E327" count="3" type="generic">3 ex.</specimenCount>
(DNA), 
<quantity id="A70DA46769A8B802A86B9EF4B2C832FF" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" metricValueMax="4.42" metricValueMin="2.7399999999999998" unit="mm" value="35.8" valueMax="44.2" valueMin="27.4">27.4–44.2 mm</quantity>
SL; same locality and collector as holotype; 
<date id="4D73320DCD035707850EA968E589C332" value="2023-03-31">
<collectingDate id="32B06E8ADA96D6BFE5B5D181609D21FC" value="2023-03-31">31 Mar. 2023</collectingDate>
</date>
; 
<specimenCode id="22510D57A1E348156D0B3839E2AE8660">
<named-content id="7CFEEF2F25A923693FE2E167C26ADE3D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-rural-do-rio-de-janeiro" xlink_title="Instituto de Biologia, Universidade Federal do Rio de Janeiro">UFRJ</named-content>
13554
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECCAI" type="diagnosis">
<paragraph id="EDCFE0BB1CB7406C41B6A4CCC798AC57">
<heading id="7BC0055AF940CDA6CDDCD16B26272FC9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="D5D0B0746B090A3A80BA1766FDB73C7C">
<taxonomicName id="01475C35BD6FB1191AAE302493C95556" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea">
<emphasis id="426A10DD5ACA040210BCE633077DE11C" italics="true">Cambeva atrobrunnea</emphasis>
</taxonomicName>
is distinguished from all other congeners by having the two posterior-most dorsal and ventral procurrent caudal-fin rays segmented (vs. only the posterior-most ray segmented). 
<taxonomicName id="2495293B9AD07EDFA333E68293781E48" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea">
<emphasis id="B9697E2716F949A5274276B70571A412" italics="true">Cambeva atrobrunnea</emphasis>
</taxonomicName>
is also distinguished from the two other species of western RISE, 
<taxonomicName id="92905CE580C7006374AE4C7EA77731C1" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="luteoreticulata" status="sp. nov.">
<emphasis id="E40DD326CFF476717B1C4CA7D974FA4E" italics="true">Cambeva luteoreticulata</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="CE9F16445356A06AE6B99FA7A64AFE4D" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="54E968BF2097B1780357C690D1B04B67" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="rotundipinna" status="sp. nov.">
<emphasis id="BBB605EB86CE54E79C6D354B1FE67E96" italics="true">Cambeva rotundipinna</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="0DB031B0978929BC284CDCA11789AAA8" rank="species">sp. nov.</taxonomicNameLabel>
, by having subtruncate caudal fin (vs. rounded), fewer interopercular odontodes (20–22 vs. 29–34), and specimens below about 
<quantity id="571A567A6492B3BEC0A5264FA02EDFF4" metricMagnitude="-2" metricUnit="m" metricValue="4.0" unit="mm" value="40.0">40 mm</quantity>
SL having flank light grey with small black dots that are arranged in irregular rows, coalesced on the anterior portion of the longitudinal midline to form a stripe (Fig. 
<figureCitation id="A142CABA14C257322908771A8E13F6DF" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov., paratype, UFRJ 13821, 33.4 mm SL. A. Lateral view; B. Dorsal view; C. Ventral view." httpUri="https://binary.pensoft.net/fig/1131389">7 A</figureCitation>
; vs. pale yellow with large, irregularly shaped dark brown to black blotches, sometimes forming longitudinal stripes in the area between dorsum and flank in 
<taxonomicName id="BF070C98B71FA092CB9E8691CEDF67B3" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="luteoreticulata" status="sp. nov.">
<emphasis id="586C87B7D82AC6667790A04E194257F8" italics="true">C. luteoreticulata</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="92B6468C394265BC2D51158231BC033B" rank="species">sp. nov.</taxonomicNameLabel>
, and light brownish yellow with small black dots irregularly arranged in 
<taxonomicName id="590A745889F95B1401EBFDD1AE105115" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="rotundipinna" status="sp. nov.">
<emphasis id="9E38B8A307B7A39D709457D0BC630B07" italics="true">C. rotundipinna</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="76766F22D92D4AEED727FD07A8A0155A" rank="species">sp. nov.</taxonomicNameLabel>
(see descriptions below); from 
<taxonomicName id="7D0E26A2A2320F05422F84413B801810" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="luteoreticulata" status="sp. nov.">
<emphasis id="33AD1799F01BCC5B0E048F18181CB4E8" italics="true">C. luteoreticulata</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="856D1889846ED20475C70647AAFB196E" rank="species">sp. nov.</taxonomicNameLabel>
by specimens above 
<quantity id="CA910C8A9AD4A992644AB7890BB22E30" metricMagnitude="-2" metricUnit="m" metricValue="4.0" unit="mm" value="40.0">40 mm</quantity>
SL having the nasal barbel reaching area just anterior to opercle (vs. reaching area anterior to orbit), fewer ventral procurrent caudal-fin rays (13 or 14 vs. 15 or 16), and jaw teeth irregularly arranged (vs. arranged in three rows); and from 
<taxonomicName id="8533862A325C38FEB1A0BF5E197908AF" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="rotundipinna" status="sp. nov.">
<emphasis id="677EB6B20CB2A42B63D4CEBC017B1A1C" italics="true">C. rotundipinna</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="1CE63FC8F522943711BEFD4B578A2D37" rank="species">sp. nov.</taxonomicNameLabel>
by having more procurrent caudal-fin rays (21 dorsal and 13 or 14 ventral, vs. 15–17 and 10 or 11, respectively), and more opercular odontodes (12 vs. eight or nine). 
<taxonomicName id="A0376D3693E711543E5B41808AD3255F" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea">
<emphasis id="9BDFEE2285FD33091B9E8AA7722BF2F1" italics="true">Cambeva atrobrunnea</emphasis>
</taxonomicName>
is also distinguished from all the remaining congeners from the Rio Iguaçu drainage by the following combination of character states: seven pectoral-fin rays (vs. eight in 
<taxonomicName id="9A4003EC3905A40E76A749199527E6A3" authority="(de Pinna, 1992)" baseAuthorityName="de Pinna" baseAuthorityYear="1992" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="castroi">
<emphasis id="B5242B811B7E721211749A0A927C7018" italics="true">Cambeva castroi</emphasis>
(
<bibRefCitation id="7D27FE887DD0D24B39A1EF87D8344C2F" author="de Pinna MCC" journalOrPublisher="Ichthyological Exploration of Freshwaters" pagination="89-95" refId="B24" refString="de Pinna MCC (1992) Trichomycterus castroi, a new species of trichomycterid catfish from the Rio Iguacu of southeastern Brazil (Teleostei, Siluriformes). Ichthyological Exploration of Freshwaters 3: 89 - 95." title="Trichomycterus castroi, a new species of trichomycterid catfish from the Rio Iguaçu of southeastern Brazil (Teleostei, Siluriformes)." volume="3" year="1992">
de 
<emphasis id="B74DB95BB03CD1F80BE4F8D079AA56B7" italics="true">Pinna</emphasis>
, 1992
</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="1296FFCC22FA4A9D74706993AD4334FD" authority="Costa, Abilhoa, Dalcin &amp; Katz, 2022" authorityName="Costa, Abilhoa, Dalcin &amp; Katz" authorityYear="2022" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="melanoptera">
<emphasis id="8F6A97563BD2A54194B15A7DD29361BF" italics="true">Cambeva melanoptera</emphasis>
<bibRefCitation id="3D17CB01C42E662540D761ACFE98B034" DOI="10.1111/jfb.15071" author="Costa WJEM &amp; Abilhoa V &amp; Dalcin RH &amp; Katz AM" journalOrPublisher="Journal of Fish Biology" pagination="69-76" refId="B18" refString="Costa WJEM, Abilhoa V, Dalcin RH, Katz AM (2022) A new catfish species of the genus Cambeva (Siluriformes: Trichomycteridae) from the Rio Iguacu drainage, southern Brazil, with a remarkable unique colour pattern. Journal of Fish Biology 101: 69 - 76. https: // doi. org / 10.1111 / jfb. 15071" title="A new catfish species of the genus Cambeva (Siluriformes: Trichomycteridae) from the Rio Iguaçu drainage, southern Brazil, with a remarkable unique colour pattern." volume="101" year="2022">Costa, Abilhoa, Dalcin &amp; Katz, 2022</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="427B2F85815B2E7F7EE2D611439FC624" authority="(Wosiacki &amp; de Pinna, 2008)" baseAuthorityName="Wosiacki &amp; de Pinna" baseAuthorityYear="2008" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="crassicaudata">
<emphasis id="4053FE84F75849CD5D691953E372D699" italics="true">Cambeva crassicaudata</emphasis>
(Wosiacki &amp; de 
<emphasis id="E8F1E22EDF3079EB1324F01DC5DC8A12" italics="true">Pinna</emphasis>
, 2008)
</taxonomicName>
, 
<taxonomicName id="DA12C8C49DD9E57C47C287DD014E2523" authority="(Miranda Ribeiro, 1968)" authorityName="" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1968" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="stawiarski">
<emphasis id="ECD6392422AE5F70935B07FF6F5B76FF" italics="true">Cambeva stawiarski</emphasis>
(Miranda Ribeiro, 1968)
</taxonomicName>
, and six in 
<taxonomicName id="2ECD201D1D414F92E5F385988CFA9FCC" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="galactica">
<emphasis id="09FE9653E5C5EB801E87716D3F3D249A" italics="true">C. galactica</emphasis>
</taxonomicName>
, 
<taxonomicName id="5CDBFFF582AD9CCDEEBB3DD2D2E4E71E" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="naipi">
<emphasis id="5B82EFEE33EACC5F695B723175ED6062" italics="true">C. naipi</emphasis>
</taxonomicName>
, and 
<taxonomicName id="6604F75BED6B959E24DB918A8C50C130" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taroba">
<emphasis id="BD49ABA40634F288ED81A9F024BD5242" italics="true">C. taroba</emphasis>
</taxonomicName>
); absence of the anterior infraorbital (vs. presence in 
<taxonomicName id="CC6345BAACEE8D1E2DD3B9CCE9F3B2F4" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="galactica">
<emphasis id="77044E55F2A3B73F6675F3290055F1B1" italics="true">C. galactica</emphasis>
</taxonomicName>
, 
<taxonomicName id="47695461D3201E579AF23C2AAB9CFE4C" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="naipi">
<emphasis id="BEAB8C1C9F6EF63AF8974CA834EB16A9" italics="true">C. naipi</emphasis>
</taxonomicName>
, 
<taxonomicName id="70CB87D7CD22C28C7BA64283DD39FCC4" authority="(Wosiacki &amp; Garavello, 2004)" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="papillifera">
<emphasis id="AEDC736AE7F701AF61E5844A1A693877" italics="true">Cambeva papillifera</emphasis>
(
<bibRefCitation id="C46A2D8A95E0B659C6F7B0701074BE1C" author="Wosiacki WB &amp; Garavello JC" journalOrPublisher="Ichthyological Exploration of Freshwaters" pagination="1-16" refId="B61" refString="Wosiacki WB, Garavello JC (2004) Five new species of Trichomycterus from the Iguacu (rio Parana Basin), southern Brazil (Siluriformes: Trichomycteridae). Ichthyological Exploration of Freshwaters 15: 1 - 16." title="Five new species of Trichomycterus from the Iguaçu (rio Paraná Basin), southern Brazil (Siluriformes: Trichomycteridae)." volume="15" year="2004">Wosiacki &amp; Garavello, 2004</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="C2FBAF32B4DBAC04B6F28F6F99C8E708" authority="(Wosiacki &amp; Garavello, 2004)" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="plumbea">
<emphasis id="9264B3B28571C0B96A80887C5208AD41" italics="true">Cambeva plumbea</emphasis>
(
<bibRefCitation id="A75D4760CD2C54B0956412330EE27FED" author="Wosiacki WB &amp; Garavello JC" journalOrPublisher="Ichthyological Exploration of Freshwaters" pagination="1-16" refId="B61" refString="Wosiacki WB, Garavello JC (2004) Five new species of Trichomycterus from the Iguacu (rio Parana Basin), southern Brazil (Siluriformes: Trichomycteridae). Ichthyological Exploration of Freshwaters 15: 1 - 16." title="Five new species of Trichomycterus from the Iguaçu (rio Paraná Basin), southern Brazil (Siluriformes: Trichomycteridae)." volume="15" year="2004">Wosiacki &amp; Garavello, 2004</bibRefCitation>
)
</taxonomicName>
, and 
<taxonomicName id="AB5D9F9E9F4C8B01D9062A05F9BBF383" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taroba">
<emphasis id="AAC80F8F8DFED9C07EEC43A739C8D6EC" italics="true">C. taroba</emphasis>
</taxonomicName>
); posterior margin of the caudal fin slightly convex (vs. about straight in 
<taxonomicName id="6E148B6191E3198AE0F73517FFAEC405" baseAuthorityName="Wosiacki &amp; de Pinna" baseAuthorityYear="2008" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="crassicaudata">
<emphasis id="0F42705337898AFFA8825F5665E3A4D5" italics="true">C. crassicaudata</emphasis>
</taxonomicName>
, 
<taxonomicName id="74818807D22E092BB4C2BAE794BB7141" baseAuthorityName="Haseman" baseAuthorityYear="1911" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="davisi">
<emphasis id="CAFD04B01DF4B8C50DEF29AC7381425C" italics="true">C. davisi</emphasis>
</taxonomicName>
, 
<taxonomicName id="7CA512C9C431FFD123BFBC34AC9B618B" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="galactica">
<emphasis id="F1CF731FF374D55AECFBADD7959551FF" italics="true">C. galactica</emphasis>
</taxonomicName>
, 
<taxonomicName id="7B4339F59496550E0A68187C0B6B63AC" authorityName="Costa, Abilhoa, Dalcin &amp; Katz" authorityYear="2022" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="melanoptera">
<emphasis id="0C2CC67D61786A6B708BE76205430C5B" italics="true">C. melanoptera</emphasis>
</taxonomicName>
, 
<taxonomicName id="605543DDA2A5B263DF46D22A69D4A2C8" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="papillifera">
<emphasis id="5ED881C23E35607A2D1A3220F975F83C" italics="true">C. papillifera</emphasis>
</taxonomicName>
, 
<taxonomicName id="43E9218F2755F1C4083A77F9C06DB7E7" authority="dos Reis, Wosiacki, Ferrer, Donin &amp; da Graca, 2022" authorityName="dos Reis, Wosiacki, Ferrer, Donin &amp; da Graca" authorityYear="2022" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="piraquara">
<emphasis id="7D43DA25EACBD3B12113A20EF514D7B2" italics="true">Cambeva piraquara</emphasis>
dos Reis, Wosiacki, Ferrer, Donin &amp; da Graça, 2022
</taxonomicName>
, 
<taxonomicName id="0777F44411DD6E5D1724DE794BE96E2E" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="plumbea">
<emphasis id="7F77A61B783E3EC914D1415A1E70D966" italics="true">C. plumbea</emphasis>
</taxonomicName>
, 
<taxonomicName id="384BDA53B07322BB401FA475D82E167F" authority="(Wosiacki &amp; de Pinna, 2008)" baseAuthorityName="Wosiacki &amp; de Pinna" baseAuthorityYear="2008" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="igobi">
<emphasis id="2183C35EB548AC2592F2C1A063A4DD4E" italics="true">C. igobi</emphasis>
(Wosiacki &amp; de 
<emphasis id="04A807B5C5FF726D0265AE052C1B5A87" italics="true">Pinna</emphasis>
, 2008)
</taxonomicName>
; straight to slightly concave in 
<taxonomicName id="B4434A5EC694C54463068344AEC0BD07" authorityName="" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1968" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="stawiarski">
<emphasis id="8DB83BF8C3BD61B51F1F76F5CF356805" italics="true">C. stawiarski</emphasis>
</taxonomicName>
; bilobed in 
<taxonomicName id="9D1BF489D8943F862C3319DF78944014" baseAuthorityName="Wosiacki &amp; de Pinna" baseAuthorityYear="2008" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="crassicaudata">
<emphasis id="6E7B74376A6F0119A79B2346747212B7" italics="true">C. crassicaudata</emphasis>
</taxonomicName>
; and emarginate in 
<taxonomicName id="DEAB2181E4F490DAA9AEB02F403A81DF" authority="dos Reis, Ferrer &amp; Graca, 2021" authorityName="dos Reis, Ferrer &amp; Graca" authorityYear="2021" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="cauim">
<emphasis id="F96A2088F08A0C9E9E81E3071356EA28" italics="true">Cambeva cauim</emphasis>
<bibRefCitation id="2D766677732AA73210733BD01E75732B" DOI="10.1111/jfb.14204" author="dos Reis RB &amp; Ferrer J &amp; da Graça WJ" journalOrPublisher="Journal of Fish Biology" pagination="350-363" refId="B27" refString="dos Reis RB, Ferrer J, da Graca WJ (2021) A new species of Cambeva (Siluriformes, Trichomycteridae) from the Rio Iguacu basin, Parana state, Brazil and redescription of Cambeva stawiarski (Miranda Ribeiro 1968). Journal of Fish Biology 96: 350 - 363. https: // doi. org / 10.1111 / jfb. 14204" title="A new species of Cambeva (Siluriformes, Trichomycteridae) from the Rio Iguaçu basin, Paraná state, Brazil and redescription of Cambeva stawiarski (Miranda Ribeiro 1968)." volume="96" year="2021">dos Reis, Ferrer &amp; Graça, 2021</bibRefCitation>
</taxonomicName>
); absence of hypertrophied papillae on the ventral surface of the head (vs. presence in 
<taxonomicName id="69CB2BD0C8BC0EBF089D547337D35644" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="papillifera">
<emphasis id="6847C7E9E6FBBD467643C88844C7D356" italics="true">C. papillifera</emphasis>
</taxonomicName>
); absence of pectoral-fin filament (vs. presence a well-developed filament in 
<taxonomicName id="2EEC7F6903043FFA6E11768AF34A88B6" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taroba">
<emphasis id="1F096992F622285840893117C3B694C3" italics="true">C. taroba</emphasis>
</taxonomicName>
and a rudimentary filament in 
<taxonomicName id="C13D8CA1DA6887064A0099F08055DDB4" baseAuthorityName="Haseman" baseAuthorityYear="1911" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="davisi">
<emphasis id="7B7BF61823070709F7435B157D85C9D0" italics="true">C. davisi</emphasis>
</taxonomicName>
, 
<taxonomicName id="641F6148414F57F0B96D527E1915B789" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="galactica">
<emphasis id="AE06340EB1A6876D704C65AAED88874F" italics="true">C. galactica</emphasis>
</taxonomicName>
, and 
<taxonomicName id="BB1D203B19C36687BF78487AA0DA5CBC" authorityName="dos Reis, Wosiacki, Ferrer, Donin &amp; da Graca" authorityYear="2022" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="piraquara">
<emphasis id="8EEBCB65E94041B7505125EDAB1F4FA8" italics="true">C. piraquara</emphasis>
</taxonomicName>
); 12 opercular odontodes (vs. broad, with 17 or 
<quantity id="C89FD1A60048EF850C9DF361D8893982" metricMagnitude="-1" metricUnit="m" metricValue="4.572" unit="in" value="18.0">18 in</quantity>
<taxonomicName id="4C10C43D1E412A866D5C4FBD165BDB05" authority="(Wosiacki &amp; Garavello, 2004)" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="mboycy">
<emphasis id="3D3A1980F8AEB00221B675A4478274E9" italics="true">Cambeva mboycy</emphasis>
(
<bibRefCitation id="0FB18ACC9951620AC79057456D25430E" author="Wosiacki WB &amp; Garavello JC" journalOrPublisher="Ichthyological Exploration of Freshwaters" pagination="1-16" refId="B61" refString="Wosiacki WB, Garavello JC (2004) Five new species of Trichomycterus from the Iguacu (rio Parana Basin), southern Brazil (Siluriformes: Trichomycteridae). Ichthyological Exploration of Freshwaters 15: 1 - 16." title="Five new species of Trichomycterus from the Iguaçu (rio Paraná Basin), southern Brazil (Siluriformes: Trichomycteridae)." volume="15" year="2004">Wosiacki &amp; Garavello, 2004</bibRefCitation>
)
</taxonomicName>
; 15 or 
<quantity id="1389B214AEC9B0BFAE41C3CD4791CDDA" metricMagnitude="-1" metricUnit="m" metricValue="4.064" unit="in" value="16.0">16 in</quantity>
<taxonomicName id="C5500916EDAD41017BCBED4912F68E05" baseAuthorityName="Haseman" baseAuthorityYear="1911" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="davisi">
<emphasis id="E6ED89FE5545EFAAEEC0F6D37C41191E" italics="true">C. davisi</emphasis>
</taxonomicName>
; seven or eight in 
<taxonomicName id="963AC0655E53B73C797800A9B949A67B" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taroba">
<emphasis id="4AADAF1C0C1A80E5CA7D51EF5AB1DD86" italics="true">C. taroba</emphasis>
</taxonomicName>
); 20–22 interopercular odontodes (vs. 12 or 
<quantity id="4A41103E10C200E6E7124946BD8FAED6" metricMagnitude="-1" metricUnit="m" metricValue="3.302" unit="in" value="13.0">13 in</quantity>
<taxonomicName id="CFF5AB764F8601BD7DE22F734C9D43D1" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="naipi">
<emphasis id="177A1AE96F002589ADA1D01C102EB024" italics="true">C. naipi</emphasis>
</taxonomicName>
); 21 dorsal procurrent caudal-fin rays (vs. 
<quantity id="279F8E4F81891FB150435C87BD9142D6" metricMagnitude="-1" metricUnit="m" metricValue="4.064" metricValueMax="4.318" metricValueMin="3.81" unit="in" value="16.0" valueMax="17.0" valueMin="15.0">15–17 in</quantity>
<taxonomicName id="07A24A957DDF86F75952E35D4499FB8C" baseAuthorityName="de Pinna" baseAuthorityYear="1992" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="castroi">
<emphasis id="94618C23A046E094971B33A2B69A9CB1" italics="true">C. castroi</emphasis>
</taxonomicName>
and 
<taxonomicName id="E9C618967D9B73CA3697C5756C1C9CC4" authorityName="Costa, Abilhoa, Dalcin &amp; Katz" authorityYear="2022" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="melanoptera">
<emphasis id="6A42D958A6864D6576DCFC8165131E3D" italics="true">C. melanoptera</emphasis>
</taxonomicName>
; 18 or 
<quantity id="F9F57D0EE889C22ACD65A989A1CA0ABB" metricMagnitude="-1" metricUnit="m" metricValue="4.826" unit="in" value="19.0">19 in</quantity>
<taxonomicName id="91943FF4E0D4592BEA48FDDFFA450007" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="naipi">
<emphasis id="98360B00A179EE3ADF670D42BC246CCC" italics="true">C. naipi</emphasis>
</taxonomicName>
; 
<quantity id="3F8A0D9BB256744392E0D87986AEFF2F" metricMagnitude="-1" metricUnit="m" metricValue="6.858" metricValueMax="7.366" metricValueMin="6.35" unit="in" value="27.0" valueMax="29.0" valueMin="25.0">25–29 in</quantity>
<taxonomicName id="77F83F11D7875AF1EB088D95592F4089" baseAuthorityName="Wosiacki &amp; de Pinna" baseAuthorityYear="2008" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="crassicaudata">
<emphasis id="071B0D2F5C604FAC25CA5E773B721DC5" italics="true">C. crassicaudata</emphasis>
</taxonomicName>
, 
<taxonomicName id="33EE2320C0177674AE428CD783D31079" baseAuthorityName="Wosiacki &amp; de Pinna" baseAuthorityYear="2008" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="igobi">
<emphasis id="D585AB1155031E6002A69C49391080C2" italics="true">C. igobi</emphasis>
</taxonomicName>
, 
<taxonomicName id="F0D17D7367F3E173FBF851D29985E5ED" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="mboycy">
<emphasis id="52EF3ABB056953D14D841569B623F6BC" italics="true">C. mboycy</emphasis>
</taxonomicName>
, 
<taxonomicName id="347460FE2CF3CDF192B1ACD80860E0B1" authorityName="" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1968" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="stawiarski">
<emphasis id="4DE2AC4C500446CC31207DE95A3F8828" italics="true">C. stawiarski</emphasis>
</taxonomicName>
, and 
<taxonomicName id="14B2F5C049590E6C46A4A2946487D431" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taroba">
<emphasis id="E89B41AFD489B65DACC1FB64F4E827D0" italics="true">C. taroba</emphasis>
</taxonomicName>
; 30 or 
<quantity id="8616A0A16E421BE2982AD906D5310C57" metricMagnitude="-1" metricUnit="m" metricValue="7.874" unit="in" value="31.0">31 in</quantity>
<taxonomicName id="56C9859975ADE32433F4D1CB82519AED" authorityName="dos Reis, Ferrer &amp; Graca" authorityYear="2021" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="cauim">
<emphasis id="E9F71DA16DEF52D41726FD2D39C1FB18" italics="true">C. cauim</emphasis>
</taxonomicName>
); 13 or 14 ventral procurrent caudal-fin rays (vs. 
<quantity id="3FA97DDB44B1EB59955083D473B3C6E3" metricMagnitude="-1" metricUnit="m" metricValue="5.588" metricValueMax="5.842" metricValueMin="5.334" unit="in" value="22.0" valueMax="23.0" valueMin="21.0">21 – 23 in</quantity>
<taxonomicName id="36914437308A03153E79AB54F93FE101" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taroba">
<emphasis id="3B9C5223FEABE939F87AEE8192AB46BB" italics="true">C. taroba</emphasis>
</taxonomicName>
); dorsal-fin origin at a vertical through the centrum of the 21 
<superScript id="83215A74CECE995A327BE52BB5FDCFFB">st</superScript>
or 22 
<superScript id="51F5C5254BAC167E92C5FAA1F04F6E4E">nd</superScript>
vertebra (vs. 19 
<superScript id="4AADA39C3527643C8743FAD6BF33A804">th</superScript>
or 20 
<superScript id="F6CBB3B630435A495ADC29F01982D1DB">th</superScript>
in 
<taxonomicName id="A1809341C975FF4A8804EE8153E383B1" baseAuthorityName="Wosiacki &amp; de Pinna" baseAuthorityYear="2008" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="crassicaudata">
<emphasis id="38068E7A4F4B2725BCC1A4F38A29D6C8" italics="true">C. crassicaudata</emphasis>
</taxonomicName>
, 
<taxonomicName id="B867DFAA4B63E4D74CE56FEB359EFECA" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="mboycy">
<emphasis id="1EA52175545A0CBFC71992B6825046A6" italics="true">C. mboycy</emphasis>
</taxonomicName>
, and 
<taxonomicName id="96A132DC8235C10D6904831515ED5C86" authorityName="" baseAuthorityName="Miranda Ribeiro" baseAuthorityYear="1968" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="stawiarski">
<emphasis id="BF4DB05E3841F2430C8C570E5D7FB263" italics="true">C. stawiarski</emphasis>
</taxonomicName>
); jaw teeth pointed, irregularly arranged (vs. incisiform and arranged in rows in 
<taxonomicName id="33C672D82E71D5F58404B825E81E6E6E" baseAuthorityName="Haseman" baseAuthorityYear="1911" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="davisi">
<emphasis id="40FFAE85F666EDDE3B31B1501281FE9A" italics="true">C. davisi</emphasis>
</taxonomicName>
); and 39 or 40 vertebrae (
<quantity id="9ACBB96F61C4777D061CF53F624472B2" metricMagnitude="-1" metricUnit="m" metricValue="9.144" unit="in" value="36.0">36 in</quantity>
<taxonomicName id="3F57B75B091D5DA144C4B589E297F03A" baseAuthorityName="Wosiacki &amp; Garavello" baseAuthorityYear="2004" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="taroba">
<emphasis id="583A5365DEC280323103FE7A6D02F848" italics="true">C. taroba</emphasis>
</taxonomicName>
). Molecular diagnosis: combination of 17 nucleotide substitutions, five of them unique among taxa analysed *, and four unique for the 
<taxonomicName id="3FE01E6401328D198C358FEDD893555B" authorityName="Katz, Barbosa, Mattos &amp; Costa" authorityYear="2018" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="genus">
<emphasis id="B8204A87EBB565D09EFAE9556AEA8EFF" italics="true">Cambeva</emphasis>
</taxonomicName>
beta-clade **: 
<abbrev id="ABBRID0ESWAI" xlink_title="cytochrome c oxidase I">COX 1</abbrev>
216 (A → C) *, 
<abbrev id="ABBRID0EWWAI" xlink_title="cytochrome c oxidase I">COX 1</abbrev>
249 (T → C), 
<abbrev id="ABBRID0E1WAI" xlink_title="cytochrome c oxidase I">COX 1</abbrev>
252 (G → A), 
<abbrev id="ABBRID0E5WAI" xlink_title="cytochrome c oxidase I">COX 1</abbrev>
471 (G → A), 
<abbrev id="ABBRID0ECXAI" xlink_title="cytochrome c oxidase I">COX 1</abbrev>
534 (G → A), 
<abbrev id="ABBRID0EGXAI" xlink_title="genes cytochrome b">CYTB</abbrev>
108 (C → T), 
<abbrev id="ABBRID0EKXAI" xlink_title="genes cytochrome b">CYTB</abbrev>
186 (C → T) **, 
<abbrev id="ABBRID0EOXAI" xlink_title="genes cytochrome b">CYTB</abbrev>
237 (C → T) *, 
<abbrev id="ABBRID0ESXAI" xlink_title="genes cytochrome b">CYTB</abbrev>
360 (T → C) **, 
<abbrev id="ABBRID0EWXAI" xlink_title="genes cytochrome b">CYTB</abbrev>
378 (C → T) *, 
<abbrev id="ABBRID0E1XAI" xlink_title="genes cytochrome b">CYTB</abbrev>
442 (C → T) *, 
<abbrev id="ABBRID0E5XAI" xlink_title="genes cytochrome b">CYTB</abbrev>
765 (C → T), 
<abbrev id="ABBRID0ECYAI" xlink_title="genes cytochrome b">CYTB</abbrev>
909 (A → T) **, 
<abbrev id="ABBRID0EGYAI" xlink_title="genes cytochrome b">CYTB</abbrev>
942 (A → G), 
<abbrev id="ABBRID0EKYAI" xlink_title="genes cytochrome b">CYTB</abbrev>
960 (A → T) *, 
<abbrev id="ABBRID0EOYAI" xlink_title="genes cytochrome b">CYTB</abbrev>
994 (A → C). 
<abbrev id="ABBRID0ESYAI" xlink_title="cytochrome c oxidase I">COX 1</abbrev>
p-distances among congeners of the 
<taxonomicName id="16D8DC8C6607564B6EF9750546245544" authorityName="Katz, Barbosa, Mattos &amp; Costa" authorityYear="2018" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="genus">
<emphasis id="7F20A4BB7CF396BC0667BF568FAADB6E" italics="true">Cambeva</emphasis>
</taxonomicName>
beta-clade ranging from 1.2 (
<taxonomicName id="0F6A29D6875EE91BEE09357301693211" authority="Costa, Feltrin &amp; Katz, 2022" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2022" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="chrysornata">
<emphasis id="BB792BEA950DB78B285B30713041E0F4" italics="true">Cambeva chrysornata</emphasis>
Costa, Feltrin &amp; Katz, 2022
</taxonomicName>
and 
<taxonomicName id="09B49328046638C4B41A65741D1310EA" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="rotundipinna" status="sp. nov.">
<emphasis id="79A35705B4B8C19709437FA6D2809B25" italics="true">Cambeva rotundipinna</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="D99F27F00DE41CAF4B44A217EE13F1C5" rank="species">sp. nov.</taxonomicNameLabel>
) and 3.9.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETZAI" type="description">
<paragraph id="AB2AC276F77DAA920146E0B864477F5F">
<heading id="4501A07662C5F60971F4AD08EB96AC2C" reason="title">Description.</heading>
</paragraph>
<paragraph id="C81D67CE81B83B7FFB2CE7E1F760F1F0">
Morphometric data appear in Table 
<tableCitation id="5680BB56108DEDF63AE6648AC4F8575D" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphometric data of Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov." httpUri="http://table.plazi.org/id/D596199D62DCD4AF7E683F5341BA35E7" tableUuid="D596199D62DCD4AF7E683F5341BA35E7">4</tableCitation>
.
</paragraph>
<caption id="D596199D62DCD4AF7E683F5341BA35E7" ID-Table-UUID="D596199D62DCD4AF7E683F5341BA35E7" httpUri="http://table.plazi.org/id/D596199D62DCD4AF7E683F5341BA35E7" startId="T4" targetIsTable="true">
<paragraph id="FAFFA99DA7E0CC0C11514997347879C1">
<label id="EE91EF2DCAB684CCBFB0A4E65FF3750B">Table 4.</label>
</paragraph>
<paragraph id="1BDE5411E1DE82C8C88913934242E934">
Morphometric data of 
<taxonomicName id="CC642F47687AE67564E58F4506B6FF78" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea" status="sp. nov.">
<emphasis id="0023A0E6903E204BCB6164FB2EB1D73D" italics="true">Cambeva atrobrunnea</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="3F7063611DA2327F726A2378B5407464" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="1A73A0A646457D1CE2B04E9B27609DCD">
<table id="TID0EWPCI" rules="all">
<tbody id="F4830A47903531C80B2618D36FFEB18D">
<tr id="D9F700AA4F3EF1585487E345F28487C3">
<th id="212244E1B63F780479ED7EA02207177E" colspan="1" isEmpty="true" rowspan="1">-</th>
<th id="087A96D19DA0E7C51CA3D10C5B6BEA94" colspan="1" rowspan="1">Holotype</th>
<th id="7B31945646604BA259A80965DF2FCCB1" colspan="1" rowspan="1">Paratypes (n = 5)</th>
</tr>
<tr id="5F0AC3632540B4E672B26D17489E3156">
<td id="235B8771F28B46DE4858ADE2037AA7A5" colspan="1" rowspan="1">Standard length (SL)</td>
<td id="1E8FE954A121E94E28F231C3E0AFF991" colspan="1" rowspan="1">70.1</td>
<td id="BB0DF40D01A9D622630A3498571C4AB9" colspan="1" rowspan="1">46.8–78.5</td>
</tr>
<tr id="79B8EBCF236F9E0C274BAA4881C568E4">
<td id="D42FD23EAF3A01E7DE3229AA0B8D0ADF" colspan="3" rowspan="1">
<emphasis id="CEDCBE20E6D40C76377F72F74F26ECAD" bold="true">Percentage of standard length</emphasis>
</td>
</tr>
<tr id="46B99BA1FBE1D3B53A8C9ECE645E5530">
<td id="4A8DAB1E7D59027D7E6EE01CF928F975" colspan="1" rowspan="1">Body depth</td>
<td id="75D460A7EE94B2B1A988B37A98599857" colspan="1" rowspan="1">14.2</td>
<td id="271571221F8327AF7BC523707D2AB2F2" colspan="1" rowspan="1">12.2–15.0</td>
</tr>
<tr id="A91410AEDC3C30E4D983B78A286409AF">
<td id="EF6105F7B023431A4BFFB095F4E6D32B" colspan="1" rowspan="1">Caudal peduncle depth</td>
<td id="83675B742CDFA000BCBDEA2FBC7EE46D" colspan="1" rowspan="1">13.7</td>
<td id="AFCBACF42FDAA26A385A8B79553F716D" colspan="1" rowspan="1">11.2–13.1</td>
</tr>
<tr id="C16A1A4156667722097CF84A81CA427C">
<td id="6FFB4AC30FE2208D97E69792C9F38D63" colspan="1" rowspan="1">Body width</td>
<td id="1CC4D64CD916F654F5221566CA75BFDB" colspan="1" rowspan="1">12.1</td>
<td id="3FC81DCB82D2561691B13C4F38A0A90B" colspan="1" rowspan="1">10.3–12.8</td>
</tr>
<tr id="AA0F83D0752ED9605005B47816095DF5">
<td id="74B22EBBCFECA5950326CEAFDA1DAFD4" colspan="1" rowspan="1">Caudal peduncle width</td>
<td id="324B4C1A6DDA20DC41197889C2054F4F" colspan="1" rowspan="1">4.3</td>
<td id="FE54409FA6CF76C0C37802C76C62CF26" colspan="1" rowspan="1">3.0–5.0</td>
</tr>
<tr id="A3CB1D903F27ECFC61D5E8DFCD714FC2">
<td id="837FE790B395A56AFAA7392B12A7F7E5" colspan="1" rowspan="1">Pre-dorsal length</td>
<td id="768B93CEA47AA31128296AAC2A669866" colspan="1" rowspan="1">67.7</td>
<td id="12D7C8459FA0307B17473354B9235E0E" colspan="1" rowspan="1">63.5–67.2</td>
</tr>
<tr id="97A64D7773AA7A0F1303DA82E4BBD258">
<td id="67A2924DF0227BA2A46FB32D6E568395" colspan="1" rowspan="1">Pre-pelvic length</td>
<td id="2A81C53FF5B941D6FD9AB9E582335DC7" colspan="1" rowspan="1">62.7</td>
<td id="A2CE91C5E75731B77670730967EC405C" colspan="1" rowspan="1">57.9–60.3</td>
</tr>
<tr id="D99EFD5377DE737DDEF8D2D66D7CDA65">
<td id="9FA0B915C52FBFD26636A986A1648D8A" colspan="1" rowspan="1">Dorsal-fin base length</td>
<td id="D13A6EB74F8665753EE0B7A6C4C565D6" colspan="1" rowspan="1">13.4</td>
<td id="AA3EBC26B3DEA33F1584C8CDC0B2E091" colspan="1" rowspan="1">10.9–12.5</td>
</tr>
<tr id="59C93CE0B8C573EA80E5F2ED7FD75D3C">
<td id="D366A937213BDB44F61FD6590EBA3DA9" colspan="1" rowspan="1">Anal-fin base length</td>
<td id="75E7A636D7FBA48006655B9C3BE83997" colspan="1" rowspan="1">10.0</td>
<td id="AF1BAECCB2E9C779D07934549ED48D86" colspan="1" rowspan="1">9.0–12.7</td>
</tr>
<tr id="D0248845B0FD4238A798951B9B90F06F">
<td id="08485C1EA0F871310B00E3C4E7AD4667" colspan="1" rowspan="1">Caudal-fin length</td>
<td id="31D795FA58AF7242DB152695AE408DE1" colspan="1" rowspan="1">17.1</td>
<td id="643F2FB6AA31C930B9AA8CCE0414A9F0" colspan="1" rowspan="1">16.0–19.0</td>
</tr>
<tr id="DAFFC780195940B6FB4C55A9480BE8AE">
<td id="736E968A5A99D6FF1ABB2A6D401E27A9" colspan="1" rowspan="1">Pectoral-fin length</td>
<td id="7284F598FF3060C84483967F5F83B50A" colspan="1" rowspan="1">12.0</td>
<td id="0CE10E455E32681F4C8D13FA6E8E68AA" colspan="1" rowspan="1">11.6–13.1</td>
</tr>
<tr id="1B7E5AE6664D269330807E8398FFE194">
<td id="48069B8D4D9F2964DC2619C75305DECD" colspan="1" rowspan="1">Pelvic-fin length</td>
<td id="919DA3F30CE7DA6EF4BC25F243D47C3A" colspan="1" rowspan="1">8.7</td>
<td id="080EC6C322836E7CA4F3ED61078DCB8A" colspan="1" rowspan="1">9.1–12.4</td>
</tr>
<tr id="20429EDF3892C49ABBB7E5F1D6E50FAD">
<td id="4097A1A9467DD33E7F815655DED35A62" colspan="1" rowspan="1">Head length</td>
<td id="A19E0F3821FD7356CF8D57648FABD350" colspan="1" rowspan="1">22.3</td>
<td id="60115DEEB05E2206C50A1842479F7BFB" colspan="1" rowspan="1">19.5–24.7</td>
</tr>
<tr id="FF3B72FC112A65DCD0850F127EAE9EEB">
<td id="95AEADA24D2C5581933988D619B1D566" colspan="3" rowspan="1">
<emphasis id="A882E33E0AFAFF4F70835A9A7C46CF31" bold="true">Percentage of head length</emphasis>
</td>
</tr>
<tr id="F347454D39AC0684D5AA63EE2C574181">
<td id="F7F5187D6F52AD265DC899C562AC01F8" colspan="1" rowspan="1">Head depth</td>
<td id="D2043C20B2020AC9F36DA7C2A4ED418D" colspan="1" rowspan="1">43.9</td>
<td id="898474FC30B3746CC6A58D496C9FD603" colspan="1" rowspan="1">39.7–51.9</td>
</tr>
<tr id="F950444AC466B86C930238E9B43E7868">
<td id="E01C0912148F5650E02D09E69D2BC9DB" colspan="1" rowspan="1">Head width</td>
<td id="B70EEB1743D4AB15A37A760598E31C49" colspan="1" rowspan="1">82.7</td>
<td id="47B924E607FC0660D6365E3B33A9300E" colspan="1" rowspan="1">65.6–88.0</td>
</tr>
<tr id="4857EAA4C98E11C8389EDE81C4E68A0D">
<td id="ACAF65858859D869839BB86159FA2147" colspan="1" rowspan="1">Snout length</td>
<td id="95266F585222C53FDF420EA529792EE7" colspan="1" rowspan="1">39.9</td>
<td id="768996A7F78B919D47D381F6642B6CF6" colspan="1" rowspan="1">36.1–46.6</td>
</tr>
<tr id="BC78EB0D83C9C8A5E3721B5AF08F934A">
<td id="AD8EF5F77FC0B13E7FC33AAC71ADD716" colspan="1" rowspan="1">Interorbital width</td>
<td id="60CABDC5A6869FD32E610CF9C88259B4" colspan="1" rowspan="1">20.3</td>
<td id="0AF31523A5BB487FF688B542DE49BDA0" colspan="1" rowspan="1">18.3–23.6</td>
</tr>
<tr id="0AEA2ADEF5AC7A0D9A9234623518D823">
<td id="96BD0E304AF17D405D1E2D9F206B3AF2" colspan="1" rowspan="1">Preorbital length</td>
<td id="2E00DBB96FFA6A0538CDDEF171ECC76C" colspan="1" rowspan="1">12.2</td>
<td id="ECCAD8268A116400DD69EA980620D88A" colspan="1" rowspan="1">10.6–15.4</td>
</tr>
<tr id="763029B3AD8B3CB1F8AF007795146D8B">
<td id="17B7B98F719320891C5A8808C12626C1" colspan="1" rowspan="1">Eye diameter</td>
<td id="BE428CE3D2AA1DEA480600E03DBB871E" colspan="1" rowspan="1">8.4</td>
<td id="EC5B9873A93AA480D07DB40B7A5CDF40" colspan="1" rowspan="1">8.2–12.4</td>
</tr>
</tbody>
</table>
</paragraph>
<paragraph id="20F96C3D166D8BE35BE2105BBC5A3D3B">
<emphasis id="8B881B1219D93DC6E5F89F201CBA0428" bold="true">
<emphasis id="621654A90A68EE475404E3302D15FBCB" bold="true" italics="true">Head morphology</emphasis>
.
</emphasis>
Barbels moderate in length. Nasal barbel reaching between orbit and opercular patch of odontodes, maxillary barbel reaching posterior portion of interopercular patch of odontodes, and rictal barbel reaching middle of interopercular patch of odontodes. Jaw teeth variable in shape, smaller teeth slightly pointed, larger teeth sub-incisiform with slightly rounded tip, irregularly arranged. Premaxillary teeth 39–41, dentary teeth 40. Opercular and interopercular odontodes pointed, about straight. Opercular odontodes 12; interopercular odontodes 20–22. Anterior infraorbital sensory canal absent.
</paragraph>
<paragraph id="ECF766521D6ABA4EEE4E50F24F14E9F9">
<emphasis id="8993EE6CBD5EF2D88B23B4E9B06C1E90" bold="true">
<emphasis id="6BD5E20FCA522FFCEB5BCD4B6523EAD8" bold="true" italics="true">Fin morphology</emphasis>
.
</emphasis>
Dorsal and anal fins subtriangular, distal margin slightly convex. Total dorsal-fin rays 10 or 11 (ii + II – III + 6–7), total anal-fin rays 9 or 10 (ii + II + 5–6). Anal-fin origin at vertical just posterior to middle of dorsal-fin base, at base of 4 
<superScript id="9002F7500BDBE5803C5BDBE024FEC8CD">th</superScript>
branched dorsal-fin ray. Pectoral fin sub-triangular in dorsal view, margins slightly convex, first pectoral-fin ray shorter than second ray, not forming terminal filament. Total pectoral-fin rays 7 (I + 6). Pelvic fin rounded, its tip reaching vertical through anterior portion of dorsal-fin base. Total pelvic-fin rays 5 (I + 4). Caudal fin subtruncate, posterior margin weakly convex. Total principal caudal-fin rays 13 (I + 11 + I), total dorsal procurrent rays 21 (xix + II), total ventral procurrent rays 13 or 14 (xi – xii + II).
</paragraph>
<paragraph id="85D0DAC5FD3E9AF9B8E313590726AA27">
<emphasis id="7CB070D83FEEB29FB6F151D726A04921" bold="true">
<emphasis id="8D38B00F8903B9CD1AC6596A5314EDA4" bold="true" italics="true">Osteology</emphasis>
(Fig. 
<figureCitation id="8F52EE58C845E1BC3EB414503BEDDA1D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Osteological structures of: A – C. Cambeva galactica Costa, Feltrin &amp; Katz, sp. nov.; D – F. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov.; G – I. Cambeva luteoreticulata Costa, Feltrin &amp; Katz, sp. nov., and J – L. Cambeva rotundipinna Costa, Feltrin &amp; Katz, sp. nov. A, D, G, J. Mesethmoidal region and adjacent structures, left and middle portions, dorsal view; B, E, H, K. Left jaw suspensorium and opercular series, lateral view. C, F, I, L. Parurohyal, ventral view. Abbreviation of structure indicated by arrow: aes, anteromedial expansion of sesamoid supraorbital. Larger stippling represents cartilaginous areas." httpUri="https://binary.pensoft.net/fig/1131386">4 D – F</figureCitation>
).
</emphasis>
Mesethmoid broader anteriorly, without lateral expansions in its main axis, anterior mesethmoid margin slightly convex. Mesethmoid cornu extremity rounded. Lateral ethmoid with small lateral projection immediately posterior to articular facet for autopalatine and small, twisted expansion on anterior margin. Lacrimal narrow, short and thin. Sesamoid supraorbital rod-shaped, narrower than lacrimal, its longitudinal length about two times and half longer than lacrimal longitudinal length, without lateral expansions. Premaxilla long, laterally narrowing, slightly curved. Maxilla slender, with rudimentary posterior process, slightly curved, its length about three fourths of premaxilla. Autopalatine sub-trapezoidal in dorsal view, medial margin sinuous, lateral margin weakly concave. Autopalatine postero-lateral process triangular, short, its length about half autopalatine length excluding anterior cartilage. Autopalatine articulation for lateral ethmoid with laminar shovel-shaped expansion.
</paragraph>
<paragraph id="755A5F01039C416CDB32C6871DB0B9CF">Metapterygoid sub-rectangular, longer than deep, relatively large, its surface greater than quadrate lateral surface. Areas anterior and posterior to cartilaginous articulation between metapterygoid and quadrate with small laminar overlapped expansions forming additional points of articulation. Quadrate with deep anterior constriction at dorsal process base. Hyomandibula long, anterior outgrowth horizontal length slightly longer than largest horizontal metapterygoid length; dorsal margin of hyomandibula outgrowth concave. Opercle elongate, longer than interopercle. Opercular odontode patch slender, its depth about half hyomandibula articular facet length. Dorsal process of opercle short, subtriangular, its extremity rounded. Opercular articular facet for hyomandibula with dorsal, trapezoidal laminar projection, articular facet for preopercle small, rounded. Interopercle moderate in length, interopercular odontode patch length about equal hyomandibula outgrowth length. Preopercle slender, with minute ventral projection.</paragraph>
<paragraph id="D77279C7C52A0A79A4F9308F153D282B">
Parurohyal robust, lateral process subtriangular, slightly curved posteriorly, with pointed tip. Parurohyal head with prominent anterolateral paired process. Parurohyal middle foramen relatively large, oval. Parurohyal posterior process moderate in length, about half of distance between anterior margin of parurohyal and anterior insertion of posterior process. Branchiostegal rays 9. Vertebrae 39 or 40. Ribs 14 or 15. Dorsal-fin origin at vertical through centrum of 21 
<superScript id="AFDA2DA6AFC7821B4B34E9D1303E859F">st</superScript>
or 22 
<superScript id="4F5F31294E408FBE2D32231E38881499">nd</superScript>
vertebra; anal-fin origin at vertical through centrum of 25 
<superScript id="DEBBC68CC8825A78F88BA599D57B1059">th</superScript>
or 26 
<superScript id="EF2CE3B5AD4088D3AB8C175A643A07BA">th</superScript>
vertebra. Two or one dorsal and single ventral hypural plate.
</paragraph>
<paragraph id="FB85C5FFB8545647412E68A5AF18586C">
<emphasis id="C33A10F59848B4563B5BAE0C1DAF6FD2" bold="true">
<emphasis id="BD9764969EB655214980EB9B96D3EF44" bold="true" italics="true">Colouration in alcohol</emphasis>
.
</emphasis>
In adult specimens (Fig. 
<figureCitation id="D4506FF1AE8E0D3820CB069B96CE1F69" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov., holotype, UFRJ 14066, 70.1 mm SL. A. Lateral view; B. Dorsal view; C. Ventral view." httpUri="https://binary.pensoft.net/fig/1131388">6</figureCitation>
), flank, dorsum, and head side light brownish yellow, with great concentration of small dark brown to black dots. Venter and ventral surface of head brownish white, with minute dark brown dots in area just anterior to pelvic fin. Nasal and maxillary barbels brown, rictal barbel brownish white. Fins hyaline, with minute dark brown dots on basal region of dorsal, anal and pectoral fins, and on whole caudal fin. In juvenile specimens below about 
<quantity id="1A35BCEC5B7499D54B5E2A5F5F77ED61" metricMagnitude="-2" metricUnit="m" metricValue="5.0" unit="mm" value="50.0">50 mm</quantity>
SL (Fig. 
<figureCitation id="E39B2B4791D499E7D3768AA6F5ABC435" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov., paratype, UFRJ 13821, 33.4 mm SL. A. Lateral view; B. Dorsal view; C. Ventral view." httpUri="https://binary.pensoft.net/fig/1131389">7</figureCitation>
), flank, dorsum and head side light grey, with small black dots arranged in irregular longitudinal rows, coalesced on anterior portion of longitudinal midline of flank to form black stripe.
</paragraph>
<caption id="3810844AA9D93E0669A104C69BDAF0A1" ID-arpha="DC2C66F5-0A1C-5112-8CFC-429F5C89D8BC" httpUri="https://binary.pensoft.net/fig/1131388" startId="F6">
<paragraph id="8EE7ECBFE5CE9596C3D32B69BA211300">
<label id="2C8D862460E2F38A0750CAEDFF832408">Figure 6.</label>
</paragraph>
<paragraph id="B4EF87F1854B7CBF95B410570992DFC9">
<taxonomicName id="017DE60F086157363C240A97BBA713C8" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea" status="sp. nov.">
<emphasis id="93B7AD1DBBB4A12EC046E5C7264C8547" italics="true">Cambeva atrobrunnea</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="9F79A279C4A123F3BE15BDB11251C5A2" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, 
<specimenCode id="D76DE22C6B863377B97CB8153B79D169">
<named-content id="42E5E741679A0CD0BC7B5B261398DDB7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-rural-do-rio-de-janeiro" xlink_title="Instituto de Biologia, Universidade Federal do Rio de Janeiro">UFRJ</named-content>
14066
</specimenCode>
, 70.1 mm SL. 
<emphasis id="B5A96B867D39C337B267C96AC8D312BE" bold="true">A.</emphasis>
Lateral view; 
<emphasis id="21EE3F00B2DFD2F90D379EDEF3701015" bold="true">B.</emphasis>
Dorsal view; 
<emphasis id="D74FC3817B177CED9E21FA518E2A08E9" bold="true">C.</emphasis>
Ventral view.
</paragraph>
</caption>
<caption id="7C08A8786EDE550B30C26DBE5F3E0815" ID-arpha="C8E71696-D3D1-5CDE-BC97-91799CC9A9C3" httpUri="https://binary.pensoft.net/fig/1131389" startId="F7">
<paragraph id="7F094EB8D5B056162B79285423C3091B">
<label id="32E6FE8211CAC9FB75E59B3D55EF6A1B">Figure 7.</label>
</paragraph>
<paragraph id="FEE044C0FABDE33578DADF7A7AC00A79">
<taxonomicName id="C404B04F47619E495F27661DB940E3A7" authority="Costa, Feltrin &amp; Katz" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea" status="sp. nov.">
<emphasis id="458E9F5728E027B0AE6ACE409FF3C59B" italics="true">Cambeva atrobrunnea</emphasis>
Costa, Feltrin &amp; Katz
</taxonomicName>
, 
<taxonomicNameLabel id="FC05AD6A375DC893DB4E80E324E10D74" rank="species">sp. nov.</taxonomicNameLabel>
, paratype, 
<specimenCode id="FEB64BF311A07F887298A0B6D73E5CD7">
<named-content id="39F3AE312D50EEA4EB7F3758D1EEFE32" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/universidade-federal-rural-do-rio-de-janeiro" xlink_title="Instituto de Biologia, Universidade Federal do Rio de Janeiro">UFRJ</named-content>
13821
</specimenCode>
, 33.4 mm SL. 
<emphasis id="4965B9D6C0C7FED55402CCE4FBFD2EED" bold="true">A.</emphasis>
Lateral view; 
<emphasis id="1D626347FADEA0E411576B42A0AABE39" bold="true">B.</emphasis>
Dorsal view; 
<emphasis id="D4A853369B20178DCCE5B850D16A503E" bold="true">C.</emphasis>
Ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EEFBI" type="distribution">
<paragraph id="2CE512B17D17313E31DB194A48BE606D">
<heading id="1B2827D2EEBBEE39949ECD4A592F7A3C" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="E0D51BE2781A438FD72FB1B04DD2E716">
<taxonomicName id="A14E65C911372AB9C40F580F25A4D375" authorityName="Costa, Feltrin &amp; Katz" authorityYear="2024" family="Trichomycteridae" genus="Cambeva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" phylum="Chordata" rank="species" species="atrobrunnea">
<emphasis id="BE0CC0F88F31CB2671A666B45869E73D" italics="true">Cambeva atrobrunnea</emphasis>
</taxonomicName>
is known from a single locality in a stream tributary of the Rio Timbó, Rio Iguaçu drainage, Rio 
<collectingRegion id="23BBB7B3AC549A24790A660EEFF29E68" country="Brazil" name="Parana">Paraná</collectingRegion>
basin, at about 
<quantity id="9AA15CCFA47925F3380F2037E4ADF5A2" metricMagnitude="2" metricUnit="m" metricValue="9.7" unit="m" value="970.0">970 m</quantity>
asl (Fig. 
<figureCitation id="CAAC6E381A61F148E3A7BEE90AC14053" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Geographical distribution of four new species of Cambeva from the Rio Iguaçu drainage, Serra do Espigão, southern Brazil: 1. Cambeva galactica Costa, Feltrin &amp; Katz, sp. nov.; 2. Cambeva atrobrunnea Costa, Feltrin &amp; Katz, sp. nov.; 3. Cambeva luteoreticulata Costa, Feltrin &amp; Katz, sp. nov.; 4. Cambeva rotundipinna Costa, Feltrin &amp; Katz, sp. nov." httpUri="https://binary.pensoft.net/fig/1131387">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZFBI" type="etymology">
<paragraph id="65F959C793877A97BA5EA1A2DD9F24FE">
<heading id="2425DA381795EA78A2025442A638BA06" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="DDD6276EC773F1A389813B064CA216BA">From the Latin ater (dull black, dark) and brunneus (brown), referring to the predominant colour of the flank in adult specimens of the new species.</paragraph>
</subSubSection>
</treatment>
</document>