184 lines
24 KiB
XML
184 lines
24 KiB
XML
<document id="A28086DA844C42165904B05723A231CB" ID-DOI="10.5281/zenodo.5286268" ID-ISSN="0253-116X" ID-Zenodo-Dep="5286268" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630047656076" checkinUser="marcus" docAuthor="Zwick, Peter" docDate="2015" docId="951DB80C6B29FFF886C8FADDDFF80C0F" docLanguage="en" docName="LinzerbiolBeitr.2015.47.2.1885-1895.pdf" docOrigin="Linzer biologische Beiträge 47 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Prionocyphon alexanderi Zwick 2015, nov.sp." docType="treatment" docVersion="3" lastPageNumber="1889" masterDocId="6924C0746B28FFFC867AFFCDDD5D0F7B" masterDocTitle="Three new Marsh Beetles (Col.: Scirtidae) from New Guinea and Java" masterLastPageNumber="1895" masterPageNumber="1885" pageNumber="1886" updateTime="1699140026292" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="7533B6927B58B187E182DCD1FC246081">
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<mods:title id="8CB3D42E0DBF0AD331E26E74FFAECDFC">Three new Marsh Beetles (Col.: Scirtidae) from New Guinea and Java</mods:title>
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<mods:name id="08016C00AD0E00E7DA19DC412B20A01A" type="personal">
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<mods:namePart id="2395C5A5F1B921E57719DD5825125ACF">Zwick, Peter</mods:namePart>
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<mods:title id="9D976A2B2B2F8213315D76C667B85495">Linzer biologische Beiträge</mods:title>
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<mods:part id="F0DB3876CFB89B8E53BACC48006EFB41">
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<mods:date id="79254695C06A9B450E0F6BB289797A2B">2015</mods:date>
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<mods:number id="E9FF261F593B5FD602CB8A150BF427B6">2015-12-30</mods:number>
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<mods:number id="C845B358D45D3E9301777D7F96BC4D4F">47</mods:number>
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<mods:number id="9462E448C279A996CC37702ECC176D5D">2</mods:number>
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<mods:start id="C9E9BEF7127702B6B5E05607B968175C">1885</mods:start>
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<mods:end id="CD74A7FE7FF4591517285A698DFBCB3E">1895</mods:end>
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<mods:identifier id="CF19B0FC8D7DF0165A576F807F6F6A02" type="DOI">10.5281/zenodo.5286268</mods:identifier>
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<mods:identifier id="9AFA64E30B15864A2485F097DA99394B" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="BE3B7F344BF05CB26E15A9E2AE37DD9B" type="Zenodo-Dep">5286268</mods:identifier>
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<treatment id="951DB80C6B29FFF886C8FADDDFF80C0F" LSID="urn:lsid:plazi:treatment:951DB80C6B29FFF886C8FADDDFF80C0F" httpUri="http://treatment.plazi.org/id/951DB80C6B29FFF886C8FADDDFF80C0F" lastPageId="4" lastPageNumber="1889" pageId="1" pageNumber="1886">
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<subSubSection id="55AE5A916B29FFFD86C8FADDD94D0A30" pageId="1" pageNumber="1886" type="nomenclature">
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<paragraph id="1D0B091A6B29FFFD86C8FADDDF240A5D" blockId="1.[178,633,1296,1318]" box="[178,633,1296,1318]" pageId="1" pageNumber="1886">
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<heading id="4643BE766B29FFFD86C8FADDDF240A5D" box="[178,633,1296,1318]" fontSize="8" level="2" pageId="1" pageNumber="1886" reason="2">
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<emphasis id="2FC0D5086B29FFFD86C8FADDDF240A5D" bold="true" box="[178,633,1296,1318]" pageId="1" pageNumber="1886">
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<taxonomicName id="DAB472996B29FFFD86C8FADDDCE60A5D" authority="Zwick, 2015" authorityName="Zwick" authorityYear="2015" box="[178,443,1296,1318]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="1886" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
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<emphasis id="2FC0D5086B29FFFD86C8FADDDCE60A5D" bold="true" box="[178,443,1296,1318]" italics="true" pageId="1" pageNumber="1886">Prionocyphon alexanderi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34F368736B29FFFD87BBFADDDF530A5E" box="[449,526,1296,1317]" pageId="1" pageNumber="1886" rank="species">nov.sp.</taxonomicNameLabel>
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(Figs 1-9)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="1D0B091A6B29FFFD86C8FAFBD94D0A30" blockId="1.[178,1040,1333,1355]" box="[178,1040,1333,1355]" pageId="1" pageNumber="1886">
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H o l o t y p e:
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<collectingCountry id="65A3498A6B29FFFD8733FAFBDCA10A30" box="[329,508,1333,1355]" name="Papua New Guinea" pageId="1" pageNumber="1886">Papua New Guinea</collectingCountry>
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:
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<collectingRegion id="DF70C7F86B29FFFD847FFAF8DF100A30" box="[517,589,1333,1355]" country="Papua New Guinea" name="Morobe" pageId="1" pageNumber="1886">Morobe</collectingRegion>
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<quantity id="DA4CA4FF6B29FFFD8429FAF8DF9A0A31" box="[595,711,1333,1354]" metricMagnitude="3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="1" pageNumber="1886" unit="m" value="1150.0" valueMax="1300.0" valueMin="1000.0">1000-1300m</quantity>
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<date id="690A2FDA6B29FFFD84B6FAF8DE6E0A31" box="[716,819,1333,1354]" pageId="1" pageNumber="1886" value="1992-10-13">13.10.1992</date>
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leg. A. Riedel (SMNS).
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</paragraph>
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</subSubSection>
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<subSubSection id="55AE5A916B29FFF886C8FA97DFF80C0F" lastPageId="4" lastPageNumber="1889" pageId="1" pageNumber="1886" type="description">
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<paragraph id="1D0B091A6B29FFFD86C8FA97D9610A9D" blockId="1.[178,1086,1370,1510]" pageId="1" pageNumber="1886">
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Habitus. Almost hemispherical, rounded and strongly domed, BL
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<quantity id="DA4CA4FF6B29FFFD85EEFA97DE800A14" box="[916,989,1370,1391]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="1" pageNumber="1886" unit="mm" value="3.8">3.8mm</quantity>
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, BL/BW ~1.4, PW corresponds to 87%, HCW to 50% of BW. Punctures on head, pronotum and scutellum very fine and widely spaced, on elytra coarse and very dense, about 1 diameter apart. Head, pronotum, scutellum and antennae orange-red, legs brown, elytra black except caudal fifth orange. All of dorsal side covered with semi-erect whitish hairs (Fig. 1).
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</paragraph>
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<paragraph id="1D0B091A6B29FFFD86C8FA38DEFD093D" blockId="1.[178,1085,1525,1606]" pageId="1" pageNumber="1886">Frontoclypeus with straight front edge, no projecting lobes. Eyes large, bulging. Mandibles not studied, maxillary palpus unmodified, last segment of labial palpus inserting at right angle near middle of the transversely extended penultimate segment.</paragraph>
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<paragraph id="1D0B091A6B2AFFFE84FAFF16DEC10C41" blockId="2.[640,1086,219,826]" pageId="2" pageNumber="1887">
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Antenna with large flat scape, the small tubular pedicel inserted on the sharp edge of the scape. Antennomere 3 minute, looking like an annulus at the base of antennomere 4. Antennomeres 4-7 strongly, 8 moderately serrate, 9-11 about tubular (Fig. 1). SAR ending where it meets the eye, not descending in front of the eye. The wide area between eye and edge of oral cavity shallowly concave. End of SGR on lower face of head slightly raised, separated from edge of oral cavity by a small gap ("buttonhole configuration",
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<bibRefCitation id="792574EB6B2AFFFE851DFDF7D95D0D2B" author="ZWICK P" box="[871,1024,570,592]" pageId="2" pageNumber="1887" pagination="163 - 189" refId="ref5081" refString="ZWICK P. (2013 b): Australian Marsh Beetles (Coleoptera: Scirtidae). 3. A restricted concept of genus Cyphon, Australian species of Cyphon s. str., and the new Australasian genus Nanocyphon. - Genus 24: 163 - 189." type="journal article" year="2013">ZWICK 2013b</bibRefCitation>
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), not connected to the edge of the oral cavity, nor to other ridges and sutures. Prosternal process and mesoventral groove not studied. Abdomen unmodified, all sternites densely covered with strong socketed hairs, no obvious sensilla except a few along the front edge of S3. Density of pilosity does not differ between sternites.
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</paragraph>
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<paragraph id="1D0B091A6B2AFFFE84FAFC86D9600C1B" blockId="2.[178,1086,842,1159]" box="[640,1085,842,864]" pageId="2" pageNumber="1887">M a l e. T8 with wide plate, a large forward</paragraph>
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<paragraph id="1D0B091A6B2AFFFE84FAFCAAD9600C06" blockId="2.[178,1086,842,1159]" box="[640,1085,871,893]" pageId="2" pageNumber="1887">directed tongue-shaped lobe between the</paragraph>
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<paragraph id="1D0B091A6B2AFFFE84FAFC48D9600CE0" blockId="2.[178,1086,842,1159]" box="[640,1085,901,923]" pageId="2" pageNumber="1887">straight apodemes. Caudal margin triangu-</paragraph>
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<paragraph id="1D0B091A6B2AFFFE84FAFC6FD9600C88" blockId="2.[178,1086,842,1159]" pageId="2" pageNumber="1887">
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lar, setal fringe near middle very long (Fig.
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<emphasis id="2FC0D5086B2AFFFE86C2FC04DDB30CA5" bold="true" box="[184,238,969,991]" pageId="2" pageNumber="1887">Fig. 1</emphasis>
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:
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<taxonomicName id="DAB472996B2AFFFE8680FC04DCBF0CA5" authority="Zwick, 2015" authorityName="Zwick" authorityYear="2015" box="[250,482,969,990]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1887" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
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<emphasis id="2FC0D5086B2AFFFE8680FC04DCBF0CA5" box="[250,482,969,990]" italics="true" pageId="2" pageNumber="1887">Prionocyphon alexanderi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34F368736B2AFFFE8792FC07DF730CA4" box="[488,558,970,991]" pageId="2" pageNumber="1887" rank="species">nov.sp.</taxonomicNameLabel>
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: male, 2). S8 large, a wide U-shaped sclerite, no habitus, dorsal. setae (Fig. 3). Apodemes of T9 closer to-
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</paragraph>
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<paragraph id="1D0B091A6B2AFFFE84FAFC36DE840BFD" blockId="2.[178,1086,842,1159]" pageId="2" pageNumber="1887">gether, connected by a sclerite arch. Plate unpigmented, very wide, its sides partly folded mediad. Caudally stand a few very fine setae on either side of a transverse soft field covered with weak microtrichia (Fig. 4). S9 long and slender, with strong concave sclerite strips supporting the caudally widening plate. A tuft of long setae on either side of midline, near caudal margin (Fig. 5).</paragraph>
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<paragraph id="1D0B091A6B2AFFFE86C8FB5ADCC30AA8" blockId="2.[178,1086,1174,1491]" pageId="2" pageNumber="1887">Tegmen (Fig. 6) with bilobed wide base, the sclerotized area about as long as wide, narrowing caudally and supporting two slender caudally tapering straight parameres. Their soft tips bear some sensory pores, converge, and meet. On the outside the parameres bear numerous thick socketed spines, the mesal edge is smooth. Each spine is parallel over most of its length and apically pointed. In front, spines form an oval patch on the base of the paramere. Near the widest point of the tegmen insert exceptionally strong and long styles whose distinct sclerotized bases curve mediad. Each style is about tubular, with a weak constriction near middle. Apex gently curved outward, outer face with a soft membranous subterminal area. A fringe of long hair-like microtrichia along medial edge takes a hairpin bend at the apex and continues forward for a short distance along a low middle rib.</paragraph>
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<paragraph id="1D0B091A6B2AFFFF86C8FA2EDFD10A8B" blockId="2.[178,1085,1507,1587]" lastBlockId="3.[178,1086,1453,1520]" lastPageId="3" lastPageNumber="1888" pageId="2" pageNumber="1887">
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Penis long and slender, parallel over 2/3 of its length, the slender parameroids converge a little (Fig. 7). A basal lobe of the parameroid extends slightly mediad, over the basal arm of the trigonium. The parameroids are slightly waisted near middle and end in narrow
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<emphasis id="2FC0D5086B2BFFFF86C8FA60DC5C0AB9" bold="true" box="[178,257,1453,1475]" pageId="3" pageNumber="1888">Figs 2-9</emphasis>
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:
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<taxonomicName id="DAB472996B2BFFFF876BFA60DCA10AB9" authority="Zwick, 2015" authorityName="Zwick" authorityYear="2015" box="[273,508,1453,1474]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1888" phylum="Arthropoda" rank="species" species="alexanderi" status="sp. nov.">
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<emphasis id="2FC0D5086B2BFFFF876BFA60DCA10AB9" box="[273,508,1453,1474]" italics="true" pageId="3" pageNumber="1888">Prionocyphon alexanderi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34F368736B2BFFFF847FFA60DF140AB9" box="[517,585,1453,1474]" pageId="3" pageNumber="1888" rank="species">nov.sp.</taxonomicNameLabel>
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, male terminalia: (
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<emphasis id="2FC0D5086B2BFFFF8487FA60DE550AB9" bold="true" box="[765,776,1453,1474]" pageId="3" pageNumber="1888">2</emphasis>
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) T8; (
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<emphasis id="2FC0D5086B2BFFFF8533FA60DE090AB9" bold="true" box="[841,852,1453,1474]" pageId="3" pageNumber="1888">3</emphasis>
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) S8; (
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<emphasis id="2FC0D5086B2BFFFF85EEFA60DEC20AB9" bold="true" box="[916,927,1453,1474]" pageId="3" pageNumber="1888">4</emphasis>
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) T9; (
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<emphasis id="2FC0D5086B2BFFFF859AFA60DEB60AB9" bold="true" box="[992,1003,1453,1474]" pageId="3" pageNumber="1888">5</emphasis>
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) S9; (
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<emphasis id="2FC0D5086B2BFFFF8250FA60D9680AB9" bold="true" box="[1066,1077,1453,1474]" pageId="3" pageNumber="1888">6</emphasis>
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) tegmen with parameres and styli; (
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<emphasis id="2FC0D5086B2BFFFF878AFA09DCA60AA2" bold="true" box="[496,507,1476,1497]" pageId="3" pageNumber="1888">7</emphasis>
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) penis; (
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<emphasis id="2FC0D5086B2BFFFF8434FA09DF040AA2" bold="true" box="[590,601,1476,1497]" pageId="3" pageNumber="1888">8</emphasis>
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) trigonium, enlarged; (
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<emphasis id="2FC0D5086B2BFFFF8555FA09DE670AA2" bold="true" box="[815,826,1476,1497]" pageId="3" pageNumber="1888">9</emphasis>
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) detail of armature on paramere. 2-7 to the same scale. pe, paramere; st, stylus.
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</paragraph>
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<paragraph id="1D0B091A6B2CFFF886C8FF16DE940EC4" blockId="4.[178,1086,219,447]" pageId="4" pageNumber="1889">flat lobes. Along the medial face the parameroids are covered with flat rounded scales. These are directed backward on the wide basal lobes, are unordered at the narrowest point of the parameroid and point forward on the narrow caudal section. The basal arms of the trigonium converge and meet, forming a parallel sclerite with subterminal triangular expansions and ending in a slender cone (Fig. 8). Ventrally, the distal part of the trigonium bears two irregular rows of strong pointed cones giving it a serrate appearance (Fig. 9). Base of endophallus (Fig. 7) without visible armature. Soft endophallus tissue seems to fill the entire space between the trigonium and the long parameroids.</paragraph>
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<paragraph id="1D0B091A6B2CFFF886C8FE02DCD70E9E" blockId="4.[178,394,463,485]" box="[178,394,463,485]" pageId="4" pageNumber="1889">F e m a l e. Unknown.</paragraph>
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<paragraph id="1D0B091A6B2CFFF886C8FE38DE2A0C4A" blockId="4.[178,1086,501,817]" pageId="4" pageNumber="1889">
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N o t e s. There are several colourful
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<taxonomicName id="DAB472996B2CFFF8845CFE38DE550D70" box="[550,776,501,523]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1889" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="2FC0D5086B2CFFF8845CFE38DFE80D71" box="[550,693,501,522]" italics="true" pageId="4" pageNumber="1889">Prionocyphon</emphasis>
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species
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</taxonomicName>
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,
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<taxonomicName id="DAB472996B2CFFF88568FE38DEC70D71" authorityName="Zwick" authorityYear="2015" box="[786,922,501,522]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1889" phylum="Arthropoda" rank="species" species="alexanderi">
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<emphasis id="2FC0D5086B2CFFF88568FE38DEC70D71" box="[786,922,501,522]" italics="true" pageId="4" pageNumber="1889">P. alexanderi</emphasis>
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</taxonomicName>
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resembles none of them. Details of genitalia differ clearly from all congeners.
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<taxonomicName id="DAB472996B2CFFF88540FDDFD9600D5C" authorityName="Zwick" authorityYear="2015" box="[826,1085,530,551]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1889" phylum="Arthropoda" rank="species" species="alexanderi">
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<emphasis id="2FC0D5086B2CFFF88540FDDFD9600D5C" box="[826,1085,530,551]" italics="true" pageId="4" pageNumber="1889">Prionocyphon alexanderi</emphasis>
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</taxonomicName>
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is more similar to the Holarctic congeners (
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<bibRefCitation id="792574EB6B2CFFF884BBFDFDDE3B0D3E" author="NYHOLM T" box="[705,870,560,581]" pageId="4" pageNumber="1889" pagination="28 - 42" refId="ref4604" refString="NYHOLM T. (1971): Beitrage zur Kenntnis der palaarktischen Helodiden 2. Bemerkungen uber die Gattung Prionocyphon REDTB. Mit Beschreibung zweier neuer Arten aus dem Mittelmeergebiet Studien uber die Familie Helodidae. XII. - Entomologisk Tidskrift 92: 28 - 42." type="journal article" year="1971">NYHOLM 1971</bibRefCitation>
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,
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<bibRefCitation id="792574EB6B2CFFF88507FDFDD9640D3E" author="YOSHITOMI H" box="[893,1081,559,581]" pageId="4" pageNumber="1889" pagination="1 - 212" refId="ref4909" refString="YOSHITOMI H. (2005): Systematic revision of the family Scirtidae of Japan, with phylogeny, morphology and bionomics (Insecta: Coleoptera, Scirtoidea). - Japanese Journal of Systematic Entomology, Monographic Series 3: 1 - 212." type="book chapter" year="2005">YOSHITOMI 2005</bibRefCitation>
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,
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<bibRefCitation id="792574EB6B2CFFF886C8FD80DC260D18" author="KLAUSNITZER B" box="[178,379,589,611]" pageId="4" pageNumber="1889" refId="ref4368" refString="KLAUSNITZER B. (2009): Insecta: Coleoptera: Scirtidae. - Susswasserfauna von Mitteleuropa 20 / 17. Heidelberg: Spektrum Akademischer Verlag, XIV + 326 pp." type="book" year="2009">KLAUSNITZER 2009</bibRefCitation>
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) than to the Taiwanese species (
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<bibRefCitation id="792574EB6B2CFFF884B6FD80DE190D19" author="RUTA R" box="[716,836,589,610]" pageId="4" pageNumber="1889" pagination="52 - 60" refId="ref4707" refString="RUTA R. (2010): Two new species of Prionocyphon REDTENBACHER from Taiwan (Coleoptera: Scirtidae), with notes on Prionocyphon, Mescirtes MOTSCHULSKY and Prionoscirtes CHAMPION from East and Southeast Asia. - Zootaxa 2402: 52 - 60." type="journal article" year="2010">RUTA 2010</bibRefCitation>
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,
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<bibRefCitation id="792574EB6B2CFFF8852AFD80D95E0D18" author="YOSHITOMI H" box="[848,1027,589,611]" pageId="4" pageNumber="1889" pagination="529 - 533" refId="ref4954" refString="YOSHITOMI H. (2010): A new species of Prionocyphon from Taiwan (Coleoptera: Scirtidae: Scirtinae). - Acta Entomologica Musei Nationalis Pragae 50: 529 - 533." type="journal article" year="2010">YOSHITOMI 2010</bibRefCitation>
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). The forward extension of T8 and the bilobed base of tegmen resemble
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<emphasis id="2FC0D5086B2CFFF885D0FDA6D9600DFB" box="[938,1085,619,640]" italics="true" pageId="4" pageNumber="1889">P. serricornis</emphasis>
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(
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<bibRefCitation id="792574EB6B2CFFF886C3FD45DC080DE5" author="MULLER Ph" box="[185,341,646,670]" pageId="4" pageNumber="1889" pagination="184 - 230" refId="ref4559" refString="MULLER Ph. W. J. (1821): Neue Insekten, beschrieben von Ph. W. J. Muller. - Magazin der Entomologie. Herausgegeben von E. F. Germar (Halle) 4: 184 - 230." type="journal article" year="1821">MÜLLER, 1821</bibRefCitation>
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) and the other west-palaearctic species. No Holarctic species seems to have styles, but the "Seitenplättchen" or "Lateralgriffel" which
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<bibRefCitation id="792574EB6B2CFFF8852BFD6BDEA90DC1" author="NYHOLM T" box="[849,1012,677,699]" pageId="4" pageNumber="1889" pagination="28 - 42" refId="ref4604" refString="NYHOLM T. (1971): Beitrage zur Kenntnis der palaarktischen Helodiden 2. Bemerkungen uber die Gattung Prionocyphon REDTB. Mit Beschreibung zweier neuer Arten aus dem Mittelmeergebiet Studien uber die Familie Helodidae. XII. - Entomologisk Tidskrift 92: 28 - 42." type="journal article" year="1971">NYHOLM (1971</bibRefCitation>
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,
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<bibRefCitation id="792574EB6B2CFFF88278FD68D96B0DC0" author="NYHOLM T" box="[1026,1078,677,699]" pageId="4" pageNumber="1889" pagination="81 - 119" refId="ref4651" refString="NYHOLM T. (1972): Zur Morphologie und Funktion des Helodiden-Aedoeagus (Col.). - Entomologica Scandinavica 3: 81 - 119." type="journal article" year="1972">1972</bibRefCitation>
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) described are their homologues; see ZWICK (in prep.) Styles occur in neotropical
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<taxonomicName id="DAB472996B2CFFF886C8FD2DDCCA0D8D" box="[178,407,736,758]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1889" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="2FC0D5086B2CFFF886C8FD2DDC1C0D8E" box="[178,321,736,757]" italics="true" pageId="4" pageNumber="1889">Prionocyphon</emphasis>
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species
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</taxonomicName>
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(
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<bibRefCitation id="792574EB6B2CFFF887D6FD2DDFD60D8D" author="KLAUSNITZER B" box="[428,651,736,758]" pageId="4" pageNumber="1889" pagination="1177 - 1194" refId="ref4475" refString="KLAUSNITZER B. (2012 b): Zur Kenntnis der Gattung Prionocyphon L. REDTENBACHER 1858 sensu lato (Coleoptera, Scirtidae) aus Sudamerika (181. Beitrag zur Kenntnis der Scirtidae). - Linzer biologische Beitrage 44 (2): 1177 - 1194." type="journal article" year="2012">KLAUSNITZER 2012b</bibRefCitation>
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), in
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<taxonomicName id="DAB472996B2CFFF884BEFD2CDE150D8D" authority="Zwick, 2015" authorityName="Zwick" authorityYear="2015" box="[708,840,737,758]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1889" phylum="Arthropoda" rank="species" species="papuanus" status="sp. nov.">
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<emphasis id="2FC0D5086B2CFFF884BEFD2CDE150D8D" box="[708,840,737,758]" italics="true" pageId="4" pageNumber="1889">P. papuanus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="34F368736B2CFFF8852CFD2DDEFF0D8E" box="[854,930,736,757]" pageId="4" pageNumber="1889" rank="species">nov.sp.</taxonomicNameLabel>
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, and in many Australian
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<taxonomicName id="DAB472996B2CFFF8875EFD30DCEE0C69" box="[292,435,765,786]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1891" phylum="Arthropoda" rank="genus">
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<emphasis id="2FC0D5086B2CFFF8875EFD30DCEE0C69" box="[292,435,765,786]" italics="true" pageId="4" pageNumber="1889">Prionocyphon</emphasis>
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</taxonomicName>
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(
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<bibRefCitation id="792574EB6B2CFFF887BEFD33DF100C68" author="WATTS C" box="[452,589,766,787]" pageId="4" pageNumber="1889" pagination="53 - 88" refId="ref4752" refString="WATTS C. H. S. (2010): Revision of Australian Prionocyphon REDTENBACHER (Scirtidae: Coleoptera). - Transactions of the Royal Society of South Australia (2010) 134: 53 - 88." type="journal article" year="2010">WATTS 2010</bibRefCitation>
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, ZWICK in prep.). However, the complex structure and the strength of the styles of
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<taxonomicName id="DAB472996B2CFFF8845EFCD1DFF10C4B" authorityName="Zwick" authorityYear="2015" box="[548,684,795,817]" class="Insecta" family="Scirtidae" genus="Prionocyphon" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1889" phylum="Arthropoda" rank="species" species="alexanderi">
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<emphasis id="2FC0D5086B2CFFF8845EFCD1DFF10C4B" box="[548,684,795,817]" italics="true" pageId="4" pageNumber="1889">P. alexanderi</emphasis>
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</taxonomicName>
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(Fig. 8) are unique.
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</paragraph>
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<paragraph id="1D0B091A6B2CFFF886C8FC8CDFF80C0F" blockId="4.[178,1085,833,884]" pageId="4" pageNumber="1889">
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E t y m o l o g y. Named for ALEXANDER RIEDEL who collected not only this specimen but many other
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<taxonomicName id="DAB472996B2CFFF88751FC93DF470C0F" authority="(ZWICK 2014)" baseAuthorityName="ZWICK" baseAuthorityYear="2014" box="[299,538,862,884]" class="Insecta" family="Scirtidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1889" phylum="Arthropoda" rank="family">
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Scirtidae (
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<bibRefCitation id="792574EB6B2CFFF887E9FC93DF4E0C0F" author="ZWICK P" box="[403,531,862,884]" pageId="4" pageNumber="1889" pagination="131 - 152" refId="ref5131" refString="ZWICK P. (2014): Australian Marsh Beetles (Coleoptera: Scirtidae). 5. New species of genus Cyphon (s. str.), and Papuacyphon gen. nov. from New Guinea. - Stuttgarter Beitrage zur Naturkunde, Neue Serie 7: 131 - 152." type="journal article" year="2014">ZWICK 2014</bibRefCitation>
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)
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</taxonomicName>
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in Irian Jaya.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |