236 lines
35 KiB
XML
236 lines
35 KiB
XML
<document id="AC7CB934F55AB4F85A67B596ED75747D" ID-DOI="10.5281/zenodo.210858" ID-GBIF-Dataset="6c13241a-b0ff-420a-acf0-8b9cbd3b9703" ID-ISSN="1175-5326" ID-Zenodo-Dep="210858" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460478186346" checkinUser="plazi" docAuthor="Eldredge, Taro" docDate="2012" docId="973687D6FF86663CFF1F34B4A0ECFC65" docLanguage="en" docName="zt03300p054.pdf" docOrigin="Zootaxa 3300" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Drusilla nearctica Eldredge, 2012, new species" docType="treatment" docVersion="9" lastPageNumber="50" masterDocId="6B0FFFAEFF856639FF883423A44DFFA2" masterDocTitle="Review of the Nearctic species of Drusilla Leach (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="54" masterPageNumber="45" pageNumber="48" updateTime="1698307154067" updateUser="plazi">
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<mods:mods id="E9D37EC7D543B1CF570B3DAD38DACADA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="07887F9269FC075F976EA0DACC8ED1DF">
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<mods:title id="22968778051ECB6669263B7955FCFE8D">Review of the Nearctic species of Drusilla Leach (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="5C60CD5FB267B551488DD5CA27C736F1" type="personal">
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<mods:roleTerm id="8F12607FE201E2906BA681FE70401FB9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E329D731BA5963B90C6674BC41501D73">Eldredge, Taro</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="20C96A4406E17674D373FEC8D8EBD363">text</mods:typeOfResource>
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<mods:titleInfo id="FE89C58F6A2C8020716480AD37D207E8">
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<mods:title id="DCAA4FF35BA4F6C581637CCC21E3DA51">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BD4331AEEFF137050E0DD3DD0A18045A">
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<mods:date id="C715E37751EF0D6835F6D6691FD24B3C">2012</mods:date>
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<mods:detail id="403D4B075B7278D93D2A6EB7AFA532DA" type="volume">
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<mods:number id="E549C4DF974680236B3B166273C51CCA">3300</mods:number>
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</mods:detail>
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<mods:extent id="06CA3A817CE6E5EB9A3A6349338F532C" unit="page">
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<mods:start id="637D6A0EDA93A374744450180782CBE4">45</mods:start>
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<mods:end id="9E5D54B38DAAFCDA9AA7F8D8362CBEF0">54</mods:end>
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<mods:classification id="01426167FC7AF39F96CF6035E5D200D8">journal article</mods:classification>
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<mods:identifier id="F5E621D53FE7D6B48286F66ECDE47083" type="DOI">10.5281/zenodo.210858</mods:identifier>
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<mods:identifier id="4065CF808E9EB159CA2B0720AA86ADEC" type="GBIF-Dataset">6c13241a-b0ff-420a-acf0-8b9cbd3b9703</mods:identifier>
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<mods:identifier id="8F90877746B1647748EDE76673AEBD7E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="68B80321E5D3485C443AB9ED40B43213" type="Zenodo-Dep">210858</mods:identifier>
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<treatment id="973687D6FF86663CFF1F34B4A0ECFC65" ID-DOI="http://doi.org/10.5281/zenodo.6176688" ID-GBIF-Taxon="119516767" ID-Zenodo-Dep="6176688" LSID="urn:lsid:plazi:treatment:973687D6FF86663CFF1F34B4A0ECFC65" httpUri="http://treatment.plazi.org/id/973687D6FF86663CFF1F34B4A0ECFC65" lastPageId="5" lastPageNumber="50" pageId="3" pageNumber="48">
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<subSubSection id="5785654BFF86663AFF1F34B4A656FF12" box="[151,539,151,176]" pageId="3" pageNumber="48" type="nomenclature">
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<paragraph id="1F2036C0FF86663AFF1F34B4A656FF12" blockId="3.[151,539,151,212]" box="[151,539,151,176]" pageId="3" pageNumber="48">
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<heading id="446881ACFF86663AFF1F34B4A656FF12" bold="true" box="[151,539,151,176]" fontSize="10" level="2" pageId="3" pageNumber="48" reason="2">
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<emphasis id="2DEBEAD2FF86663AFF1F34B4A656FF12" bold="true" box="[151,539,151,176]" pageId="3" pageNumber="48">
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2.
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<taxonomicName id="D89F4D43FF86663AFF3B34B4A5C9FF12" ID-CoL="37TGR" box="[179,388,151,176]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="nearctica" status="sp. nov.">
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<emphasis id="2DEBEAD2FF86663AFF3B34B4A5C9FF12" bold="true" box="[179,388,151,176]" italics="true" pageId="3" pageNumber="48">Drusilla nearctica</emphasis>
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</taxonomicName>
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,
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||
<taxonomicNameLabel id="36D857A9FF86663AFE1A34BBA656FF12" box="[402,539,151,176]" pageId="3" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5785654BFF86663AFF1F349FA5E1FF76" box="[151,428,188,212]" pageId="3" pageNumber="48" type="description">
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<paragraph id="1F2036C0FF86663AFF1F349FA5E1FF76" blockId="3.[151,539,151,212]" box="[151,428,188,212]" pageId="3" pageNumber="48">
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||
(
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<figureCitation id="87A42A45FF86663AFF17349FA547FF76" box="[159,266,188,212]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1962,1985]" captionTargetBox="[174,1413,772,1939]" captionTargetId="figure@2.[174,1413,772,1941]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Holotypes of Drusilla. Dorsal habitus: 1) D. nearctica; 2) D. ashei. 3) D. nearctica lateral view of head and thorax." httpUri="https://zenodo.org/record/210859/files/figure.png" pageId="3" pageNumber="48">Figs. 1, 3</figureCitation>
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,
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<figureCitation id="87A42A45FF86663AFE9E349FA51CFF76" box="[278,337,188,212]" captionStart-0="FIGURES 4 – 5" captionStart-1="FIGURES 6 – 10" captionStartId-0="3.[151,264,1934,1957]" captionStartId-1="4.[151,264,1664,1687]" captionTargetBox-0="[197,1388,858,1905]" captionTargetBox-1="[194,1389,190,1623]" captionTargetId-0="figure@3.[185,1402,844,1913]" captionTargetId-1="figure@4.[184,1403,177,1641]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURES 4 – 5. Mouthparts of Drusilla nearctica. 4) prementum, ventral; 5) right maxilla, ventral. Scale bars (0.1 mm): I = 5; II = 4." captionText-1="FIGURES 6 – 10. Mouthparts of Drusilla nearctica. 6) labrum; 7) epipharynx; 8) left mandible, ventral; 9) right mandible, dorsal; 10) mentum, ventral; 11) left antenna, inner surface with dorsal surface facing down the page. Scale bars (0.1 mm): I = 11; II = 8 – 9; III = 6 – 7, 10." httpUri-0="https://zenodo.org/record/210860/files/figure.png" httpUri-1="https://zenodo.org/record/210861/files/figure.png" pageId="3" pageNumber="48">4–10</figureCitation>
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,
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<figureCitation id="87A42A45FF86663AFED5349FA5EEFF76" box="[349,419,188,212]" captionStart-0="FIGURES 12 – 19" captionStart-1="FIGURES 20 – 25" captionStartId-0="6.[151,264,1916,1939]" captionStartId-1="7.[151,264,1468,1491]" captionTargetBox-0="[181,1397,421,1887]" captionTargetBox-1="[164,1418,202,1445]" captionTargetId-0="figure@6.[168,1418,411,1895]" captionTargetId-1="figure@7.[158,1428,193,1457]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." captionText-1="FIGURES 20 – 25. Details of Drusilla nearctica (20 – 23) and D. ashei (24 – 25). 20) tergite VIII; 21) sternite VIII; 22) female genital segments (tergites IX and X), showing genital plate and spermathecal duct, dorsal; 23) spermatheca; 24) male genital segments (sternite IX and tergites IX and X), dorsal; 25) detail of sternite IX, ventral. Scale bars (0.1 mm): I = 20 – 22, 24 – 25." httpUri-0="https://zenodo.org/record/210862/files/figure.png" httpUri-1="https://zenodo.org/record/210863/files/figure.png" pageId="3" pageNumber="48">16–23</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="5785654BFF86663AFF1F3527A51EFE0E" pageId="3" pageNumber="48" type="materials_examined">
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<paragraph id="1F2036C0FF86663AFF1F3527A51EFE0E" blockId="3.[151,1436,259,789]" pageId="3" pageNumber="48">
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<emphasis id="2DEBEAD2FF86663AFF1F3527A66CFEBE" bold="true" box="[151,545,259,284]" pageId="3" pageNumber="48">
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<typeStatus id="C0248862FF86663AFF1F3527A49FFEBE" box="[151,210,260,284]" pageId="3" pageNumber="48">Type</typeStatus>
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material.
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<typeStatus id="C0248862FF86663AFEC53527A5F3FEBE" box="[333,446,260,284]" pageId="3" pageNumber="48" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="67887650FF86663AFE433520A656FEBE" box="[459,539,259,284]" name="United States of America" pageId="3" pageNumber="48">U.S.A.</collectingCountry>
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:
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</emphasis>
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3, “ARKANSAS: Pulaski Co./Pinnacle Ridge Estates/
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<quantity id="D8679B25FF86663AFB1C3527A080FEBE" box="[1172,1229,260,284]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="48" unit="km" value="2.0">2 km</quantity>
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W Pinnacle Mtn./ St. Pk.,
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<date id="6B211000FF86663AFF7B3504A5C6FE9D" box="[243,395,295,319]" pageId="3" pageNumber="48" value="1989-02-13">13 Feb. 1989</date>
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/R. Anderson, ex:berl./hardwood litter” (
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<collectionCode id="798EAE05FF86663AFCD33504A7E0FE9D" box="[859,941,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="3" pageNumber="48">SEMC</collectionCode>
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).
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<emphasis id="2DEBEAD2FF86663AFC4A350BA077FEE2" bold="true" box="[962,1082,296,320]" pageId="3" pageNumber="48">
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<typeStatus id="C0248862FF86663AFC4A350BA077FEE2" box="[962,1082,296,320]" pageId="3" pageNumber="48" type="paratype">Paratypes</typeStatus>
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</emphasis>
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||
(12
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<collectionCode id="798EAE05FF86663AFBE0350BA0DFFE9D" box="[1128,1170,296,319]" pageId="3" pageNumber="48">ƤƤ</collectionCode>
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, 153):
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||
<emphasis id="2DEBEAD2FF86663AFA8B3504A114FEE2" bold="true" box="[1283,1369,295,320]" pageId="3" pageNumber="48">
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||
<collectingCountry id="67887650FF86663AFA8B3504A11EFEE2" box="[1283,1363,295,320]" name="United States of America" pageId="3" pageNumber="48">U.S.A.</collectingCountry>
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:
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||
</emphasis>
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||
same data as
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<typeStatus id="C0248862FF86663AFF62356FA501FEC6" box="[234,332,332,356]" pageId="3" pageNumber="48" type="holotype">holotype</typeStatus>
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||
(12
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<collectionCode id="798EAE05FF86663AFEF0356FA5EFFEC1" box="[376,418,332,355]" pageId="3" pageNumber="48">ƤƤ</collectionCode>
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||
, 143;
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<collectionCode id="798EAE05FF86663AFD8D356FA61BFEC6" box="[517,598,332,356]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="3" pageNumber="48">SEMC</collectionCode>
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||
); “
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||
<collectingCountry id="67887650FF86663AFDFB356FA6E0FEC6" box="[627,685,332,356]" name="United States of America" pageId="3" pageNumber="48">USA</collectingCountry>
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||
AR/Logan Co./Mt. Magazine//
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<date id="6B211000FF86663AFB86356FA0DEFEC6" box="[1038,1171,332,356]" pageId="3" pageNumber="48" value="1986-06-02">2 June 1986</date>
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||
/
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<taxonomicName id="D89F4D43FF86663AFB14356EA14AFEC6" box="[1180,1287,333,356]" class="Actinopterygii" family="Lactariidae" genus="Lactarius" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="48" phylum="Chordata" rank="genus">
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||
<emphasis id="2DEBEAD2FF86663AFB14356EA14AFEC6" box="[1180,1287,333,356]" italics="true" pageId="3" pageNumber="48">Lactarius</emphasis>
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||
</taxonomicName>
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||
/R. Leschen// RL213” (3, specimen looks as though extruded genitalia were removed or broken off and were therefore never located;
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<collectionCode id="798EAE05FF86663AFF7E35B7A50AFE0E" box="[246,327,404,428]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="3" pageNumber="48">SEMC</collectionCode>
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).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5785654BFF86663AFF4F3594A003FD9E" pageId="3" pageNumber="48" type="diagnosis">
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<paragraph id="1F2036C0FF86663AFF4F3594A003FD9E" blockId="3.[151,1436,259,789]" pageId="3" pageNumber="48">
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<emphasis id="2DEBEAD2FF86663AFF4F3594A50CFE72" bold="true" box="[199,321,439,464]" pageId="3" pageNumber="48">Diagnosis.</emphasis>
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<taxonomicName id="D89F4D43FF86663AFECF359BA65EFE72" box="[327,531,440,464]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="nearctica">
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<emphasis id="2DEBEAD2FF86663AFECF359BA65EFE72" box="[327,531,440,464]" italics="true" pageId="3" pageNumber="48">Drusilla nearctica</emphasis>
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||
</taxonomicName>
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most closely resembles
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<taxonomicName id="D89F4D43FF86663AFCAC359AA7CCFE72" box="[804,897,440,464]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="ashei">
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<emphasis id="2DEBEAD2FF86663AFCAC359AA7CCFE72" box="[804,897,440,464]" italics="true" pageId="3" pageNumber="48">D. ashei</emphasis>
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</taxonomicName>
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, but can be confidently separated by the following male characters: shape of the median lobe (
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<figureCitation id="87A42A45FF86663AFD2635FFA778FE56" box="[686,821,476,500]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="3" pageNumber="48">Figs. 16–17</figureCitation>
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); distal apodeme short, less than four times as long as its width (
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<figureCitation id="87A42A45FF86663AFE8E35DCA514FDB5" box="[262,345,511,535]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="3" pageNumber="48">Fig. 16</figureCitation>
|
||
); aedeagal costal morphology of parameral surface (
|
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<figureCitation id="87A42A45FF86663AFC2935DCA7BEFDB5" box="[929,1011,511,535]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="3" pageNumber="48">Fig. 17</figureCitation>
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||
); paramere with velum obscuring apical lobe in outer lateral view (
|
||
<figureCitation id="87A42A45FF86663AFE6E3607A674FD9E" box="[486,569,548,572]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="3" pageNumber="48">Fig. 18</figureCitation>
|
||
); chaetotaxy of parameral apical lobe (
|
||
<figureCitation id="87A42A45FF86663AFC653607A00DFD9E" box="[1005,1088,548,572]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="3" pageNumber="48">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5785654BFF86663CFF4F3664A51CFC06" lastPageId="5" lastPageNumber="50" pageId="3" pageNumber="48" type="description">
|
||
<paragraph id="1F2036C0FF86663AFF4F3664A0FAFCB6" blockId="3.[151,1436,259,789]" pageId="3" pageNumber="48">
|
||
<emphasis id="2DEBEAD2FF86663AFF4F3664A514FDC2" bold="true" box="[199,345,583,608]" pageId="3" pageNumber="48">Description.</emphasis>
|
||
Body (
|
||
<figureCitation id="87A42A45FF86663AFE393664A5B8FDFD" box="[433,501,583,607]" captionStart="FIGURES 1 – 3" captionStartId="2.[151,264,1962,1985]" captionTargetBox="[174,1413,772,1939]" captionTargetId="figure@2.[174,1413,772,1941]" captionTargetPageId="2" captionText="FIGURES 1 – 3. Holotypes of Drusilla. Dorsal habitus: 1) D. nearctica; 2) D. ashei. 3) D. nearctica lateral view of head and thorax." httpUri="https://zenodo.org/record/210859/files/figure.png" pageId="3" pageNumber="48">Fig. 1</figureCitation>
|
||
) length on average
|
||
<quantity id="D8679B25FF86663AFD6A3664A776FDFD" box="[738,827,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="3" pageNumber="48" unit="mm" value="3.3">3.3 mm</quantity>
|
||
(
|
||
<quantity id="D8679B25FF86663AFCC53664A791FDFD" box="[845,988,583,607]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.6" metricValueMin="3.1" pageId="3" pageNumber="48" unit="mm" value="3.35" valueMax="3.6" valueMin="3.1">3.1–3.6 mm</quantity>
|
||
, n = 10).
|
||
<collectingCountry id="67887650FF86663AFBDA3664A0C7FDFD" box="[1106,1162,583,607]" name="India" pageId="3" pageNumber="48">Hind</collectingCountry>
|
||
wings degenerated and absent. Body color with head, pronotum, elytra and most of abdomen dark brown; usually most (if partial, mediobasal portion) of abdominal tergite VI black; frequently tergites V, VII and VIII brown to approaching black and usually not as dark as tergite VI; segments III–IV sometimes with a small basal portion of tergal disk approaching brown compared with rest of given segment; abdominal sternites usually corresponding with tergites in overall color patterning amongst segments; antennae, legs and mouthparts yellowish-brown to brown.
|
||
</paragraph>
|
||
<caption id="4BE06648FF86663AFF1F33ADA49BF860" httpUri="https://zenodo.org/record/210860/files/figure.png" pageId="3" pageNumber="48" targetBox="[197,1388,858,1905]" targetPageId="3">
|
||
<paragraph id="1F2036C0FF86663AFF1F33ADA49BF860" blockId="3.[151,1436,1934,1987]" pageId="3" pageNumber="48">
|
||
<emphasis id="2DEBEAD2FF86663AFF1F33ADA57EF801" bold="true" box="[151,307,1934,1957]" pageId="3" pageNumber="48">FIGURES 4–5</emphasis>
|
||
. Mouthparts of
|
||
<taxonomicName id="D89F4D43FF86663AFE5F33ADA6DDF806" box="[471,656,1934,1956]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="48" phylum="Arthropoda" rank="species" species="nearctica">
|
||
<emphasis id="2DEBEAD2FF86663AFE5F33ADA6DDF806" box="[471,656,1934,1956]" italics="true" pageId="3" pageNumber="48">Drusilla nearctica</emphasis>
|
||
</taxonomicName>
|
||
. 4) prementum, ventral; 5) right maxilla, ventral. Scale bars (0.1 mm): I = 5; II = 4.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="4BE06648FF81663DFF1F32A3A530F971" httpUri="https://zenodo.org/record/210861/files/figure.png" pageId="4" pageNumber="49" targetBox="[194,1389,190,1623]" targetPageId="4">
|
||
<paragraph id="1F2036C0FF81663DFF1F32A3A530F971" blockId="4.[151,1436,1664,1748]" pageId="4" pageNumber="49">
|
||
<emphasis id="2DEBEAD2FF81663DFF1F32A3A50DF934" bold="true" box="[151,320,1664,1687]" pageId="4" pageNumber="49">FIGURES 6–10</emphasis>
|
||
. Mouthparts of
|
||
<taxonomicName id="D89F4D43FF81663DFE6B32A3A6D0F935" box="[483,669,1664,1687]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="49" phylum="Arthropoda" rank="species" species="nearctica">
|
||
<emphasis id="2DEBEAD2FF81663DFE6B32A3A6D0F935" box="[483,669,1664,1687]" italics="true" pageId="4" pageNumber="49">Drusilla nearctica</emphasis>
|
||
</taxonomicName>
|
||
. 6) labrum; 7) epipharynx; 8) left mandible, ventral; 9) right mandible, dorsal; 10) mentum, ventral; 11) left antenna, inner surface with dorsal surface facing down the page. Scale bars (0.1 mm): I = 11; II = 8–9; III = 6–7, 10.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1F2036C0FF81663DFF4F3323A6B3F852" blockId="4.[151,1436,1792,2033]" pageId="4" pageNumber="49">
|
||
Head subcircular (HW =
|
||
<quantity id="D8679B25FF81663DFE6A3323A60AF8BA" box="[482,583,1792,1816]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="4" pageNumber="49" unit="mm" value="0.47">0.47 mm</quantity>
|
||
,
|
||
<quantity id="D8679B25FF81663DFDDB3323A6B4F8BA" box="[595,761,1792,1816]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="4.8" metricValueMin="4.699999999999999" pageId="4" pageNumber="49" unit="mm" value="0.475" valueMax="0.48" valueMin="0.47">0.47–0.48 mm</quantity>
|
||
; HL =
|
||
<quantity id="D8679B25FF81663DFCCF3323A7E3F8BA" box="[839,942,1792,1816]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="49" unit="mm" value="0.48">0.48 mm</quantity>
|
||
,
|
||
<quantity id="D8679B25FF81663DFC313323A012F8BA" box="[953,1119,1792,1816]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="5.0" metricValueMin="4.699999999999999" pageId="4" pageNumber="49" unit="mm" value="0.485" valueMax="0.5" valueMin="0.47">0.47–0.50 mm</quantity>
|
||
; HW/HL = 0.98, 0.94–1.00) with faint imbricate microsculpture; eyes small and occupying approximately a third of head length (OL = 0.16,
|
||
<quantity id="D8679B25FF81663DFF1F336BA572F8C2" box="[151,319,1864,1888]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="4" pageNumber="49" unit="mm" value="0.16" valueMax="0.17" valueMin="0.15">0.15–0.17 mm</quantity>
|
||
; OL/HL = 0.34, 0.30–0.35); setae on vertex directed posteriomedially; labrum (
|
||
<figureCitation id="87A42A45FF81663DFB46336BA15EF8C2" box="[1230,1299,1864,1888]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1664,1687]" captionTargetBox="[194,1389,190,1623]" captionTargetId="figure@4.[184,1403,177,1641]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Mouthparts of Drusilla nearctica. 6) labrum; 7) epipharynx; 8) left mandible, ventral; 9) right mandible, dorsal; 10) mentum, ventral; 11) left antenna, inner surface with dorsal surface facing down the page. Scale bars (0.1 mm): I = 11; II = 8 – 9; III = 6 – 7, 10." httpUri="https://zenodo.org/record/210861/files/figure.png" pageId="4" pageNumber="49">Fig. 6</figureCitation>
|
||
) with distal seta 2 (ds2) posterior of medial setae; epipharynx (
|
||
<figureCitation id="87A42A45FF81663DFD5C334FA755F826" box="[724,792,1900,1924]" captionStart="FIGURES 6 – 10" captionStartId="4.[151,264,1664,1687]" captionTargetBox="[194,1389,190,1623]" captionTargetId="figure@4.[184,1403,177,1641]" captionTargetPageId="4" captionText="FIGURES 6 – 10. Mouthparts of Drusilla nearctica. 6) labrum; 7) epipharynx; 8) left mandible, ventral; 9) right mandible, dorsal; 10) mentum, ventral; 11) left antenna, inner surface with dorsal surface facing down the page. Scale bars (0.1 mm): I = 11; II = 8 – 9; III = 6 – 7, 10." httpUri="https://zenodo.org/record/210861/files/figure.png" pageId="4" pageNumber="49">Fig. 7</figureCitation>
|
||
) with a generalized athetine form; maxillary (
|
||
<figureCitation id="87A42A45FF81663DFA96334FA12CF826" box="[1310,1377,1900,1924]" captionStart="FIGURES 4 – 5" captionStartId="3.[151,264,1934,1957]" captionTargetBox="[197,1388,858,1905]" captionTargetId="figure@3.[185,1402,844,1913]" captionTargetPageId="3" captionText="FIGURES 4 – 5. Mouthparts of Drusilla nearctica. 4) prementum, ventral; 5) right maxilla, ventral. Scale bars (0.1 mm): I = 5; II = 4." httpUri="https://zenodo.org/record/210860/files/figure.png" pageId="4" pageNumber="49">Fig. 5</figureCitation>
|
||
) lacinia with a distal comb consisting of approximately five spines, palpomere IV subulate with filamentous sensillae that are fitted against segment, palpomere IV distally lacking an apical process; labial palpus (
|
||
<figureCitation id="87A42A45FF81663DFB383397A0BEF86E" box="[1200,1267,1972,1996]" captionStart="FIGURES 4 – 5" captionStartId="3.[151,264,1934,1957]" captionTargetBox="[197,1388,858,1905]" captionTargetId="figure@3.[185,1402,844,1913]" captionTargetPageId="3" captionText="FIGURES 4 – 5. Mouthparts of Drusilla nearctica. 4) prementum, ventral; 5) right maxilla, ventral. Scale bars (0.1 mm): I = 5; II = 4." httpUri="https://zenodo.org/record/210860/files/figure.png" pageId="4" pageNumber="49">Fig. 4</figureCitation>
|
||
) lacking setula β and δ, see generic diagnosis for a further description.
|
||
</paragraph>
|
||
<paragraph id="1F2036C0FF80663CFF4F34BBA6E9FF55" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="50">
|
||
Pronotum slightly transverse (PW =
|
||
<quantity id="D8679B25FF80663CFDED34B4A680FF0D" box="[613,717,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="5" pageNumber="50" unit="mm" value="0.54">0.54 mm</quantity>
|
||
,
|
||
<quantity id="D8679B25FF80663CFD5134B4A732FF0D" box="[729,895,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="5.35" metricValueMax="5.6" metricValueMin="5.1" pageId="5" pageNumber="50" unit="mm" value="0.535" valueMax="0.56" valueMin="0.51">0.51–0.56 mm</quantity>
|
||
; PL =
|
||
<quantity id="D8679B25FF80663CFC4434B4A079FF0D" box="[972,1076,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="50" unit="mm" value="0.48">0.48 mm</quantity>
|
||
,
|
||
<quantity id="D8679B25FF80663CFBB734B4A0ABFF0D" box="[1087,1254,151,175]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" metricValueMax="5.1" metricValueMin="4.5" pageId="5" pageNumber="50" unit="mm" value="0.48" valueMax="0.51" valueMin="0.45">0.45–0.51 mm</quantity>
|
||
; PW/PL = 1.12, 1.09–1.17) with faint imbricate microsculpture; three pairs of macrosetae present, consisting of one apical pair and two lateral pairs anterior of horizontal midline.
|
||
</paragraph>
|
||
<paragraph id="1F2036C0FF80663CFF4F3527A75DFE0E" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="50">
|
||
Elytra, together, transverse (EW = 0.59,
|
||
<quantity id="D8679B25FF80663CFD023527A77DFEBE" box="[650,816,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="6.3" metricValueMin="5.699999999999999" pageId="5" pageNumber="50" unit="mm" value="0.6" valueMax="0.63" valueMin="0.57">0.57–0.63 mm</quantity>
|
||
; EL = 0.44,
|
||
<quantity id="D8679B25FF80663CFC313527A012FEBE" box="[953,1119,260,284]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="5" pageNumber="50" unit="mm" value="0.435" valueMax="0.45" valueMin="0.42">0.42–0.45 mm</quantity>
|
||
; EW/EL = 1.36, 1.27–1.41); each elytron broadly rounded with posterior-most extension medially; elytron with pleural surface forming an approximate right angle with disk; disk with longitudinal basolateral impression; disk of elytron with conspicuously stronger and more irregularly imbricate microsculpture compared with head and pronotum; macroseta on each elytron at subapicolateral margin on dorsal surface.
|
||
</paragraph>
|
||
<paragraph id="1F2036C0FF80663CFF4F3594A0DDFE56" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="50">
|
||
Abdomen with dorsal surface subglabrous; tergite VIII (
|
||
<figureCitation id="87A42A45FF80663CFCDE3594A7E6FE6D" box="[854,939,439,463]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,264,1468,1491]" captionTargetBox="[164,1418,202,1445]" captionTargetId="figure@7.[158,1428,193,1457]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Details of Drusilla nearctica (20 – 23) and D. ashei (24 – 25). 20) tergite VIII; 21) sternite VIII; 22) female genital segments (tergites IX and X), showing genital plate and spermathecal duct, dorsal; 23) spermatheca; 24) male genital segments (sternite IX and tergites IX and X), dorsal; 25) detail of sternite IX, ventral. Scale bars (0.1 mm): I = 20 – 22, 24 – 25." httpUri="https://zenodo.org/record/210863/files/figure.png" pageId="5" pageNumber="50">Fig. 20</figureCitation>
|
||
) with apical margin sinuate, resulting in a broad medial and acute lateral projections; sternite VIII (
|
||
<figureCitation id="87A42A45FF80663CFC9B35FFA728FE56" box="[787,869,476,500]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,264,1468,1491]" captionTargetBox="[164,1418,202,1445]" captionTargetId="figure@7.[158,1428,193,1457]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Details of Drusilla nearctica (20 – 23) and D. ashei (24 – 25). 20) tergite VIII; 21) sternite VIII; 22) female genital segments (tergites IX and X), showing genital plate and spermathecal duct, dorsal; 23) spermatheca; 24) male genital segments (sternite IX and tergites IX and X), dorsal; 25) detail of sternite IX, ventral. Scale bars (0.1 mm): I = 20 – 22, 24 – 25." httpUri="https://zenodo.org/record/210863/files/figure.png" pageId="5" pageNumber="50">Fig. 21</figureCitation>
|
||
) with a blunt medial apex.
|
||
</paragraph>
|
||
<paragraph id="1F2036C0FF80663CFF4F35DCA152FCB6" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="50">
|
||
Males with vertex of head variably impressed. Pronotum variably, medially impressed. Median lobe ovular in lateral (
|
||
<figureCitation id="87A42A45FF80663CFF643607A570FD9E" box="[236,317,548,572]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 16</figureCitation>
|
||
) and parameral (
|
||
<figureCitation id="87A42A45FF80663CFE733607A601FD9E" box="[507,588,548,572]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 17</figureCitation>
|
||
) view; apical process of median lobe short, extending paramerally; athetine bridge incomplete at aparameral surface, with length of distal apodeme less then four times its width; foramen situated approximately in middle; proximal costa extending most of surface posterior to foramen; median costa extending to apical lobe; arcuate costa sinuate, diverging and closely approximating either side of median costa, again broadly diverging towards apicolateral base of apical lobe. Paramere (
|
||
<figureCitation id="87A42A45FF80663CFC323697A043FD6E" box="[954,1038,692,716]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 18</figureCitation>
|
||
), in outer lateral view, with length approximately two times width; velum large and obscuring apical lobe in outer lateral view; apical lobe of paramere (
|
||
<figureCitation id="87A42A45FF80663CFE8636DFA52DFCB6" box="[270,352,764,788]" captionStart="FIGURES 12 – 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 – 19. Male genitalia of Drusilla ashei (12 – 15) and D. nearctica (16 – 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 – 14, 16 – 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 19</figureCitation>
|
||
) in inner lateral view with four setulae, subequal in length, crowded at apical margin.
|
||
</paragraph>
|
||
<paragraph id="1F2036C0FF80663CFF4F373CA51CFC06" blockId="5.[151,1436,151,968]" pageId="5" pageNumber="50">
|
||
Females with vertex of head with shallow impression that is less pronounced than in males; pronotum usual not medially impressed, but when impressed, to a much lesser degree than in males; spermatheca (
|
||
<figureCitation id="87A42A45FF80663CFB7B3767A10BFCFE" box="[1267,1350,836,860]" captionStart="FIGURES 20 – 25" captionStartId="7.[151,264,1468,1491]" captionTargetBox="[164,1418,202,1445]" captionTargetId="figure@7.[158,1428,193,1457]" captionTargetPageId="7" captionText="FIGURES 20 – 25. Details of Drusilla nearctica (20 – 23) and D. ashei (24 – 25). 20) tergite VIII; 21) sternite VIII; 22) female genital segments (tergites IX and X), showing genital plate and spermathecal duct, dorsal; 23) spermatheca; 24) male genital segments (sternite IX and tergites IX and X), dorsal; 25) detail of sternite IX, ventral. Scale bars (0.1 mm): I = 20 – 22, 24 – 25." httpUri="https://zenodo.org/record/210863/files/figure.png" pageId="5" pageNumber="50">Fig. 23</figureCitation>
|
||
) with a simple bend away from side of spermathecal gland, proximal end bulbous in form, distal end slightly dilated with an internal cone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5785654BFF80663CFF4F3793A0ECFC65" box="[199,1185,943,968]" pageId="5" pageNumber="50" type="etymology">
|
||
<paragraph id="1F2036C0FF80663CFF4F3793A0ECFC65" blockId="5.[151,1436,151,968]" box="[199,1185,943,968]" pageId="5" pageNumber="50">
|
||
<emphasis id="2DEBEAD2FF80663CFF4F3793A503FC6A" bold="true" box="[199,334,944,968]" pageId="5" pageNumber="50">Etymology.</emphasis>
|
||
In commemoration, as the first endemic species of
|
||
<taxonomicName id="D89F4D43FF80663CFC183793A7A0FC6A" box="[912,1005,944,968]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="2DEBEAD2FF80663CFC183793A7A0FC6A" box="[912,1005,944,968]" italics="true" pageId="5" pageNumber="50">Drusilla</emphasis>
|
||
</taxonomicName>
|
||
to the Nearctic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |