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<document id="AB785CCFFB5961E5311504ECBDDAC544" ID-DOI="10.5281/zenodo.5742183" ID-ISSN="1536-9307" ID-Zenodo-Dep="5742183" ID-ZooBank="18E6176F-7B73-434C-8DE2-782068C222BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638266808478" checkinUser="felipe" docAuthor="de Armas, Luis F. &amp; Teruel, Rolando" docDate="2021" docId="BB1287AE0310E113FEA99CB0E25BF87B" docLanguage="en" docName="Euscorpius.339.1-5.pdf" docOrigin="Euscorpius 339" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Centruroides concordia de Armas &amp; Teruel 2021, sp. n." docType="treatment" docVersion="3" lastPageNumber="3" masterDocId="472BFFD60314E117FFBE9D1DE74EFF98" masterDocTitle="The correct identity of Centruroides hoffmanni Armas, 1996 (Scorpiones: Buthidae), with the description of a new species from Chiapas, Mexico" masterLastPageNumber="5" masterPageNumber="1" pageNumber="3" updateTime="1699436824662" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="87068C89C287A414E8E27A7ED39AFC36" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E66FD5322035E7542B162A11CE347FB8">
<mods:title id="31DE6B06BA869592031CF2518F05706B">The correct identity of Centruroides hoffmanni Armas, 1996 (Scorpiones: Buthidae), with the description of a new species from Chiapas, Mexico</mods:title>
</mods:titleInfo>
<mods:name id="04D61926620201F2E535FAF8F9627C2E" type="personal">
<mods:role id="34C47F85D21F70A98789AD70C117AFB8">
<mods:roleTerm id="EE12C05F73A779CB74C704AD1A0317BC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A67321DFF29C2E24013F28746192A172">de Armas, Luis F.</mods:namePart>
</mods:name>
<mods:name id="F97061C759626091C594D5867EFA1988" type="personal">
<mods:role id="62739A330B42997A44610977A37E3FC8">
<mods:roleTerm id="2DD69FD7689E855DEA80C6174815A4EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E8B9B34A1F63229A2BCD85B6C23B6FA0">Teruel, Rolando</mods:namePart>
</mods:name>
<mods:typeOfResource id="0D9738402A8D8DD7188926B98DC83452">text</mods:typeOfResource>
<mods:relatedItem id="A9EEE6C84BC50FBD69B9AA3B7619C3A6" type="host">
<mods:titleInfo id="3F6993EA5A88F9E1AD341EAB1CE131A1">
<mods:title id="315B403A1C3BF41DEC2A74ADF96F5F8C">Euscorpius</mods:title>
</mods:titleInfo>
<mods:part id="A5849447793096A7E15BC9B9CF430199">
<mods:date id="DD7153EECA8990AA9654E920C7387080">2021</mods:date>
<mods:detail id="B3E09AFAAD58263209596189CAA796B1" type="volume">
<mods:number id="1803E6A63F9483AE4FDD920A7C39EFD8">339</mods:number>
</mods:detail>
<mods:extent id="169808D3EBAA405E83A57F6B2E60E0C3" unit="page">
<mods:start id="3F17231EEFC66C65AB05173F83A0A4F2">1</mods:start>
<mods:end id="F978B11B9E1AB258B6C7C3CBF921AE5F">5</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="AF607A61EBC3678DA2B1D90B78359B41">journal article</mods:classification>
<mods:identifier id="81A0FE374416E17CBFB8702E2FCBD4DB" type="DOI">10.5281/zenodo.5742183</mods:identifier>
<mods:identifier id="A15F8BFBBE5F09E73158BC5D63008198" type="ISSN">1536-9307</mods:identifier>
<mods:identifier id="4A5DC4A4C47EF6B5B19FC0C38E39B410" type="Zenodo-Dep">5742183</mods:identifier>
<mods:identifier id="CA23B2F5F0D33DA8C1D994FF4080A7A3" type="ZooBank">18E6176F-7B73-434C-8DE2-782068C222BA</mods:identifier>
</mods:mods>
<treatment id="BB1287AE0310E113FEA99CB0E25BF87B" LSID="urn:lsid:plazi:treatment:BB1287AE0310E113FEA99CB0E25BF87B" httpUri="http://treatment.plazi.org/id/BB1287AE0310E113FEA99CB0E25BF87B" lastPageNumber="3" pageId="4" pageNumber="3">
<subSubSection id="7BA165330310E113FEA99CB0E510FE5D" box="[279,606,429,454]" pageId="4" pageNumber="3" type="nomenclature">
<paragraph id="330436B80310E113FEA99CB0E510FE5D" blockId="4.[124,760,429,550]" box="[279,606,429,454]" pageId="4" pageNumber="3">
<heading id="684C81D40310E113FEA99CB0E510FE5D" box="[279,606,429,454]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="3" reason="2">
<taxonomicName id="F4BB4D3B0310E113FEA99CB0E555FE5D" authority="de Armas &amp; Teruel, 2021" authorityName="de Armas &amp; Teruel" authorityYear="2021" box="[279,539,429,453]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="concordia" status="sp. nov.">Centruroides concordia</taxonomicName>
<taxonomicNameLabel id="1AFC57D10310E113FD9C9CB3E510FE5D" box="[546,606,430,454]" pageId="4" pageNumber="3" rank="species">
<emphasis id="01CFEAAA0310E113FD9C9CB3E575FE5E" bold="true" box="[546,571,430,454]" pageId="4" pageNumber="3">sp</emphasis>
.
<emphasis id="01CFEAAA0310E113FDF69CB3E519FE5E" bold="true" box="[584,599,430,454]" pageId="4" pageNumber="3">n</emphasis>
.
</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7BA165330310E113FE3B9CD3E67FFC9E" pageId="4" pageNumber="3" type="description">
<paragraph id="330436B80310E113FE3B9CD3E5B6FD9E" blockId="4.[124,760,429,550]" pageId="4" pageNumber="3">
(
<figureCitation id="AB802A3D0310E113FE329CD3E6A6FE7E" box="[396,488,462,486]" captionStart="Figure 2" captionStartId="5.[113,181,1349,1371]" captionTargetBox="[113,1474,234,1342]" captionTargetId="figure-289@5.[113,1474,234,1342]" captionTargetPageId="5" captionText="Figure 2. Geographical distribution of the authentic Centruroides hoffmanni (circles) and C. concordia sp. n. (triangles), with the tYpe localities indicated in red. Image frame represents 3,800 x 1,550 km (a) and 600 x 245 km (b)." figureDoi="http://doi.org/10.5281/zenodo.5742187" httpUri="https://zenodo.org/record/5742187/files/figure.png" pageId="4" pageNumber="3">Figure 2</figureCitation>
)
<emphasis id="01CFEAAA0310E113FFC29CF3E5B6FD9E" bold="true" box="[124,760,494,518]" pageId="4" pageNumber="3">http://zoobank.org/urn:lsid:zoobank.org:act:F3624B67-</emphasis>
</paragraph>
<paragraph id="330436B80310E113FF409F13E538FDBE" blockId="4.[124,760,429,550]" box="[254,630,526,550]" pageId="4" pageNumber="3">
<emphasis id="01CFEAAA0310E113FF409F13E538FDBE" bold="true" box="[254,630,526,550]" pageId="4" pageNumber="3">996A-43F8-B898-77F14F2B2D13</emphasis>
</paragraph>
<paragraph id="330436B80310E113FFCF9F53E67FFC9E" blockId="4.[113,772,590,774]" pageId="4" pageNumber="3">
<taxonomicName id="F4BB4D3B0310E113FFCF9F53E6C6FDFE" authorityName="Armas" authorityYear="1996" box="[113,392,590,614]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="01CFEAAA0310E113FFCF9F53E6C6FDFE" box="[113,392,590,614]" italics="true" pageId="4" pageNumber="3">Centruroides hoffmanni</emphasis>
</taxonomicName>
[misidentification]: Goodman, Prendini &amp; Esposito, 2021: 3, 6–7, 9–10, 13, 15; fig. 2; tabs. 1–2. Goodman, Prendini, Francke &amp; Esposito, 2021: 8, 53–55, figs. 2, 4, 7a–b, 10 a–b, 15a, 16a, 17c–f, 18c–f, 19c–f, 20c–f, 21c–f, 22c–f, 23c–f, 24c–f, 25c–f, 38–39; tabs. 1, 8, 10.
</paragraph>
</subSubSection>
<subSubSection id="7BA165330310E113FFCF9E33E5A2FBFE" pageId="4" pageNumber="3" type="materials_examined">
<paragraph id="330436B80310E113FFCF9E33E6FFFC1E" blockId="4.[113,771,814,902]" pageId="4" pageNumber="3">
<materialsCitation id="83D33CE50310E113FFCF9E33E608FCDE" box="[113,326,814,838]" collectionCode="LOCALITY, AND" pageId="4" pageNumber="3" specimenCount="1" typeStatus="holotype">
<typeStatus id="EC00881A0310E113FFCF9E33E7E9FCDD" box="[113,167,814,837]" pageId="4" pageNumber="3">TYPE</typeStatus>
<collectionCode id="55AAAE7D0310E113FF139E33E608FCDE" box="[173,326,814,838]" pageId="4" pageNumber="3">LOCALITY AND</collectionCode>
</materialsCitation>
<materialsCitation id="83D33CE50310E113FEF29E33E6E3FC1E" collectionCode="REPOSITORY" collectorName="La Concordia" country="Mexico" location="Chiapas State" pageId="4" pageNumber="3" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<typeStatus id="EC00881A0310E113FEF29E33E630FCDD" box="[332,382,814,837]" pageId="4" pageNumber="3">TYPE</typeStatus>
<collectionCode id="55AAAE7D0310E113FE3A9E33E54EFCDE" box="[388,512,814,838]" pageId="4" pageNumber="3">REPOSITORY</collectionCode>
.
<emphasis id="01CFEAAA0310E113FDB69E33E515FCDE" bold="true" box="[520,603,814,838]" pageId="4" pageNumber="3">
<collectingCountry id="4BAC76280310E113FDB69E33E515FCDE" box="[520,603,814,838]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
</emphasis>
,
<emphasis id="01CFEAAA0310E113FDD69E33E5B2FCDD" box="[616,764,814,838]" italics="true" pageId="4" pageNumber="3">
<collectingRegion id="F17FF85A0310E113FDD69E33E5B2FCDD" box="[616,764,814,838]" country="Mexico" name="Chiapas" pageId="4" pageNumber="3">Chiapas State</collectingRegion>
</emphasis>
,
<collectorName id="9E4E536E0310E113FFCF9E53E643FCFE" box="[113,269,846,870]" pageId="4" pageNumber="3">La Concordia</collectorName>
MunicipalitY, Villa Corzo-La Tigrilla, San Julián, Revolución Mexicana
</materialsCitation>
.
</paragraph>
<paragraph id="330436B80310E113FFCF9EB3E5A2FBFE" blockId="4.[113,771,942,1126]" pageId="4" pageNumber="3">
<materialsCitation id="83D33CE50310E113FFCF9EB3E5D9FC5E" box="[113,663,942,966]" collectionCode="TYPE, MATERIAL, CNAN" country="Mexico" pageId="4" pageNumber="3" specimenCode="SC3998" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<collectionCode id="55AAAE7D0310E113FFCF9EB3E655FC5E" box="[113,283,942,966]" pageId="4" pageNumber="3">TYPE MATERIAL</collectionCode>
.
<specimenCount id="25BDFD310310E113FE8F9EB3E61CFC5E" box="[305,338,942,966]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
<typeStatus id="EC00881A0310E113FEDC9EB3E6F1FC5E" box="[354,447,942,966]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
(
<collectionCode id="55AAAE7D0310E113FE689EB3E56DFC5E" box="[470,547,942,966]" pageId="4" pageNumber="3">CNAN</collectionCode>
<specimenCode id="631D9EC30310E113FD8D9EB3E5C1FC5E" box="[563,655,942,966]" collectionCode="CNAN" pageId="4" pageNumber="3">SC3998</specimenCode>
)
</materialsCitation>
:
<materialsCitation id="83D33CE50310E113FD179EB3E6AAFBDD" collectingDate="2007-04-17" collectionCode="CNAN" collectorName="C. MaYorga &amp; G. Ortega &amp; L. Cervantes." country="Mexico" county="La Concordia" elevation="544" latitude="16.0" location="La Tigrilla" longLatPrecision="21" longitude="-92.84639" municipality="Villa" pageId="4" pageNumber="3" specimenCount="1" stateProvince="Chiapas" typeStatus="paratype">
<emphasis id="01CFEAAA0310E113FD179EB3E5B2FC5E" bold="true" box="[681,764,942,966]" pageId="4" pageNumber="3">
<collectingCountry id="4BAC76280310E113FD179EB3E5B2FC5E" box="[681,764,942,966]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
</emphasis>
,
<emphasis id="01CFEAAA0310E113FFCF9ED3E65EFC7D" box="[113,272,974,998]" italics="true" pageId="4" pageNumber="3">
<collectingRegion id="F17FF85A0310E113FFCF9ED3E65EFC7D" box="[113,272,974,998]" country="Mexico" name="Chiapas" pageId="4" pageNumber="3">Chiapas State</collectingRegion>
</emphasis>
,
<collectingCounty id="DA654E340310E113FE969ED3E68AFC7E" box="[296,452,974,998]" pageId="4" pageNumber="3">La Concordia</collectingCounty>
MunicipalitY,
<collectingMunicipality id="D360ACC20310E113FDCA9ED3E5E6FC7E" box="[628,680,974,998]" pageId="4" pageNumber="3">Villa</collectingMunicipality>
Corzo-
<location id="366460630310E113FFCF9EF3E7A5FB9E" LSID="urn:lsid:plazi:treatment:BB1287AE0310E113FEA99CB0E25BF87B:366460630310E113FFCF9EF3E7A5FB9E" box="[113,235,1006,1030]" country="Mexico" county="La Concordia" latitude="16.0" longLatPrecision="21" longitude="-92.84639" municipality="Villa" name="La Tigrilla" pageId="4" pageNumber="3" stateProvince="Chiapas">La Tigrilla</location>
,
<location id="366460630310E113FF459EF3E63EFB9E" LSID="urn:lsid:plazi:treatment:BB1287AE0310E113FEA99CB0E25BF87B:366460630310E113FF459EF3E63EFB9E" box="[251,368,1006,1030]" country="Mexico" county="La Concordia" latitude="16.0" longLatPrecision="21" longitude="-92.84639" municipality="Villa" name="San Julian" pageId="4" pageNumber="3" stateProvince="Chiapas">San Julián</location>
,
<location id="366460630310E113FE3F9EF3E53FFB9E" LSID="urn:lsid:plazi:treatment:BB1287AE0310E113FEA99CB0E25BF87B:366460630310E113FE3F9EF3E53FFB9E" box="[385,625,1006,1030]" country="Mexico" county="La Concordia" latitude="16.0" longLatPrecision="21" longitude="-92.84639" municipality="Villa" name="Revolucion Mexicana" pageId="4" pageNumber="3" stateProvince="Chiapas">Revolución Mexicana</location>
(
<geoCoordinate id="568F507F0310E113FD3A9EF3E44DFB9E" box="[644,771,1006,1030]" degrees="16" direction="north" minutes="00" orientation="latitude" pageId="4" pageNumber="3" precision="15" seconds="00" value="16.0">16°00'00&quot;N</geoCoordinate>
<geoCoordinate id="568F507F0310E113FFCF9913E7B6FBBE" box="[113,248,1038,1062]" degrees="92" direction="west" minutes="50" orientation="longitude" pageId="4" pageNumber="3" precision="15" seconds="47" value="-92.84639">92°50'47&quot;W</geoCoordinate>
),
<quantity id="F4439B5D0310E113FEB49913E603FBBD" box="[266,333,1038,1062]" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" pageId="4" pageNumber="3" unit="m" value="544.0">
<elevation id="B896D18B0310E113FEB49913E603FBBD" box="[266,333,1038,1062]" metricMagnitude="2" metricUnit="m" metricValue="5.4399999999999995" pageId="4" pageNumber="3" unit="m" value="544.0">544 m</elevation>
</quantity>
a. s. l,
<date id="470510780310E113FE2A9913E567FBBE" box="[404,553,1038,1062]" pageId="4" pageNumber="3" value="2007-04-17">
<collectingDate id="5741E9900310E113FE2A9913E567FBBE" box="[404,553,1038,1062]" pageId="4" pageNumber="3" value="2007-04-17">17 April 2007</collectingDate>
</date>
, leg.
<collectorName id="9E4E536E0310E113FDE19913E5AEFBBE" box="[607,736,1038,1062]" pageId="4" pageNumber="3">C. MaYorga</collectorName>
,
<collectorName id="9E4E536E0310E113FD579913E7F4FBDE" pageId="4" pageNumber="3">G. Ortega</collectorName>
&amp;
<collectorName id="9E4E536E0310E113FF619933E63FFBDE" box="[223,369,1070,1094]" pageId="4" pageNumber="3">L. Cervantes.</collectorName>
<typeStatus id="EC00881A0310E113FEC69933E6AAFBDD" box="[376,484,1070,1093]" pageId="4" pageNumber="3" type="paratype">ParatYpes</typeStatus>
</materialsCitation>
: all other specimens
<materialsCitation id="83D33CE50310E113FD779933E5A9FBFE" collectionCode="CNAN" collectorName="Goodman" country="Mexico" location="Material" pageId="4" pageNumber="3" specimenCount="1" typeStatus="paratype">
listed under “
<location id="366460630310E113FF009953E655FBFE" LSID="urn:lsid:plazi:treatment:BB1287AE0310E113FEA99CB0E25BF87B:366460630310E113FF009953E655FBFE" box="[190,283,1102,1126]" country="Mexico" name="Material" pageId="4" pageNumber="3">Material</location>
examined” bY
<collectorName id="9E4E536E0310E113FE039953E568FBFE" box="[445,550,1102,1126]" pageId="4" pageNumber="3">Goodman</collectorName>
et al. (2021b: 55)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7BA165330310E113FFCF9993E533FB7E" pageId="4" pageNumber="3" type="etymology">
<paragraph id="330436B80310E113FFCF9993E533FB7E" blockId="4.[113,771,1166,1254]" pageId="4" pageNumber="3">
ETYMOLOGY. The specific name is an indeclinable noun in apposition, taken straight from the Spanish name of the municipalitY from which the
<typeStatus id="EC00881A0310E113FE1699D3E54BFB7E" box="[424,517,1230,1254]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
originates.
</paragraph>
</subSubSection>
<subSubSection id="7BA165330310E113FFCF9813E639F91E" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="330436B80310E113FFCF9813E639F91E" blockId="4.[113,771,1294,1670]" pageId="4" pageNumber="3">
DIAGNOSIS. A medium-sized species (
<quantity id="F4439B5D0310E113FD9B9813E5EBFABE" box="[549,677,1294,1318]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" metricValueMax="6.0" metricValueMin="4.4" pageId="4" pageNumber="3" unit="mm" value="52.0" valueMax="60.0" valueMin="44.0">44–60 mm</quantity>
) of the “
<emphasis id="01CFEAAA0310E113FFC39833E79CFADE" box="[125,210,1326,1350]" italics="true" pageId="4" pageNumber="3">thorellii</emphasis>
” group of the genus
<taxonomicName id="F4BB4D3B0310E113FE6B9833E52FFADE" authorityName="Marx" authorityYear="1890" box="[469,609,1326,1350]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="01CFEAAA0310E113FE6B9833E52FFADE" box="[469,609,1326,1350]" italics="true" pageId="4" pageNumber="3">Centruroides</emphasis>
</taxonomicName>
, characterized for the following combination of characters:
<emphasis id="01CFEAAA0310E113FDE39853E53AFAFE" bold="true" box="[605,628,1358,1382]" pageId="4" pageNumber="3">1)</emphasis>
tergites with complete median carinae;
<emphasis id="01CFEAAA0310E113FE2D9873E6E4FA1E" bold="true" box="[403,426,1390,1414]" pageId="4" pageNumber="3">2)</emphasis>
pectinal tooth counts
<quantity id="F4439B5D0310E113FD1C9873E44DFA1E" box="[674,771,1390,1414]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="4" pageNumber="3" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
both sexes;
<emphasis id="01CFEAAA0310E113FF499893E643FA3E" bold="true" box="[247,269,1422,1446]" pageId="4" pageNumber="3">3)</emphasis>
metasomal segments I-II with strong ventral submedian carinae;
<emphasis id="01CFEAAA0310E113FEE998B3E623FA5E" bold="true" box="[343,365,1454,1478]" pageId="4" pageNumber="3">4)</emphasis>
male carapace 0.91–1.00 times as long as wide;
<emphasis id="01CFEAAA0310E113FEB498D3E66EFA7E" bold="true" box="[266,288,1486,1510]" pageId="4" pageNumber="3">5)</emphasis>
carapace with posterior median carinae well developed, granular;
<emphasis id="01CFEAAA0310E113FEEC98F3E626F99E" bold="true" box="[338,360,1518,1542]" pageId="4" pageNumber="3">6)</emphasis>
pedipalp chelae with prodorsal carina;
<emphasis id="01CFEAAA0310E113FFCF9B13E7C6F9BE" bold="true" box="[113,136,1550,1574]" pageId="4" pageNumber="3">7)</emphasis>
male telson vesicle elongate, 2.1–2.9 times longer than wide. Its closest relative seems to be
<taxonomicName id="F4BB4D3B0310E113FDA69B33E54AF9FE" authority="Goodman, Prendini, Francke &amp; Esposito, 2001" authorityName="Goodman, Prendini, Francke &amp; Esposito" authorityYear="2001" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="chanae">
<emphasis id="01CFEAAA0310E113FDA69B33E5C7F9DE" box="[536,649,1582,1606]" italics="true" pageId="4" pageNumber="3">C. chanae</emphasis>
Goodman, Prendini, Francke &amp; Esposito, 2001
</taxonomicName>
; for a comparison, see Goodman et al. (2021b).
</paragraph>
</subSubSection>
<subSubSection id="7BA165330310E113FFCF9BB3E24EFE3A" pageId="4" pageNumber="3" type="discussion">
<paragraph id="330436B80310E113FFCF9BB3E5BBF87D" blockId="4.[113,772,1710,2022]" pageId="4" pageNumber="3">
COMMENTS. Below we will discuss in detail the complete mismatch between the
<typeStatus id="EC00881A0310E113FEC39BD3E694F97E" box="[381,474,1742,1766]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
of
<taxonomicName id="F4BB4D3B0310E113FDB39BD3E5D1F97E" authorityName="Armas" authorityYear="1996" box="[525,671,1742,1766]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="01CFEAAA0310E113FDB39BD3E5D1F97E" box="[525,671,1742,1766]" italics="true" pageId="4" pageNumber="3">C. hoffmanni</emphasis>
</taxonomicName>
and the species identified as such bY Goodman (2021a, 2021b):
<emphasis id="01CFEAAA0310E113FFCF9A13E65DF8BE" bold="true" box="[113,275,1806,1830]" pageId="4" pageNumber="3">Color pattern.</emphasis>
The
<typeStatus id="EC00881A0310E113FEF39A13E6E6F8BE" box="[333,424,1806,1830]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
has the carapace with large and sYmmetrical dark blotches and the tergites with two broad and solid dark longitudinal stripes (see
<bibRefCitation id="572A4B490310E113FE559A53E5EDF8FE" author="ARMAS" box="[491,675,1870,1894]" pageId="4" pageNumber="3" pagination="21 - 33" refId="ref2553" refString="ARMAS, L. F. DE. 1996. Presencia de Centruroides schmidti Sissom en el sureste de Mexico Y descripcion de dos especies nuevas (Scorpiones: Buthidae). Revista Nicaraguense de Entomologia, 36: 21 - 33." type="journal article" year="1996">Armas, 1996: 29</bibRefCitation>
), exactlY as in our specimens conspecific with it (see Martín-Frías et al., 2005: 2, fig. 1). On the other hand, Goodman, Prendini, Francke &amp; Esposito (2021: 68–69, figs. 38–39) depicted the carapace and tergites denselY and irregularlY variegated in the second taxon.
</paragraph>
<paragraph id="330436B80310E113FC8E9DF7E24EFE3A" blockId="4.[816,1475,234,1827]" pageId="4" pageNumber="3">
<emphasis id="01CFEAAA0310E113FC8E9DF7E362FE9B" bold="true" box="[816,1068,234,259]" pageId="4" pageNumber="3">Pectinal tooth counts.</emphasis>
The
<typeStatus id="EC00881A0310E113FBD99DF6E38AFE9B" box="[1127,1220,235,259]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
has 20/21 (see
<bibRefCitation id="572A4B490310E113FACA9DF6E4D9FEBB" author="ARMAS" pageId="4" pageNumber="3" pagination="21 - 33" refId="ref2553" refString="ARMAS, L. F. DE. 1996. Presencia de Centruroides schmidti Sissom en el sureste de Mexico Y descripcion de dos especies nuevas (Scorpiones: Buthidae). Revista Nicaraguense de Entomologia, 36: 21 - 33." type="journal article" year="1996">Armas, 1996: 30</bibRefCitation>
), while our conspecific specimens have
<quantity id="F4439B5D0310E113FAE19C16E28CFEBB" box="[1375,1474,267,291]" metricMagnitude="-1" metricUnit="m" metricValue="5.715" metricValueMax="6.096" metricValueMin="5.334" pageId="4" pageNumber="3" unit="in" value="22.5" valueMax="24.0" valueMin="21.0">21–24 in</quantity>
males and
<quantity id="F4439B5D0310E113FC199C36E349FEDB" box="[935,1031,299,323]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="4" pageNumber="3" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20–22 in</quantity>
females, as stated bY Martín-Frías et al. (2005: 5; tab. 3). On the other hand, Goodman et al. (2021b: 48; tab. 8) recorded non-overlapping and much lower counts of
<quantity id="F4439B5D0310E113FCF39C97E4E2FE3B" box="[845,940,394,419]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="3.81" metricValueMin="3.302" pageId="4" pageNumber="3" unit="in" value="14.0" valueMax="15.0" valueMin="13.0">13–15 in</quantity>
both sexes of the second taxon.
</paragraph>
</subSubSection>
<subSubSection id="7BA165330310E113FC8E9CB6E496F8BA" pageId="4" pageNumber="3" type="description">
<paragraph id="330436B80310E113FC8E9CB6E369FCBB" blockId="4.[816,1475,234,1827]" pageId="4" pageNumber="3">
<emphasis id="01CFEAAA0310E113FC8E9CB6E417FE7B" bold="true" pageId="4" pageNumber="3">Length / width of metasomal segments I / II / III / IV / V.</emphasis>
The
<typeStatus id="EC00881A0310E113FC259CD6E4B4FE7B" box="[923,1018,459,483]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
female has the following ratios: 1.12 / 1.36 / 1.49 / 1.74 / 2.12 (calculated from
<bibRefCitation id="572A4B490310E113FABF9CF6E2F3FD9B" author="ARMAS" box="[1281,1469,491,515]" pageId="4" pageNumber="3" pagination="21 - 33" refId="ref2553" refString="ARMAS, L. F. DE. 1996. Presencia de Centruroides schmidti Sissom en el sureste de Mexico Y descripcion de dos especies nuevas (Scorpiones: Buthidae). Revista Nicaraguense de Entomologia, 36: 21 - 33." type="journal article" year="1996">Armas, 1996: 28</bibRefCitation>
, tab. 3), because of being immature, slightlY shorter than our conspecific adult females with 1.15–1.23 / 1.40–1.59 / 1.45–1.73 / 1.76–1.95 / 2.14–2.35 (see Martín-Frías et al., 2005: 4, tabs. 1–2, enhanced with our new calculations). On the other hand, Goodman et al. (2021b: 48; tab. 8) gave precise measurements of adult females of the second taxon that allowed us to calculate the following non-overlapping and much longer ratios: 1.28–1.50 / 1.94–2.20 / 2.27–2.40 / 2.34–2.86 / 2.73–3.31.
</paragraph>
<paragraph id="330436B80310E113FC8E9E36E43DFBDA" blockId="4.[816,1475,234,1827]" pageId="4" pageNumber="3">
<emphasis id="01CFEAAA0310E113FC8E9E36E485FCDB" bold="true" box="[816,971,811,835]" pageId="4" pageNumber="3">Telson shape.</emphasis>
Despite being an immature, the
<typeStatus id="EC00881A0310E113FA869E36E2DBFCDB" box="[1336,1429,811,835]" pageId="4" pageNumber="3" type="holotype">holotYpe</typeStatus>
has the vesicle short and bulbose and the subaculear tubercle spiniform (
<bibRefCitation id="572A4B490310E113FC169E76E32CFC1B" author="ARMAS" box="[936,1122,875,899]" pageId="4" pageNumber="3" pagination="21 - 33" refId="ref2553" refString="ARMAS, L. F. DE. 1996. Presencia de Centruroides schmidti Sissom en el sureste de Mexico Y descripcion de dos especies nuevas (Scorpiones: Buthidae). Revista Nicaraguense de Entomologia, 36: 21 - 33." type="journal article" year="1996">Armas, 1996: 30</bibRefCitation>
, fig. 8), just like our conspecific specimens (Martín-Frías et al., 2005: 2–3; figs. 5–6, 10; see also Santibáñez-López &amp; Ponce-Saavedra, 2009: 328, fig. 11). On the other hand, Goodman et al. (2021b: 51–52, figs. 24c,f, 25c,f) depicted the vesicle elongate and slender and the subaculear tubercle triangular and compressed in the second taxon.
</paragraph>
<paragraph id="330436B80310E113FC8E9956E305FA3B" blockId="4.[816,1475,234,1827]" pageId="4" pageNumber="3">
All differences listed above, as well as others well visible even to unaided eYe, are clearlY shown in the
<figureCitation id="AB802A3D0310E113FAA69976E23CFB1B" box="[1304,1394,1131,1155]" captionStart="Figure 2" captionStartId="5.[113,181,1349,1371]" captionTargetBox="[113,1474,234,1342]" captionTargetId="figure-289@5.[113,1474,234,1342]" captionTargetPageId="5" captionText="Figure 2. Geographical distribution of the authentic Centruroides hoffmanni (circles) and C. concordia sp. n. (triangles), with the tYpe localities indicated in red. Image frame represents 3,800 x 1,550 km (a) and 600 x 245 km (b)." figureDoi="http://doi.org/10.5281/zenodo.5742187" httpUri="https://zenodo.org/record/5742187/files/figure.png" pageId="4" pageNumber="3">Figure 2</figureCitation>
herein, which includes a topotYpe of the authentic
<taxonomicName id="F4BB4D3B0310E113FB419996E2C5FB3B" authorityName="Armas" authorityYear="1996" box="[1279,1419,1163,1187]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="01CFEAAA0310E113FB419996E2C5FB3B" box="[1279,1419,1163,1187]" italics="true" pageId="4" pageNumber="3">C. hoffmanni</emphasis>
</taxonomicName>
with the following collecting data:
<collectingCountry id="4BAC76280310E113FBCC99B6E388FB5B" box="[1138,1222,1195,1219]" name="Mexico" pageId="4" pageNumber="3">Mexico</collectingCountry>
,
<collectingRegion id="F17FF85A0310E113FB6C99B6E229FB5B" box="[1234,1383,1195,1219]" country="Mexico" name="Chiapas" pageId="4" pageNumber="3">Chiapas State</collectingRegion>
, Arriaga MunicipalitY, La Gloria Farm, collected inside inhabited house bY public health staff, no date, leg. J. G. Baldazo (
<specimenCount id="25BDFD310310E113FAE699F6E239FA9B" box="[1368,1399,1259,1283]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
RTO). The interested reader will easilY realize its perfect match to the figures published bY
<bibRefCitation id="572A4B490310E113FBF49836E3A8FADB" author="ARMAS" box="[1098,1254,1323,1347]" pageId="4" pageNumber="3" pagination="21 - 33" refId="ref2553" refString="ARMAS, L. F. DE. 1996. Presencia de Centruroides schmidti Sissom en el sureste de Mexico Y descripcion de dos especies nuevas (Scorpiones: Buthidae). Revista Nicaraguense de Entomologia, 36: 21 - 33." type="journal article" year="1996">Armas (1996)</bibRefCitation>
, Martín-Frías et al. (2005) and Santibáñez-López &amp; Ponce-Saavedra (2009) on one hand, and its absolute contradiction to those bY Goodman et al. (2021b) on the other.
</paragraph>
<paragraph id="330436B80310E113FC8E98B6E496F8BA" blockId="4.[816,1475,234,1827]" pageId="4" pageNumber="3">
With respect to the infrageneric affiliation of the authentic
<taxonomicName id="F4BB4D3B0310E113FA1798B6E4EEFA7B" authorityName="Armas" authorityYear="1996" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="01CFEAAA0310E113FA1798B6E4EEFA7B" italics="true" pageId="4" pageNumber="3">C. hoffmanni</emphasis>
</taxonomicName>
, the four diagnostic characters discussed above are enough to demonstrate beYond anY reasonable doubt that it is not a member of the “
<emphasis id="01CFEAAA0310E113FB109B16E249F9BB" box="[1198,1287,1547,1571]" italics="true" pageId="4" pageNumber="3">thorellii</emphasis>
” species-group. As first suggested implicitlY bY Martín-Frías et al. (2005) and then formallY established bY
<bibRefCitation id="572A4B490310E113FB0C9B56E2DEF9FA" author="TERUEL" box="[1202,1424,1610,1635]" pageId="4" pageNumber="3" pagination="3 - 14" refId="ref2944" refString="TERUEL, R., F. KOVARIK, J. G. BALDAZO-MONSIVAIS &amp; D. HOFEREK. 2015. A new species of Centruroides of the &quot; nigrovariatus &quot; group (Scorpiones: Buthidae) from southern Mexico. Revista Iberica de Aracnologia, 26: 3 - 14." type="journal article" year="2015">Teruel et al. (2015)</bibRefCitation>
and followed bY Santibáñez-López &amp; Ponce-Saavedra (2009), it belongs in the “
<emphasis id="01CFEAAA0310E113FBB69B96E3D1F93A" box="[1032,1183,1675,1698]" italics="true" pageId="4" pageNumber="3">nigrovariatus</emphasis>
” species-group, where it is most closelY related to
<taxonomicName id="F4BB4D3B0310E113FBE39BB6E424F97B" authority="Hoffmann, 1932" authorityName="Hoffmann" authorityYear="1932" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="baergi">
<emphasis id="01CFEAAA0310E113FBE39BB6E20FF95B" box="[1117,1345,1707,1731]" italics="true" pageId="4" pageNumber="3">Centruroides baergi</emphasis>
Hoffmann, 1932
</taxonomicName>
,
<taxonomicName id="F4BB4D3B0310E113FCC09BD6E429F89B" authority="Santibanez-Lopez &amp; Contreras-Felix, 2013" authorityName="Santibanez-Lopez &amp; Contreras-Felix" authorityYear="2013" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="franckei">
<emphasis id="01CFEAAA0310E113FCC09BD6E34FF97B" box="[894,1025,1739,1763]" italics="true" pageId="4" pageNumber="3">C. franckei</emphasis>
Santibáñez-López &amp; Contreras-Félix, 2013
</taxonomicName>
and
<taxonomicName id="F4BB4D3B0310E113FC209BF6E49AF8BA" authority="Teruel, Kovarik, Baldazo-Monsivais &amp; Hoferek, 2015" authorityName="Teruel, Kovarik, Baldazo-Monsivais &amp; Hoferek" authorityYear="2015" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="poncei">
<emphasis id="01CFEAAA0310E113FC209BF6E347F89A" box="[926,1033,1771,1794]" italics="true" pageId="4" pageNumber="3">C. poncei</emphasis>
Teruel, Kovařík, Baldazo-Monsivais &amp; Hoferek, 2015
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="7BA165330310E113FC8E9A56E25BF87B" pageId="4" pageNumber="3" type="discussion">
<paragraph id="330436B80310E113FC8E9A56E252F81B" blockId="4.[816,1474,1867,2019]" pageId="4" pageNumber="3">REMARKS. The present description is made in full agreement with Article 13.1.2 of the Code (ICZN, 1999).</paragraph>
<paragraph id="330436B80310E113FCDE9A96E25BF87B" blockId="4.[816,1474,1867,2019]" pageId="4" pageNumber="3">
For additional data of this species (measurements, pectinal tooth counts, illustrations, ecologY and distribution), see Goodman et al. (2021b: as
<taxonomicName id="F4BB4D3B0310E113FBC49AD6E24BF87B" authorityName="Armas" authorityYear="1996" box="[1146,1285,1995,2019]" class="Insecta" family="Cerambycidae" genus="Centruroides" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="hoffmanni">
<emphasis id="01CFEAAA0310E113FBC49AD6E24BF87B" box="[1146,1285,1995,2019]" italics="true" pageId="4" pageNumber="3">C. hoffmanni</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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