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<document id="EE697101602BB314D6FBC3B3E11A1B9F" ID-DOI="10.5281/zenodo.210858" ID-GBIF-Dataset="6c13241a-b0ff-420a-acf0-8b9cbd3b9703" ID-ISSN="1175-5326" ID-Zenodo-Dep="210858" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460478186346" checkinUser="plazi" docAuthor="Eldredge, Taro" docDate="2012" docId="973687D6FF80663FFF1F3032A13BFEC6" docLanguage="en" docName="zt03300p054.pdf" docOrigin="Zootaxa 3300" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Drusilla ashei Eldredge, new species" docType="treatment" docVersion="9" lastPageNumber="51" masterDocId="6B0FFFAEFF856639FF883423A44DFFA2" masterDocTitle="Review of the Nearctic species of Drusilla Leach (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="54" masterPageNumber="45" pageNumber="50" updateTime="1698307154067" updateUser="plazi">
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<mods:title id="6149AFD99CE986FA2C50F820A1CEB6A1">Review of the Nearctic species of Drusilla Leach (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="15BBBB9AAD70A4847161F9FFC21C13FE">Eldredge, Taro</mods:namePart>
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<mods:typeOfResource id="86BDA188E51B57EABD1BF9C395A570C3">text</mods:typeOfResource>
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<mods:date id="F5F656DE58DEAB4ADFDF12F690EFC078">2012</mods:date>
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<mods:identifier id="6AFDD79CDBC812E41F72DF0C7BB03118" type="DOI">10.5281/zenodo.210858</mods:identifier>
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<treatment id="973687D6FF80663FFF1F3032A13BFEC6" ID-DOI="http://doi.org/10.5281/zenodo.6176690" ID-GBIF-Taxon="119516771" ID-Zenodo-Dep="6176690" LSID="urn:lsid:plazi:treatment:973687D6FF80663FFF1F3032A13BFEC6" httpUri="http://treatment.plazi.org/id/973687D6FF80663FFF1F3032A13BFEC6" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
<subSubSection id="5785654BFF80663CFF1F3032A62DFB88" box="[151,608,1041,1066]" pageId="5" pageNumber="50" type="nomenclature">
<paragraph id="1F2036C0FF80663CFF1F3032A62DFB88" blockId="5.[151,608,1041,1101]" box="[151,608,1041,1066]" pageId="5" pageNumber="50">
<heading id="446881ACFF80663CFF1F3032A62DFB88" bold="true" box="[151,608,1041,1066]" fontSize="10" level="2" pageId="5" pageNumber="50" reason="2">
<emphasis id="2DEBEAD2FF80663CFF1F3032A62DFB88" bold="true" box="[151,608,1041,1066]" pageId="5" pageNumber="50">
3.
<taxonomicName id="D89F4D43FF80663CFF3B3032A587FB88" ID-CoL="37TDN" authority="Eldredge" authorityName="Eldredge" box="[179,458,1041,1066]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ashei" status="sp. nov.">
<emphasis id="2DEBEAD2FF80663CFF3B3032A51BFB88" bold="true" box="[179,342,1041,1066]" italics="true" pageId="5" pageNumber="50">Drusilla ashei</emphasis>
Eldredge
</taxonomicName>
,
<taxonomicNameLabel id="36D857A9FF80663CFE5F3031A62DFB88" box="[471,608,1041,1066]" pageId="5" pageNumber="50" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF80663CFF1F3016A5D3FBEF" box="[151,414,1077,1101]" pageId="5" pageNumber="50" type="description">
<paragraph id="1F2036C0FF80663CFF1F3016A5D3FBEF" blockId="5.[151,608,1041,1101]" box="[151,414,1077,1101]" pageId="5" pageNumber="50">
(
<figureCitation id="87A42A45FF80663CFF173016A4A2FBEF" box="[159,239,1077,1101]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1962,1985]" captionTargetBox="[174,1413,772,1939]" captionTargetId="figure@2.[174,1413,772,1941]" captionTargetPageId="2" captionText="FIGURES 1 3. Holotypes of Drusilla. Dorsal habitus: 1) D. nearctica; 2) D. ashei. 3) D. nearctica lateral view of head and thorax." httpUri="https://zenodo.org/record/210859/files/figure.png" pageId="5" pageNumber="50">Figs. 2</figureCitation>
,
<figureCitation id="87A42A45FF80663CFF723016A50FFBEF" box="[250,322,1077,1101]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">1215</figureCitation>
,
<figureCitation id="87A42A45FF80663CFEC63016A5DBFBEF" box="[334,406,1077,1101]" captionStart="FIGURES 20 25" captionStartId="7.[151,264,1468,1491]" captionTargetBox="[164,1418,202,1445]" captionTargetId="figure@7.[158,1428,193,1457]" captionTargetPageId="7" captionText="FIGURES 20 25. Details of Drusilla nearctica (20 23) and D. ashei (24 25). 20) tergite VIII; 21) sternite VIII; 22) female genital segments (tergites IX and X), showing genital plate and spermathecal duct, dorsal; 23) spermatheca; 24) male genital segments (sternite IX and tergites IX and X), dorsal; 25) detail of sternite IX, ventral. Scale bars (0.1 mm): I = 20 22, 24 25." httpUri="https://zenodo.org/record/210863/files/figure.png" pageId="5" pageNumber="50">2425</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF80663CFF1F305EA79CFAEB" pageId="5" pageNumber="50" type="materials_examined">
<paragraph id="1F2036C0FF80663CFF1F305EA79CFAEB" blockId="5.[151,1437,1149,2037]" pageId="5" pageNumber="50">
<emphasis id="2DEBEAD2FF80663CFF1F305EA675FB34" bold="true" box="[151,568,1149,1174]" pageId="5" pageNumber="50">
<typeStatus id="C0248862FF80663CFF1F305EA49FFB37" box="[151,210,1149,1173]" pageId="5" pageNumber="50">Type</typeStatus>
material.
<typeStatus id="C0248862FF80663CFED4305EA580FB37" box="[348,461,1149,1173]" pageId="5" pageNumber="50" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="67887650FF80663CFE69305EA67FFB34" box="[481,562,1149,1174]" name="United States of America" pageId="5" pageNumber="50">U.S.A.</collectingCountry>
:
</emphasis>
3, “
<collectionCode id="798EAE05FF80663CFDF4305EA693FB37" box="[636,734,1149,1173]" pageId="5" pageNumber="50">NORTH</collectionCode>
CAROLINA/Rockingham Co./Reidsville,
<date id="6B211000FF80663CFB53305EA1D5FB37" box="[1243,1432,1149,1173]" pageId="5" pageNumber="50" value="1979-05-05" valueMax="1979-05-13" valueMin="1979-05-05">513 May/1979</date>
, J.S.Ashe/ex., pan traps” (
<collectionCode id="798EAE05FF80663CFE393082A64EFB1B" box="[433,515,1185,1209]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="50">SEMC</collectionCode>
).
<emphasis id="2DEBEAD2FF80663CFD9E3081A6C3FB18" bold="true" box="[534,654,1186,1210]" pageId="5" pageNumber="50">
<typeStatus id="C0248862FF80663CFD9E3081A6C3FB18" box="[534,654,1186,1210]" pageId="5" pageNumber="50" type="paratype">Paratypes</typeStatus>
</emphasis>
(3
<collectionCode id="798EAE05FF80663CFD243081A69BFB1B" box="[684,726,1186,1209]" pageId="5" pageNumber="50">ƤƤ</collectionCode>
, 43):
<emphasis id="2DEBEAD2FF80663CFCBC3082A7C6FB18" bold="true" box="[820,907,1185,1210]" pageId="5" pageNumber="50">
<collectingCountry id="67887650FF80663CFCBC3082A7C8FB18" box="[820,901,1185,1210]" name="United States of America" pageId="5" pageNumber="50">U.S.A.</collectingCountry>
:
</emphasis>
same as
<typeStatus id="C0248862FF80663CFC783082A01FFB1B" box="[1008,1106,1185,1209]" pageId="5" pageNumber="50" type="holotype">holotype</typeStatus>
label (23;
<collectionCode id="798EAE05FF80663CFB6F3082A175FB1B" box="[1255,1336,1185,1209]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="50">SEMC</collectionCode>
); “Rockingham Co., Reids-/ville,
<date id="6B211000FF80663CFE3B30E6A6E8FB7F" box="[435,677,1221,1245]" pageId="5" pageNumber="50" value="1970-04-29" valueMax="1970-05-03" valueMin="1970-04-29">Apr. 29-May 3, 1970</date>
/J. Ashe, ex: pan trap” (3;
<collectionCode id="798EAE05FF80663CFC5E30E6A06AFB7F" box="[982,1063,1221,1245]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="50">SEMC</collectionCode>
); “Rockingham Co., N.C./Reidsville/
<date id="6B211000FF80663CFF5B30CAA524FAA3" box="[211,361,1257,1281]" pageId="5" pageNumber="50" value="1973-08-12">VIII-12-1973</date>
/JS, Ashe//James S. Ashe Collection” (Ƥ;
<collectionCode id="798EAE05FF80663CFCCF30CAA7D5FAA3" box="[839,920,1257,1281]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="50">SEMC</collectionCode>
); “Rockingham Co., N.C./Reidsville/
<date id="6B211000FF80663CFAB730CAA49EFA87" pageId="5" pageNumber="50" value="1973-08-14">VIII-14- 1973</date>
/J.S. Ashe//on and under
<taxonomicName id="D89F4D43FF80663CFE79312EA608FA87" box="[497,581,1293,1317]" class="Agaricomycetes" family="Boletaceae" genus="Boletus" kingdom="Fungi" order="Boletales" pageId="5" pageNumber="50" phylum="Basidiomycota" rank="genus">
<emphasis id="2DEBEAD2FF80663CFE79312EA608FA87" box="[497,581,1293,1317]" italics="true" pageId="5" pageNumber="50">Boletus</emphasis>
</taxonomicName>
sp.//James S. Ashe Collection” (2
<collectionCode id="798EAE05FF80663CFC58312DA7B7FA87" box="[976,1018,1294,1317]" pageId="5" pageNumber="50">ƤƤ</collectionCode>
, 3 [additional label reads: “will not key out/in
<bibRefCitation id="7B0E4B31FF80663CFE863112A5F9FAEB" author="Seevers" box="[270,436,1329,1353]" pageId="5" pageNumber="50" refString="Seevers, C. H. (1978) A generic and tribal revision of the North American Aleocharinae (Coleoptera: Staphylinidae). Fieldiana: Zoology, 71, vi + 275 pp." type="book" year="1978">Seevers (1978)</bibRefCitation>
/but seems closest/to
<taxonomicName id="D89F4D43FF80663CFD2E3112A71BFAEB" authority="Leach" authorityName="Leach" box="[678,854,1329,1353]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="2DEBEAD2FF80663CFD2E3112A74EFAEB" box="[678,771,1329,1353]" italics="true" pageId="5" pageNumber="50">Drusilla</emphasis>
Leach
</taxonomicName>
”];
<collectionCode id="798EAE05FF80663CFCFB3112A788FAEB" box="[883,965,1329,1353]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="5" pageNumber="50">SEMC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF80663CFF4F3176A7DFFA7B" pageId="5" pageNumber="50" type="diagnosis">
<paragraph id="1F2036C0FF80663CFF4F3176A7DFFA7B" blockId="5.[151,1437,1149,2037]" pageId="5" pageNumber="50">
<emphasis id="2DEBEAD2FF80663CFF4F3176A50CFACC" bold="true" box="[199,321,1365,1390]" pageId="5" pageNumber="50">Diagnosis.</emphasis>
<taxonomicName id="D89F4D43FF80663CFECF3176A5ABFACF" box="[327,486,1365,1389]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="2DEBEAD2FF80663CFECF3176A5ABFACF" box="[327,486,1365,1389]" italics="true" pageId="5" pageNumber="50">Drusilla ashei</emphasis>
</taxonomicName>
most closely resembles
<taxonomicName id="D89F4D43FF80663CFD7F3175A7CCFACF" box="[759,897,1366,1389]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="nearctica">
<emphasis id="2DEBEAD2FF80663CFD7F3175A7CCFACF" box="[759,897,1366,1389]" italics="true" pageId="5" pageNumber="50">D. nearctica</emphasis>
</taxonomicName>
, but can be confidently separated by the following male characters: shape of the median lobe (
<figureCitation id="87A42A45FF80663CFD23315AA778FA33" box="[683,821,1401,1425]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Figs. 1213</figureCitation>
); distal apodeme eight times its width (
<figureCitation id="87A42A45FF80663CFB79315AA108FA33" box="[1265,1349,1401,1425]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 12</figureCitation>
); costal morphology of median lobe parameral surface (
<figureCitation id="87A42A45FF80663CFD3F31BEA746FA17" box="[695,779,1437,1461]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 13</figureCitation>
); paramere with velum not obscuring apical lobe in outer lateral view (
<figureCitation id="87A42A45FF80663CFEA131E2A530FA7B" box="[297,381,1473,1497]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 14</figureCitation>
); chaetotaxy of parameral apical lobe (
<figureCitation id="87A42A45FF80663CFCB931E2A7C9FA7B" box="[817,900,1473,1497]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF80663FFF4F31C6A6D6FE9D" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50" type="description">
<paragraph id="1F2036C0FF80663CFF4F31C6A763F983" blockId="5.[151,1437,1149,2037]" pageId="5" pageNumber="50">
<emphasis id="2DEBEAD2FF80663CFF4F31C6A515FA5F" bold="true" box="[199,344,1509,1534]" pageId="5" pageNumber="50">Description.</emphasis>
<taxonomicName id="D89F4D43FF80663CFEEB31C6A64BFA5F" box="[355,518,1509,1533]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="2DEBEAD2FF80663CFEEB31C6A64BFA5F" box="[355,518,1509,1533]" italics="true" pageId="5" pageNumber="50">Drusilla ashei</emphasis>
</taxonomicName>
is externally indistinguishable from
<taxonomicName id="D89F4D43FF80663CFC3C31C5A00EFA5F" box="[948,1091,1510,1533]" class="Insecta" family="Staphylinidae" genus="Drusilla" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="50" phylum="Arthropoda" rank="species" species="nearctica">
<emphasis id="2DEBEAD2FF80663CFC3C31C5A00EFA5F" box="[948,1091,1510,1533]" italics="true" pageId="5" pageNumber="50">D. nearctica</emphasis>
</taxonomicName>
in many respects; only those characters that differ have been included in the description.
</paragraph>
<paragraph id="1F2036C0FF80663CFF4F320EA060F92F" blockId="5.[151,1437,1149,2037]" pageId="5" pageNumber="50">
Body (
<figureCitation id="87A42A45FF80663CFE9C320EA514F9E7" box="[276,345,1581,1605]" captionStart="FIGURES 1 3" captionStartId="2.[151,264,1962,1985]" captionTargetBox="[174,1413,772,1939]" captionTargetId="figure@2.[174,1413,772,1941]" captionTargetPageId="2" captionText="FIGURES 1 3. Holotypes of Drusilla. Dorsal habitus: 1) D. nearctica; 2) D. ashei. 3) D. nearctica lateral view of head and thorax." httpUri="https://zenodo.org/record/210859/files/figure.png" pageId="5" pageNumber="50">Fig. 2</figureCitation>
) length on average
<quantity id="D8679B25FF80663CFDB3320EA6DFF9E7" box="[571,658,1581,1605]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="5" pageNumber="50" unit="mm" value="3.2">3.2 mm</quantity>
(
<quantity id="D8679B25FF80663CFD29320EA707F9E7" box="[673,842,1581,1605]" metricMagnitude="-3" metricUnit="m" metricValue="3.2199999999999998" metricValueMax="3.39" metricValueMin="3.05" pageId="5" pageNumber="50" unit="mm" value="3.2199999999999998" valueMax="3.39" valueMin="3.05">3.053.39 mm</quantity>
, n = 8). Body color with head, pronotum, elytra and most of abdomen dark brown, sometimes with elytra and tergites IIIV brown; tergites VVIII piceous; legs usually concolorous with most of body and dark brown, sometimes yellowish-brown.
</paragraph>
<paragraph id="1F2036C0FF80663CFF4F32BAA494F95B" blockId="5.[151,1437,1149,2037]" pageId="5" pageNumber="50">
Head subcircular (HW =
<quantity id="D8679B25FF80663CFE6832BAA60BF913" box="[480,582,1689,1713]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" pageId="5" pageNumber="50" unit="mm" value="0.47">0.47 mm</quantity>
,
<quantity id="D8679B25FF80663CFDD932BAA6BBF913" box="[593,758,1689,1713]" metricMagnitude="-4" metricUnit="m" metricValue="4.6499999999999995" metricValueMax="4.8" metricValueMin="4.5" pageId="5" pageNumber="50" unit="mm" value="0.46499999999999997" valueMax="0.48" valueMin="0.45">0.450.48 mm</quantity>
; HL =
<quantity id="D8679B25FF80663CFCCB32BAA7E7F913" box="[835,938,1689,1713]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="5" pageNumber="50" unit="mm" value="0.48">0.48 mm</quantity>
,
<quantity id="D8679B25FF80663CFC3C32BAA014F913" box="[948,1113,1689,1713]" metricMagnitude="-4" metricUnit="m" metricValue="4.75" metricValueMax="5.0" metricValueMin="4.5" pageId="5" pageNumber="50" unit="mm" value="0.475" valueMax="0.5" valueMin="0.45">0.450.50 mm</quantity>
; HW/HL = 0.98, 0.941.07); eyes small and occupying approximately a third of head length (OL = 0.15,
<quantity id="D8679B25FF80663CFC7F329EA0D3F977" box="[1015,1182,1725,1749]" metricMagnitude="-4" metricUnit="m" metricValue="1.5500000000000003" metricValueMax="1.7000000000000002" metricValueMin="1.4000000000000001" pageId="5" pageNumber="50" unit="mm" value="0.155" valueMax="0.17" valueMin="0.14">0.140.17 mm</quantity>
; OL/HL = 0.32, 0.27 0.34).
</paragraph>
<paragraph id="1F2036C0FF80663CFF4F3325A555F8E3" blockId="5.[151,1437,1149,2037]" pageId="5" pageNumber="50">
Pronotum slightly transverse (PW =
<quantity id="D8679B25FF80663CFDED3326A680F8BF" box="[613,717,1797,1821]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="50" unit="mm" value="0.55">0.55 mm</quantity>
,
<quantity id="D8679B25FF80663CFD513326A732F8BF" box="[729,895,1797,1821]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.699999999999999" metricValueMin="5.4" pageId="5" pageNumber="50" unit="mm" value="0.5549999999999999" valueMax="0.57" valueMin="0.54">0.540.57 mm</quantity>
; PL =
<quantity id="D8679B25FF80663CFC443326A079F8BF" box="[972,1076,1797,1821]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="50" unit="mm" value="0.52">0.52 mm</quantity>
,
<quantity id="D8679B25FF80663CFBB73326A0ABF8BF" box="[1087,1254,1797,1821]" metricMagnitude="-4" metricUnit="m" metricValue="5.15" metricValueMax="5.3" metricValueMin="5.0" pageId="5" pageNumber="50" unit="mm" value="0.515" valueMax="0.53" valueMin="0.5">0.500.53 mm</quantity>
; PW/PL = 1.07, 1.061.09).
</paragraph>
<paragraph id="1F2036C0FF80663CFF4F336EA1D1F8C7" blockId="5.[151,1437,1149,2037]" box="[199,1436,1869,1893]" pageId="5" pageNumber="50">
Elytra, together, transverse (EW = 0.59,
<quantity id="D8679B25FF80663CFD02336EA77DF8C7" box="[650,816,1869,1893]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.0" metricValueMin="5.699999999999999" pageId="5" pageNumber="50" unit="mm" value="0.585" valueMax="0.6" valueMin="0.57">0.570.60 mm</quantity>
; EL = 0.44,
<quantity id="D8679B25FF80663CFC31336EA012F8C7" box="[953,1119,1869,1893]" metricMagnitude="-4" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="5" pageNumber="50" unit="mm" value="0.435" valueMax="0.45" valueMin="0.42">0.420.45 mm</quantity>
; EW/EL = 1.35, 1.311.38).
</paragraph>
<paragraph id="1F2036C0FF80663FFF4F3352A6D6FE9D" blockId="5.[151,1437,1149,2037]" lastBlockId="6.[151,1436,151,356]" lastPageId="6" lastPageNumber="51" pageId="5" pageNumber="50">
Males with median lobe elliptical in lateral view (
<figureCitation id="87A42A45FF80663CFD783352A70DF82B" box="[752,832,1905,1929]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 12</figureCitation>
) and limuloid in parameral view (
<figureCitation id="87A42A45FF80663CFB333352A140F82B" box="[1211,1293,1905,1929]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="5" pageNumber="50">Fig. 13</figureCitation>
); apical lobe of median lobe long, extending anteriorly in a nearly horizontal fashion, with apex abruptly diverting paramerally; athetine bridge incomplete at aparameral surface, with distal apodeme eight times its width; foramen situated slightly posterior of middle; proximal costa short, occupying approximately two thirds of posterior surface posterior of foramen; median costa extending to apical lobe; arcuate costa running parallel with median costa basally but never touching, broadly diverging until reaching apical lobe, distally recurving back to foramen; paramere (
<figureCitation id="87A42A45FF83663FFACB349FA1D9FF76" box="[1347,1428,188,212]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="6" pageNumber="51">Fig. 14</figureCitation>
) with length greater than two times its width, velum, in outer lateral view, not obscuring apical lobe; parameral apical lobe (
<figureCitation id="87A42A45FF83663FFF773527A51DFEBE" box="[255,336,260,284]" captionStart="FIGURES 12 19" captionStartId="6.[151,264,1916,1939]" captionTargetBox="[181,1397,421,1887]" captionTargetId="figure@6.[168,1418,411,1895]" captionTargetPageId="6" captionText="FIGURES 12 19. Male genitalia of Drusilla ashei (12 15) and D. nearctica (16 19). 12, 16) aedeagus, lateral; 13, 17) aedeagus, parameral; 14, 18) paramere, outer lateral; 15, 19) apex of apical lobe of paramere, inner lateral. Scale bars (0.1 mm): I = 12 14, 16 18; II = 15, 19." httpUri="https://zenodo.org/record/210862/files/figure.png" pageId="6" pageNumber="51">Fig. 15</figureCitation>
) in inner lateral view with four setulae: setulae a and d on inner surface, b and c on outer surface; setula a longest; b, c and d subequal in length.
</paragraph>
</subSubSection>
<subSubSection id="5785654BFF83663FFF4F356FA13BFEC6" box="[199,1398,332,356]" pageId="6" pageNumber="51" type="etymology">
<paragraph id="1F2036C0FF83663FFF4F356FA13BFEC6" blockId="6.[151,1436,151,356]" box="[199,1398,332,356]" pageId="6" pageNumber="51">
<emphasis id="2DEBEAD2FF83663FFF4F356FA503FEC6" bold="true" box="[199,334,332,356]" pageId="6" pageNumber="51">Etymology.</emphasis>
In honor of the only collector of this species, the late Dr. J.S. Ashe noun in the genitive case.
</paragraph>
</subSubSection>
</treatment>
</document>