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<document id="33F0D3FDCEF18BC46886E7930439CB9E" ID-CLB-Dataset="46853" ID-DOI="10.5281/zenodo.201129" ID-GBIF-Dataset="2cb1a478-0e9c-4d1e-b7f9-4b882a6a4fd9" ID-ISSN="1175-5326" ID-Zenodo-Dep="201129" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460358460197" checkinUser="plazi" docAuthor="Cao, Liangming, Tomokuni, Masaaki &amp; Cai, Wanzhi" docDate="2011" docId="D9598780FF99FFFB48BF89A1FA34BC20" docLanguage="en" docName="zt02888p030.pdf" docOrigin="Zootaxa 2888" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Physoderes esakii Cao, Tomokuni &amp; Cai, 2011, sp. n." docType="treatment" docVersion="11" lastPageNumber="27" masterDocId="2560FFF8FF98FFFF48288B44FFBDBE7F" masterDocTitle="Taxonomic notes on Physoderinae (Heteroptera: Reduviidae) from China" masterLastPageNumber="30" masterPageNumber="23" pageNumber="24" updateTime="1698294171813" updateUser="ExternalLinkService">
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<mods:title id="06727CF0321D40D48355016C5FEF004D">Taxonomic notes on Physoderinae (Heteroptera: Reduviidae) from China</mods:title>
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<mods:namePart id="017539AE70C33FDDA6CEF7A5F4333352">Cao, Liangming</mods:namePart>
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<mods:namePart id="024DC2EA38C610C6BE009961F979F9B2">Cai, Wanzhi</mods:namePart>
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<mods:date id="FBAF194DAD7AA289DDB580348AD7E39A">2011</mods:date>
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<treatment id="D9598780FF99FFFB48BF89A1FA34BC20" ID-DOI="http://doi.org/10.5281/zenodo.6183389" ID-GBIF-Taxon="119409199" ID-Zenodo-Dep="6183389" LSID="urn:lsid:plazi:treatment:D9598780FF99FFFB48BF89A1FA34BC20" httpUri="http://treatment.plazi.org/id/D9598780FF99FFFB48BF89A1FA34BC20" lastPageId="4" lastPageNumber="27" pageId="1" pageNumber="24">
<subSubSection id="19EA651DFF99FFFE48BF89A1FEA4BD60" pageId="1" pageNumber="24" type="nomenclature">
<paragraph id="514F3696FF99FFFE48BF89A1FE0BBC80" blockId="1.[151,438,741,799]" box="[151,438,741,767]" pageId="1" pageNumber="24">
<heading id="0A0781FAFF99FFFE48BF89A1FE0BBC80" bold="true" box="[151,438,741,767]" fontSize="11" level="1" pageId="1" pageNumber="24" reason="1">
<emphasis id="6384EA84FF99FFFE48BF89A1FE0BBC80" bold="true" box="[151,438,741,767]" pageId="1" pageNumber="24">
<taxonomicName id="96F04D15FF99FFFE48BF89A1FED0BC80" ID-CoL="4HHTQ" box="[151,365,741,767]" class="Insecta" family="Reduviidae" genus="Physoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="esakii" status="sp. nov.">
<emphasis id="6384EA84FF99FFFE48BF89A1FED0BC80" bold="true" box="[151,365,741,767]" italics="true" pageId="1" pageNumber="24">Physoderes esakii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78B757FFFF99FFFE495C89A1FE0BBC80" box="[372,438,741,767]" pageId="1" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="514F3696FF99FFFE48BF8843FEA4BD60" blockId="1.[151,438,741,799]" box="[151,281,775,799]" pageId="1" pageNumber="24">
(
<figureCitation id="C9CB2A13FF99FFFE48B78843FEADBD60" box="[159,272,775,799]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 2 12" captionStartId-0="2.[151,250,1980,2003]" captionStartId-1="3.[151,264,1952,1975]" captionTargetBox-0="[187,1412,324,1961]" captionTargetBox-1="[155,1431,208,1864]" captionTargetId-0="figure@2.[173,1413,307,1992]" captionTargetId-1="figure@3.[151,1436,193,1931]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." captionText-1="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri-0="https://zenodo.org/record/201130/files/figure.png" httpUri-1="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">Figs.112</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="19EA651DFF99FFFE48BF881DFD02BDF2" pageId="1" pageNumber="24" type="reference_group">
<paragraph id="514F3696FF99FFFE48BF881DFD02BDF2" blockId="1.[151,707,857,910]" pageId="1" pageNumber="24">
<treatmentCitationGroup id="71E011B8FF99FFFE48BF881DFD02BDF2" pageId="1" pageNumber="24">
<taxonomicName id="96F04D15FF99FFFE48BF881DFDAEBD10" authority="Esaki 1931: 212" authorityName="Esaki" authorityPageNumber="212" authorityYear="1931" box="[151,531,857,879]" class="Insecta" family="Reduviidae" genus="Epirodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="latithorax">
<emphasis id="6384EA84FF99FFFE48BF881DFED5BD10" box="[151,360,857,879]" italics="true" pageId="1" pageNumber="24">Epirodera latithorax</emphasis>
<treatmentCitation id="D0511087FF99FFFE4946881DFDAEBD10" author="Esaki" box="[366,531,857,879]" page="212" pageId="1" pageNumber="24" year="1931">
<bibRefCitation id="35614B67FF99FFFE4946881DFE5EBD10" author="Esaki" box="[366,483,857,879]" pageId="1" pageNumber="24" refString="Esaki, T. (1931) Hemiptera - Heteroptera von der Insel Botol - Tobago (Kotosho), Sud Formosa. Bulletin of the Biogeographical Society of Japan, 2, 209 - 220." type="journal article" year="1931">Esaki 1931</bibRefCitation>
: 212
</treatmentCitation>
</taxonomicName>
(nomen nudum).
<taxonomicName id="96F04D15FF99FFFE48BF883CFD07BDF2" ID-CoL="4HHTZ" authority="Maldonado-Capriles 1990: 380" authorityName="Maldonado-Capriles" authorityPageNumber="380" authorityYear="1990" box="[151,698,888,910]" class="Insecta" family="Reduviidae" genus="Physoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="latithorax">
<emphasis id="6384EA84FF99FFFE48BF883CFEC8BDF1" box="[151,373,888,910]" italics="true" pageId="1" pageNumber="24">Physoderes latithorax</emphasis>
<treatmentCitation id="D0511087FF99FFFE4953883CFD07BDF2" author="Maldonado-Capriles" box="[379,698,888,910]" page="380" pageId="1" pageNumber="24" year="1990">
<bibRefCitation id="35614B67FF99FFFE4953883CFD35BDF2" author="Maldonado-Capriles" box="[379,648,888,910]" pageId="1" pageNumber="24" refString="Maldonado-Capriles J. (1990) Systematic catalogue of the Reduviidae of the world (Insecta: Heteroptera). A special edition of Caribbean Journal of Science, Puerto Rico, 694 pp." type="book" year="1990">Maldonado-Capriles 1990</bibRefCitation>
: 380
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="19EA651DFF99FFFE48BF88FEFEEBB999" pageId="1" pageNumber="24" type="description">
<paragraph id="514F3696FF99FFFE48BF88FEFCD9BA41" blockId="1.[151,1437,954,2022]" pageId="1" pageNumber="24">
<emphasis id="6384EA84FF99FFFE48BF88FEFEC6BDAC" bold="true" box="[151,379,954,979]" pageId="1" pageNumber="24">Description. Color.</emphasis>
Body generally brown; eyes, second visible labial segment, basal half of first antennal segment, four irregular spots on anterior pronotal lobe, posterior 1/3 of each connexival segment dorsally and ventrally, two semicircular spots on seventh abdominal tergite, paramedian portions of femora, basal portion of tibiae, and tarsi yellow; anterior 2/3 of each connexival segment black.
</paragraph>
<paragraph id="514F3696FF99FFFE48ED8F0EFB86BAF9" blockId="1.[151,1437,954,2022]" pageId="1" pageNumber="24">
<emphasis id="6384EA84FF99FFFE48ED8F0EFE80BA1C" bold="true" box="[197,317,1098,1123]" pageId="1" pageNumber="24">Structure.</emphasis>
Head, first to third antennal segments, apical tubercles on lateral pronotal margins and legs with curved club-shaped setae; fourth antennal segment with short and long dense setae.
</paragraph>
<paragraph id="514F3696FF99FFFE48ED8FD6FE3EBB69" blockId="1.[151,1437,954,2022]" pageId="1" pageNumber="24">
Head (
<figureCitation id="C9CB2A13FF99FFFE49388FD6FEE2BAD5" box="[272,351,1170,1194]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Figs. 1</figureCitation>
,
<figureCitation id="C9CB2A13FF99FFFE49468FD6FEC0BAD5" box="[366,381,1170,1194]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">2</figureCitation>
) columniform, cone-shaped anteriad to antennifers, not flat ventrally; eyes (
<figureCitation id="C9CB2A13FF99FFFE4CF38FD6FA94BAD5" box="[1243,1321,1170,1194]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Figs. 1</figureCitation>
,
<figureCitation id="C9CB2A13FF99FFFE4D118FD6FAF5BAD5" box="[1337,1352,1170,1194]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">2</figureCitation>
) small. Ocelli (
<figureCitation id="C9CB2A13FF99FFFE48C38FF2FE91BAB1" box="[235,300,1206,1230]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
) large, located far from each other. Each antennal segment (
<figureCitation id="C9CB2A13FF99FFFE4BF08FF2FBA4BAB1" box="[984,1049,1206,1230]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
) subequal in length; first segment thickest, third one thinnest. First visible labial segment (
<figureCitation id="C9CB2A13FF99FFFE4B2F8F9EFCF4BA8D" box="[775,841,1242,1266]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">Fig. 2</figureCitation>
) thick, about 1/4 of second segment; second segment elongate and slender.
</paragraph>
<paragraph id="514F3696FF99FFFE48ED8E66FEF0B825" blockId="1.[151,1437,954,2022]" pageId="1" pageNumber="24">
Collar processes (
<figureCitation id="C9CB2A13FF99FFFE49A68E66FE72BB45" box="[398,463,1314,1338]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
) tuberculiform and prominent. Anterior pronotal lobe (
<figureCitation id="C9CB2A13FF99FFFE4C6A8E66FB2DBB45" box="[1090,1168,1314,1338]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Figs. 1</figureCitation>
,
<figureCitation id="C9CB2A13FF99FFFE4CB78E66FB13BB45" box="[1183,1198,1314,1338]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">2</figureCitation>
) swollen, wider than posterior lobe, with a row of small tubers along anterior and lateral margins, and centrally with a round deep depression; other parts smooth and slightly shining. Posterior pronotal lobe (
<figureCitation id="C9CB2A13FF99FFFE4BD08E2EFBFBBBFD" box="[1016,1094,1386,1410]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Figs. 1</figureCitation>
,
<figureCitation id="C9CB2A13FF99FFFE4C7D8E2EFBD9BBFD" box="[1109,1124,1386,1410]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">2</figureCitation>
) distinctly lower than anterior lobe, widely concave in anteromedial area, with a transverse ridge on discal area; posterior angles semicircular, prominent backward. Hemelytra (
<figureCitation id="C9CB2A13FF99FFFE4A398EF6FDEFBBB5" box="[529,594,1458,1482]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
) just reaching base of seventh abdominal tergite; corium (
<figureCitation id="C9CB2A13FF99FFFE4CC98EF6FA9CBBB5" box="[1249,1313,1458,1482]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
) elongatetriangular, occupied small portion of forewing. Fore femora (
<figureCitation id="C9CB2A13FF99FFFE4B798E92FC1DBB91" box="[849,928,1494,1518]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Figs. 1</figureCitation>
,
<figureCitation id="C9CB2A13FF99FFFE4B988E92FC02BB91" box="[944,959,1494,1518]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">4</figureCitation>
) each with rows of small tubers and with two rows of large spines ventrally; mid femora (
<figureCitation id="C9CB2A13FF99FFFE4A878EBEFD52B86D" box="[687,751,1530,1554]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
) with just one row of spines ventrally; tibiae (
<figureCitation id="C9CB2A13FF99FFFE4CDA8EBEFA82B86D" box="[1266,1343,1530,1554]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Figs. 1</figureCitation>
,
<figureCitation id="C9CB2A13FF99FFFE4D648EBEFAE6B86D" box="[1356,1371,1530,1554]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">4</figureCitation>
) with 7-8 rows of large or small tubers, fore tibiae each with a large process subapically, and with a cleaning comb at prominent apex.
</paragraph>
<paragraph id="514F3696FF99FFFE48ED8D22FC8AB94D" blockId="1.[151,1437,954,2022]" pageId="1" pageNumber="24">
Abdomen patulous; connexiva visible from above (
<figureCitation id="C9CB2A13FF99FFFE4B2C8D22FCFBB801" box="[772,838,1638,1662]" captionStart="FIGURE 1" captionStartId="2.[151,250,1980,2003]" captionTargetBox="[187,1412,324,1961]" captionTargetId="figure@2.[173,1413,307,1992]" captionTargetPageId="2" captionText="FIGURE 1. Physoderes esakii sp. n. Habitus. 3. Scale bar = 1.25 mm." httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="1" pageNumber="24">Fig. 1</figureCitation>
). Pygophore (
<figureCitation id="C9CB2A13FF99FFFE4BCF8D22FBD3B801" box="[999,1134,1638,1662]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">Figs. 3, 8, 9</figureCitation>
) nearly straight along posterior margin; pygophore process hemi-annular, middle thick (
<figureCitation id="C9CB2A13FF99FFFE4B7B8DCEFC2AB8DD" box="[851,919,1674,1698]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">Fig. 9</figureCitation>
). Paramere (
<figureCitation id="C9CB2A13FF99FFFE4C0E8DCEFB2CB8DD" box="[1062,1169,1674,1698]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">Figs. 57</figureCitation>
) flattened, elongate-triangular, swollen in anterior half, with apical margin nearly straight. Phallus as shown in
<figureCitation id="C9CB2A13FF99FFFE4CA58DEAFAABB8B9" box="[1165,1302,1710,1734]" captionStart="FIGURES 2 12" captionStartId="3.[151,264,1952,1975]" captionTargetBox="[155,1431,208,1864]" captionTargetId="figure@3.[151,1436,193,1931]" captionTargetPageId="3" captionText="FIGURES 2 12. Physoderes esakii sp. n. 3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 5 7, paramere; 8, 9, pygophore; 10 12, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 5 7 = 0.17 mm; of 8, 9 = 0.33 mm; of 10 12 = 0.25 mm." httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="1" pageNumber="24">Figs. 1012</figureCitation>
; basal plate slender and long; basal plate bridge thin and nearly straight; pedicel short; phallosoma oblong, slightly swollen apically, with apex concave centrally; ventral phallothecal sclerite sclerotized conspicuously; struts fused each other, reaching to subapical part of phallosoma; endosoma simple.
</paragraph>
<paragraph id="514F3696FF99FFFE48ED8C7BFEEBB999" blockId="1.[151,1437,954,2022]" pageId="1" pageNumber="24">
<emphasis id="6384EA84FF99FFFE48ED8C7BFEC6B928" bold="true" box="[197,379,1855,1879]" pageId="1" pageNumber="24">Measurements.</emphasis>
[3 (n=1)]. Body length 10.5; maximum width of abdomen 3.95; head length 2.2; length of anteocular portion 1.05; length of postocular portion 0.75; synthlipsis 0.6; interocellar space 0.35; lengths of antennal segments I-IV=0.55, 0.7, 0.6, 0.8; lengths of visible labial segments I-III=0.4, 1.7, 0.5; length of anterior pronotal lobe 1.5; length of posterior pronotal lobe 1.3; maximum width of thorax 3.5; length of scutellum 0.65; length of hemelytron 5.45.
</paragraph>
</subSubSection>
<subSubSection id="19EA651DFF9AFFFB48ED8BDCFC58BFB0" lastPageId="4" lastPageNumber="27" pageId="2" pageNumber="25" type="materials_examined">
<paragraph id="514F3696FF9AFFFD48ED8BDCFDFABF63" blockId="2.[151,1437,151,284]" pageId="2" pageNumber="25">
<emphasis id="6384EA84FF9AFFFD48ED8BDCFE58BECF" bold="true" box="[197,485,151,176]" pageId="2" pageNumber="25">
<typeStatus id="8E4B8834FF9AFFFD48ED8BDCFEBDBECF" box="[197,256,152,176]" pageId="2" pageNumber="25">Type</typeStatus>
material.
<typeStatus id="8E4B8834FF9AFFFD49508BDCFE58BECF" box="[376,485,152,176]" pageId="2" pageNumber="25" type="holotype">Holotype</typeStatus>
</emphasis>
, 3,
<collectingCountry id="29E77606FF9AFFFD4A3D8BD3FDE6BED0" box="[533,603,151,175]" name="China" pageId="2" pageNumber="25">China</collectingCountry>
,
<collectingCountry id="29E77606FF9AFFFD4A4E8BD3FD05BED0" box="[614,696,151,175]" name="Taiwan" pageId="2" pageNumber="25">Taiwan</collectingCountry>
Province, Botol-Tobago
<collectingCountry id="29E77606FF9AFFFD4BF88BD3FBAABED0" box="[976,1047,151,175]" name="Iceland" pageId="2" pageNumber="25">Island</collectingCountry>
, earlier than 1931, Tadao Kano leg (Kyushu University). Right antennal segments 3-4 missing, left antennal segments glued on the label, right mid tibia and tarsus missing.
<bibRefCitation id="35614B67FF9AFFFD49828B9BFD84BE88" author="Esaki" box="[426,569,223,247]" pageId="2" pageNumber="25" refString="Esaki, T. (1931) Hemiptera - Heteroptera von der Insel Botol - Tobago (Kotosho), Sud Formosa. Bulletin of the Biogeographical Society of Japan, 2, 209 - 220." type="journal article" year="1931">Esaki (1931)</bibRefCitation>
reported this specimen as female. He might have mistaken the patulous abdomen of the male for that of the female.
</paragraph>
<caption id="058F661EFF9AFFFD48BF8CF8FCD1B9AD" box="[151,876,1979,2003]" httpUri="https://zenodo.org/record/201130/files/figure.png" pageId="2" pageNumber="25" targetBox="[187,1412,324,1961]" targetPageId="2">
<paragraph id="514F3696FF9AFFFD48BF8CF8FCD1B9AD" blockId="2.[151,876,1979,2003]" box="[151,876,1979,2003]" pageId="2" pageNumber="25">
<emphasis id="6384EA84FF9AFFFD48BF8CF8FEAEB9AD" bold="true" box="[151,275,1980,2003]" pageId="2" pageNumber="25">FIGURE 1.</emphasis>
<taxonomicName id="96F04D15FF9AFFFD49318CF9FE73B9AC" box="[281,462,1981,2003]" class="Insecta" family="Reduviidae" genus="Physoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="esakii" status="sp. nov.">
<emphasis id="6384EA84FF9AFFFD49318CF9FE73B9AC" box="[281,462,1981,2003]" italics="true" pageId="2" pageNumber="25">Physoderes esakii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78B757FFFF9AFFFD49FC8CF9FDB0B9AD" box="[468,525,1981,2002]" pageId="2" pageNumber="25" rank="species">
<emphasis id="6384EA84FF9AFFFD49FC8CF9FDB0B9AD" bold="true" box="[468,525,1981,2002]" pageId="2" pageNumber="25">sp. n.</emphasis>
</taxonomicNameLabel>
Habitus. 3. Scale bar = 1.25 mm.
</paragraph>
</caption>
<caption id="058F661EFF9BFFFC48BF8CE4FAB3B98C" httpUri="https://zenodo.org/record/201131/files/figure.png" pageId="3" pageNumber="26" targetBox="[155,1431,208,1864]" targetPageId="3">
<paragraph id="514F3696FF9BFFFC48BF8CE4FAB3B98C" blockId="3.[151,1436,1952,2035]" pageId="3" pageNumber="26">
<emphasis id="6384EA84FF9BFFFC48BF8CE4FEFAB9C9" bold="true" box="[151,327,1952,1975]" pageId="3" pageNumber="26">FIGURES 212.</emphasis>
<taxonomicName id="96F04D15FF9BFFFC49658CE4FDBEB9C9" box="[333,515,1952,1974]" class="Insecta" family="Reduviidae" genus="Physoderes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="esakii" status="sp. nov.">
<emphasis id="6384EA84FF9BFFFC49658CE4FDBEB9C9" box="[333,515,1952,1974]" italics="true" pageId="3" pageNumber="26">Physoderes esakii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="78B757FFFF9BFFFC4A218CE5FDFEB9C9" box="[521,579,1953,1974]" pageId="3" pageNumber="26" rank="species">
<emphasis id="6384EA84FF9BFFFC4A218CE5FDFEB9C9" bold="true" box="[521,579,1953,1974]" pageId="3" pageNumber="26">sp. n.</emphasis>
</taxonomicNameLabel>
3. 2, Head and pronotum, antennae removed; 3, apical portion of abdomen; 4, right fore leg; 57, paramere; 8, 9, pygophore; 1012, phallus. 2, 4, 8, 11, lateral view; 3, 9, 12, ventral view; 10, dorsal view. Scale bar of 2 = 0.8 mm; of 3 = 1.00 mm; of 4 = 0.77 mm; of 57 = 0.17 mm; of 8, 9 = 0.33 mm; of 1012 = 0.25 mm.
</paragraph>
</caption>
<paragraph id="514F3696FF9CFFFB48ED8BD3FC58BFB0" blockId="4.[151,1436,151,608]" pageId="4" pageNumber="27">
The
<typeStatus id="8E4B8834FF9CFFFB48DF8BDCFE9ABECF" box="[247,295,152,176]" pageId="4" pageNumber="27">type</typeStatus>
specimen originally bears one white rectangular label with Esakis handwriting “
<taxonomicName id="96F04D15FF9CFFFB4C9C8BDCFF6ABEAB" authority="Esaki" authorityName="Esaki" class="Insecta" family="Reduviidae" genus="Epirodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="latithorax">
<emphasis id="6384EA84FF9CFFFB4C9C8BDCFA26BECF" box="[1204,1435,152,176]" italics="true" pageId="4" pageNumber="27">Epirodera latithorax</emphasis>
Esaki
</taxonomicName>
” and printed “Det. Teiso Esaki, 1929,” but 29 was crossed out by Esaki and replaced by 31. As noted by
<bibRefCitation id="35614B67FF9CFFFB48BF8B9BFE01BE88" author="Ishikawa" box="[151,444,223,247]" pageId="4" pageNumber="27" refString="Ishikawa, T. &amp; Yano, S. (1999) A new Polytoxus (Heteroptera, Reduviidae) from Taiwan. Japanese Journal of Systematic Entomology, 5, 341 - 345." type="journal article" year="1999">Ishikawa &amp; Yano (1999)</bibRefCitation>
,
<bibRefCitation id="35614B67FF9CFFFB49E48B9BFDDDBE88" author="Esaki" box="[460,608,223,247]" pageId="4" pageNumber="27" refString="Esaki, T. (1931) Hemiptera - Heteroptera von der Insel Botol - Tobago (Kotosho), Sud Formosa. Bulletin of the Biogeographical Society of Japan, 2, 209 - 220." type="journal article" year="1931">Esaki (1931)</bibRefCitation>
reported 26 species of true bugs from Botol-Tobago
<collectingCountry id="29E77606FF9CFFFB4CF68B9BFA9FBE88" box="[1246,1314,223,247]" name="Iceland" pageId="4" pageNumber="27">Island</collectingCountry>
(currently known as Lanyu Island) of
<collectingCountry id="29E77606FF9CFFFB49EF8A40FDA4BF63" box="[455,537,260,284]" name="Taiwan" pageId="4" pageNumber="27">Taiwan</collectingCountry>
based on the specimens collected by Tadao Kano. In Esakis paper, three species of reduviids, i.e.,
<taxonomicName id="96F04D15FF9CFFFB494A8A6CFD55BF40" authority="Ectrichodinae" authorityName="Ectrichodinae" box="[354,744,295,320]" class="Insecta" family="Reduviidae" genus="Ectrychotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="kanoi">
<emphasis id="6384EA84FF9CFFFB494A8A6CFD90BF3F" box="[354,557,296,320]" italics="true" pageId="4" pageNumber="27">Ectrychotes kanoi</emphasis>
(Ectrichodinae)
</taxonomicName>
,
<taxonomicName id="96F04D15FF9CFFFB4ADD8A6CFB28BF40" authority="Physoderinae" authorityName="Physoderinae" box="[757,1173,295,320]" class="Insecta" family="Reduviidae" genus="Epirodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="latithorax">
<emphasis id="6384EA84FF9CFFFB4ADD8A6CFC62BF3F" box="[757,991,296,320]" italics="true" pageId="4" pageNumber="27">Epirodera latithorax</emphasis>
(Physoderinae)
</taxonomicName>
, and
<taxonomicName id="96F04D15FF9CFFFB4CFB8A6CFEB6BF1B" authority="(Saicinae)" baseAuthorityName="Saicinae" class="Insecta" family="Reduviidae" genus="Polytoxus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="6384EA84FF9CFFFB4CFB8A6CFA21BF3F" box="[1235,1436,296,320]" italics="true" pageId="4" pageNumber="27">Polytoxus pallens</emphasis>
(Saicinae)
</taxonomicName>
, were reported under “in litteris” status. Those reduviids have never been properly described except for
<taxonomicName id="96F04D15FF9CFFFB48BF8A34FEE3BFF7" box="[151,350,368,392]" class="Insecta" family="Reduviidae" genus="Polytoxus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="pallens">
<emphasis id="6384EA84FF9CFFFB48BF8A34FEE3BFF7" box="[151,350,368,392]" italics="true" pageId="4" pageNumber="27">Polytoxus pallens</emphasis>
</taxonomicName>
, which was named by
<bibRefCitation id="35614B67FF9CFFFB4A4E8A2BFCC3BFF8" author="Ishikawa" box="[614,894,367,391]" pageId="4" pageNumber="27" refString="Ishikawa, T. &amp; Yano, S. (1999) A new Polytoxus (Heteroptera, Reduviidae) from Taiwan. Japanese Journal of Systematic Entomology, 5, 341 - 345." type="journal article" year="1999">Ishikawa &amp; Yano (1999)</bibRefCitation>
as
<taxonomicName id="96F04D15FF9CFFFB4B8E8A34FBE0BFF7" box="[934,1117,368,392]" class="Insecta" family="Reduviidae" genus="Polytoxus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="esakii">
<emphasis id="6384EA84FF9CFFFB4B8E8A34FBE0BFF7" box="[934,1117,368,392]" italics="true" pageId="4" pageNumber="27">Polytoxus esakii</emphasis>
</taxonomicName>
. Based on the data in
<bibRefCitation id="35614B67FF9CFFFB4D768A2BFF5CBFD3" author="Esaki" pageId="4" pageNumber="27" refString="Esaki, T. (1931) Hemiptera - Heteroptera von der Insel Botol - Tobago (Kotosho), Sud Formosa. Bulletin of the Biogeographical Society of Japan, 2, 209 - 220." type="journal article" year="1931">Esaki (1931)</bibRefCitation>
and
<bibRefCitation id="35614B67FF9CFFFB493F8AD0FD90BFD3" author="Ishikawa" box="[279,557,404,428]" pageId="4" pageNumber="27" refString="Ishikawa, T. &amp; Yano, S. (1999) A new Polytoxus (Heteroptera, Reduviidae) from Taiwan. Japanese Journal of Systematic Entomology, 5, 341 - 345." type="journal article" year="1999">Ishikawa &amp; Yano (1999)</bibRefCitation>
, we added a new label to the
<typeStatus id="8E4B8834FF9CFFFB4B5C8AD1FC19BFD2" box="[884,932,405,429]" pageId="4" pageNumber="27">type</typeStatus>
specimen with the collecting data as “
<collectingCountry id="29E77606FF9CFFFB4D788AD0FA25BFD3" box="[1360,1432,404,428]" name="China" pageId="4" pageNumber="27">China</collectingCountry>
,
<collectingCountry id="29E77606FF9CFFFB48BF8AF3FF54BFB0" box="[151,233,439,463]" name="Taiwan" pageId="4" pageNumber="27">Taiwan</collectingCountry>
Province, Botol-Tobago
<collectingCountry id="29E77606FF9CFFFB4A2B8AF3FDF7BFB0" box="[515,586,439,463]" name="Iceland" pageId="4" pageNumber="27">Island</collectingCountry>
, earlier than 1931, Tadao Kano leg.”
</paragraph>
</subSubSection>
<subSubSection id="19EA651DFF9CFFFB48ED8A9FFDA6BF8B" box="[197,539,475,500]" pageId="4" pageNumber="27" type="distribution">
<paragraph id="514F3696FF9CFFFB48ED8A9FFDA6BF8B" blockId="4.[151,1436,151,608]" box="[197,539,475,500]" pageId="4" pageNumber="27">
<emphasis id="6384EA84FF9CFFFB48ED8A9FFEE3BF8B" bold="true" box="[197,350,475,500]" pageId="4" pageNumber="27">Distribution.</emphasis>
<collectingCountry id="29E77606FF9CFFFB494D8A98FE15BF8B" box="[357,424,476,500]" name="China" pageId="4" pageNumber="27">China</collectingCountry>
(
<collectingCountry id="29E77606FF9CFFFB499F8A98FDADBF8B" box="[439,528,476,500]" name="Taiwan" pageId="4" pageNumber="27">Taiwan</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="19EA651DFF9CFFFB48ED8944FCB7BC43" pageId="4" pageNumber="27" type="etymology">
<paragraph id="514F3696FF9CFFFB48ED8944FCB7BC43" blockId="4.[151,1436,151,608]" pageId="4" pageNumber="27">
<emphasis id="6384EA84FF9CFFFB48ED8944FEF6BC67" bold="true" box="[197,331,512,536]" pageId="4" pageNumber="27">Etymology.</emphasis>
This species was named in honor of Dr. Teiso Esaki of Kyushu University for his contribution to heteropterology and the recognition of this new species.
</paragraph>
</subSubSection>
<subSubSection id="19EA651DFF9CFFFB48ED890CFA34BC20" box="[197,1417,583,608]" pageId="4" pageNumber="27" type="discussion">
<paragraph id="514F3696FF9CFFFB48ED890CFA34BC20" blockId="4.[151,1436,151,608]" box="[197,1417,583,608]" pageId="4" pageNumber="27">
<emphasis id="6384EA84FF9CFFFB48ED890CFE84BC1F" bold="true" box="[197,313,584,608]" pageId="4" pageNumber="27">Remarks.</emphasis>
This species may easily be distinguished from its congeners by the swollen anterior pronotal lobe.
</paragraph>
</subSubSection>
</treatment>
</document>