treatments-xml/data/5E/6B/87/5E6B87F2FFB5FFAA2B9EFA58FB13D66A.xml
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<document id="5B27E82773F084B59EB354CF702C18A5" ID-DOI="10.11646/zootaxa.5360.1.3" ID-GBIF-Dataset="9807050a-8ef5-4251-a385-11eecbb06227" ID-ISSN="1175-5326" ID-Zenodo-Dep="10084843" ID-ZooBank="1A2295D0-E7D6-4738-BD23-E0743E3F19CA" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1698938070764" checkinUser="plazi" docAuthor="Bang, Woo Jun &amp; Shin, Seunggwan" docDate="2023" docId="5E6B87F2FFB5FFAA2B9EFA58FB13D66A" docLanguage="en" docName="zootaxa.5360.1.3.pdf" docOrigin="Zootaxa 5360 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5360.1.3/52105" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chaoborus pseudoflavicans Bang &amp; Shin 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="63" masterDocId="A252FF8AFFB7FFAC2B09FFC9FFFED04D" masterDocTitle="A new species of the Chaoborus flavicans complex (Diptera, Chaoboridae) in South Korea" masterLastPageNumber="81" masterPageNumber="57" pageNumber="59" updateTime="1699699869657" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="9171D2010C2461A76326AEA6C335A958">A new species of the Chaoborus flavicans complex (Diptera, Chaoboridae) in South Korea</mods:title>
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<mods:namePart id="9FE6A438CC33CD70276D4E8E8B066A7E">Bang, Woo Jun</mods:namePart>
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<mods:affiliation id="1309DCCC2AED27261953E4B22C537E54">School of Biological Sciences, Seoul 08826, Seoul National University, Republic of Korea &amp; Comparative Medicine Disease Research Center, Seoul National University, Seoul 08826, Republic of Korea</mods:affiliation>
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<mods:namePart id="45CC919A8BD4B63EA12DA471EE139AC3">Shin, Seunggwan</mods:namePart>
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<mods:affiliation id="537F90EEF1791B6361EDD5BAE8EF7664">School of Biological Sciences, Seoul 08826, Seoul National University, Republic of Korea &amp; Comparative Medicine Disease Research Center, Seoul National University, Seoul 08826, Republic of Korea</mods:affiliation>
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<mods:date id="4C7A15E20ADDFBE4C2739AC026B151D7">2023</mods:date>
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<treatment id="5E6B87F2FFB5FFAA2B9EFA58FB13D66A" ID-GBIF-Taxon="217220394" LSID="urn:lsid:plazi:treatment:5E6B87F2FFB5FFAA2B9EFA58FB13D66A" httpUri="http://treatment.plazi.org/id/5E6B87F2FFB5FFAA2B9EFA58FB13D66A" lastPageId="6" lastPageNumber="63" pageId="2" pageNumber="59">
<subSubSection id="9ED8656FFFB5FFAE2B9EFA58FDB5D5E1" box="[151,587,1425,1452]" pageId="2" pageNumber="59" type="nomenclature">
<paragraph id="D67D36E4FFB5FFAE2B9EFA58FDB5D5E1" blockId="2.[151,587,1425,1452]" box="[151,587,1425,1452]" pageId="2" pageNumber="59">
<heading id="8D358188FFB5FFAE2B9EFA58FDB5D5E1" bold="true" box="[151,587,1425,1452]" fontSize="11" level="1" pageId="2" pageNumber="59" reason="1">
<emphasis id="E4B6EAF6FFB5FFAE2B9EFA58FDB5D5E1" bold="true" box="[151,587,1425,1452]" pageId="2" pageNumber="59">
<taxonomicName id="11C24D67FFB5FFAE2B9EFA58FE18D5E1" authority="Bang &amp; Shin, 2023" authorityName="Bang &amp; Shin" authorityYear="2023" box="[151,486,1425,1452]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="pseudoflavicans" status="sp. nov.">
<emphasis id="E4B6EAF6FFB5FFAE2B9EFA58FE18D5E1" bold="true" box="[151,486,1425,1452]" italics="true" pageId="2" pageNumber="59">Chaoborus pseudoflavicans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF85578DFFB5FFAE2AE4FA5BFDB5D5E1" box="[493,587,1426,1452]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9ED8656FFFB5FFAE2B9EFA1FFCFBD624" pageId="2" pageNumber="59" type="reference_group">
<paragraph id="D67D36E4FFB5FFAE2B9EFA1FFD24D5A3" blockId="2.[151,795,1494,1641]" box="[151,730,1494,1519]" pageId="2" pageNumber="59">
<treatmentCitationGroup id="F6D211CAFFB5FFAE2B9EFA1FFD24D5A3" box="[151,730,1494,1519]" pageId="2" pageNumber="59">
<taxonomicName id="11C24D67FFB5FFAE2B9EFA1FFE7AD5A3" ID-CoL="8BFQH" baseAuthorityName="De Geer" baseAuthorityYear="1776" box="[151,388,1494,1518]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="crystallinus">
<emphasis id="E4B6EAF6FFB5FFAE2B9EFA1FFE7AD5A3" box="[151,388,1494,1518]" italics="true" pageId="2" pageNumber="59">Chaoborus crystallinus</emphasis>
</taxonomicName>
(nec De Geer):
<treatmentCitation id="576310F5FFB5FFAE2921FA1EFD28D5A3" author="Komyo, E." box="[552,726,1494,1519]" page="12" pageId="2" pageNumber="59" year="1954">
<bibRefCitation id="B2534B15FFB5FFAE2921FA1EFD28D5A3" author="Komyo, E." box="[552,726,1494,1519]" pageId="2" pageNumber="59" pagination="11 - 15" refId="ref6831" refString="Komyo, E. (1954) Japanese Chaoborin Mosquitoes. The Scientific Reports of the Saikyo University Agriculture, 6, 11 - 15." type="journal article" year="1954">Komyo 1954: 12</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D67D36E4FFB5FFAE2B9EFA3CFCE5D640" blockId="2.[151,795,1494,1641]" box="[151,795,1525,1549]" pageId="2" pageNumber="59">
<treatmentCitationGroup id="F6D211CAFFB5FFAE2B9EFA3CFCE5D640" box="[151,795,1525,1549]" pageId="2" pageNumber="59">
<taxonomicName id="11C24D67FFB5FFAE2B9EFA3CFE77D640" ID-CoL="8BFQJ" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[151,393,1525,1549]" class="Insecta" family="Chaoboridae" genus="Chaoborus" isUncertain="true" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB5FFAE2B9EFA3CFEF6D640" box="[151,264,1525,1549]" italics="true" pageId="2" pageNumber="59">Chaoborus</emphasis>
cf.
<emphasis id="E4B6EAF6FFB5FFAE2A27FA3CFE77D640" box="[302,393,1525,1549]" italics="true" pageId="2" pageNumber="59">flavicans</emphasis>
</taxonomicName>
(nec Meigen):
<treatmentCitation id="576310F5FFB5FFAE292CFA3CFCE9D640" author="Dupuis, D. &amp; Svensson, J. - E. &amp; Taylor, D. J." box="[549,791,1525,1549]" page="240" pageId="2" pageNumber="59" year="2008">
<bibRefCitation id="B2534B15FFB5FFAE292CFA3CFCE9D640" author="Dupuis, D. &amp; Svensson, J. - E. &amp; Taylor, D. J." box="[549,791,1525,1549]" pageId="2" pageNumber="59" pagination="236 - 243" refId="ref6346" refString="Dupuis, D., Svensson, J. - E. &amp; Taylor, D. J. (2008) The cryptic origins of environment-indicating phantom midges (Chaoborus). Limnology and Oceanography, 53, 236 - 243. https: // doi. org / 10.4319 / lo. 2008.53.1.0236" type="journal article" year="2008">
Dupuis
<emphasis id="E4B6EAF6FFB5FFAE297CFA3FFD5CD640" box="[629,674,1525,1549]" italics="true" pageId="2" pageNumber="59">et al</emphasis>
. 2008: 240
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D67D36E4FFB5FFAE2B9EF9DDFD3CD661" blockId="2.[151,795,1494,1641]" box="[151,706,1556,1580]" pageId="2" pageNumber="59">
<treatmentCitationGroup id="F6D211CAFFB5FFAE2B9EF9DDFD3CD661" box="[151,706,1556,1580]" pageId="2" pageNumber="59">
<taxonomicName id="11C24D67FFB5FFAE2B9EF9DDFE97D661" ID-CoL="8BFQJ" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[151,361,1556,1580]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB5FFAE2B9EF9DDFE97D661" box="[151,361,1556,1580]" italics="true" pageId="2" pageNumber="59">Chaoborus flavicans</emphasis>
</taxonomicName>
(nec Meigen):
<treatmentCitation id="576310F5FFB5FFAE290DF9DDFD40D661" author="An, H. &amp; Jung, G. &amp; Kim, C. - B." box="[516,702,1556,1580]" page="39" pageId="2" pageNumber="59" year="2012">
<bibRefCitation id="B2534B15FFB5FFAE290DF9DDFD40D661" author="An, H. &amp; Jung, G. &amp; Kim, C. - B." box="[516,702,1556,1580]" pageId="2" pageNumber="59" pagination="36 - 41" refId="ref5995" refString="An, H., Jung, G. &amp; Kim, C. - B. (2012) Molecular taxonomy of a phantom midge species (Chaoborus flavicans) in Korea. Animal Systematics, Evolution and Diversity, 28 (1), 36 - 41. https: // doi. org / 10.5635 / ASED. 2012.28.1.036" type="journal article" year="2012">
An
<emphasis id="E4B6EAF6FFB5FFAE2920F9DDFDA8D661" box="[553,598,1556,1580]" italics="true" pageId="2" pageNumber="59">et al</emphasis>
. 2012: 39
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D67D36E4FFB5FFAE2B9EF9FBFD40D607" blockId="2.[151,795,1494,1641]" box="[151,702,1586,1610]" pageId="2" pageNumber="59">
<treatmentCitationGroup id="F6D211CAFFB5FFAE2B9EF9FBFD40D607" box="[151,702,1586,1610]" pageId="2" pageNumber="59">
<taxonomicName id="11C24D67FFB5FFAE2B9EF9FBFED5D607" box="[151,299,1586,1610]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E4B6EAF6FFB5FFAE2B9EF9FBFEF6D607" box="[151,264,1586,1610]" italics="true" pageId="2" pageNumber="59">Chaoborus</emphasis>
sp.
</taxonomicName>
(unidentified):
<treatmentCitation id="576310F5FFB5FFAE2AC3F9FBFD47D607" author="Zhang, X. &amp; Kang, Z. &amp; Ding, S. &amp; Wang, Y. &amp; Borkent, C. &amp; Saigusa, T. &amp; Yang, D." box="[458,697,1586,1610]" page="747" pageId="2" pageNumber="59" year="2019">
<bibRefCitation id="B2534B15FFB5FFAE2AC3F9FBFD47D607" author="Zhang, X. &amp; Kang, Z. &amp; Ding, S. &amp; Wang, Y. &amp; Borkent, C. &amp; Saigusa, T. &amp; Yang, D." box="[458,697,1586,1610]" pageId="2" pageNumber="59" pagination="747" refId="ref7482" refString="Zhang, X., Kang, Z., Ding, S., Wang, Y., Borkent, C., Saigusa, T. &amp; Yang, D. (2019) Mitochondrial Genomes Provide Insights into the Phylogeny of Culicomorpha (Insecta: Diptera). International journal of molecular sciences, 20 (3), 747. https: // doi. org / 10.3390 / ijms 20030747" type="journal article" year="2019">
Zhang
<emphasis id="E4B6EAF6FFB5FFAE2918F9FAFDC0D607" box="[529,574,1586,1610]" italics="true" pageId="2" pageNumber="59">et al</emphasis>
. 2019: 747
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D67D36E4FFB5FFAE2B9EF998FCFBD624" blockId="2.[151,795,1494,1641]" box="[151,773,1617,1641]" pageId="2" pageNumber="59">
<treatmentCitationGroup id="F6D211CAFFB5FFAE2B9EF998FCFBD624" box="[151,773,1617,1641]" pageId="2" pageNumber="59">
<taxonomicName id="11C24D67FFB5FFAE2B9EF998FEA2D625" box="[151,348,1617,1641]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E4B6EAF6FFB5FFAE2B9EF998FEF6D624" box="[151,264,1617,1641]" italics="true" pageId="2" pageNumber="59">Chaoborus</emphasis>
JPN sp.
</taxonomicName>
(unidentified):
<treatmentCitation id="576310F5FFB5FFAE2AF2F998FCFFD624" author="Salmela, J. &amp; Harma, O. &amp; Taylor, D. J." box="[507,769,1617,1641]" page="181" pageId="2" pageNumber="59" year="2021">
<bibRefCitation id="B2534B15FFB5FFAE2AF2F998FCFFD624" author="Salmela, J. &amp; Harma, O. &amp; Taylor, D. J." box="[507,769,1617,1641]" pageId="2" pageNumber="59" pagination="151 - 196" refId="ref7342" refString="Salmela, J., Harma, O. &amp; Taylor, D. J. (2021 b) Chaoborus flavicans Meigen (Diptera, Chaoboridae) is a complex of lake and pond dwelling species: a revision. Zootaxa, 4927 (2), 151 - 196. https: // doi. org / 10.11646 / zootaxa. 4927.2.1" type="journal article" year="2021">
Salmela
<emphasis id="E4B6EAF6FFB5FFAE295BF99BFD81D624" box="[594,639,1617,1641]" italics="true" pageId="2" pageNumber="59">et al</emphasis>
. 2021b: 181
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9ED8656FFFB5FFAF2B9EF95EFB96D10C" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59" type="diagnosis">
<paragraph id="D67D36E4FFB5FFAE2B9EF95EFE60D6FC" blockId="2.[151,1436,1687,2002]" box="[151,414,1687,1713]" pageId="2" pageNumber="59">
<heading id="8D358188FFB5FFAE2B9EF95EFE60D6FC" bold="true" box="[151,414,1687,1713]" fontSize="11" level="1" pageId="2" pageNumber="59" reason="1">
<emphasis id="E4B6EAF6FFB5FFAE2B9EF95EFE60D6FC" bold="true" box="[151,414,1687,1713]" pageId="2" pageNumber="59">Differential diagnosis.</emphasis>
</heading>
</paragraph>
<paragraph id="D67D36E4FFB5FFAE2BCEF972FD20D7C4" blockId="2.[151,1436,1687,2002]" pageId="2" pageNumber="59">
<emphasis id="E4B6EAF6FFB5FFAE2BCEF972FEABD698" bold="true" box="[199,341,1723,1749]" pageId="2" pageNumber="59">Adult male.</emphasis>
See Samela
<emphasis id="E4B6EAF6FFB5FFAE2AE2F975FDE1D698" box="[491,543,1723,1749]" italics="true" pageId="2" pageNumber="59">et al</emphasis>
. (2021b) to see the illustrations of
<taxonomicName id="11C24D67FFB5FFAE28BCF975FBC3D698" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[949,1085,1723,1750]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB5FFAE28BCF975FC36D69B" box="[949,968,1724,1750]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE28D1F972FBC3D698" box="[984,1085,1723,1749]" italics="true" pageId="2" pageNumber="59">flavicans</emphasis>
</taxonomicName>
species complex. This species can be distinguished from
<taxonomicName id="11C24D67FFB5FFAE2AC2F929FDD5D6B4" authorityName="Salmela" authorityYear="2021" box="[459,555,1760,1786]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="posio">
<emphasis id="E4B6EAF6FFB5FFAE2AC2F929FE20D6B7" box="[459,478,1760,1786]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2AE7F929FDD5D6B4" box="[494,555,1760,1785]" italics="true" pageId="2" pageNumber="59">posio</emphasis>
</taxonomicName>
by its yellowish coloration (dark-almost in
<taxonomicName id="11C24D67FFB5FFAE2F21F929FB76D6B4" authorityName="Salmela" authorityYear="2021" box="[1064,1160,1760,1786]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="posio">
<emphasis id="E4B6EAF6FFB5FFAE2F21F929FBC5D6B7" box="[1064,1083,1760,1786]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2F42F929FB76D6B4" box="[1099,1160,1760,1785]" italics="true" pageId="2" pageNumber="59">posio</emphasis>
</taxonomicName>
), and from
<taxonomicName id="11C24D67FFB5FFAE2E1DF929FA62D6B4" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[1300,1436,1759,1786]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB5FFAE2E1DF929FAD9D6B7" box="[1300,1319,1760,1786]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2E3EF916FA62D6B4" box="[1335,1436,1759,1785]" italics="true" pageId="2" pageNumber="59">flavicans</emphasis>
</taxonomicName>
and
<taxonomicName id="11C24D67FFB5FFAE2BC3F8CDFEC0D750" baseAuthorityName="Johannsen" baseAuthorityYear="1903" box="[202,318,1795,1822]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="E4B6EAF6FFB5FFAE2BC3F8CDFF23D753" box="[202,221,1796,1822]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2BE4F8CAFEC0D750" box="[237,318,1795,1821]" italics="true" pageId="2" pageNumber="59">albipes</emphasis>
</taxonomicName>
by the following combination of characters: Tergal bands absent (present in
<taxonomicName id="11C24D67FFB5FFAE2FB4F8CDFAB8D750" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[1213,1350,1795,1822]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB5FFAE2FB4F8CDFB2ED753" box="[1213,1232,1796,1822]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2FE8F8CAFAB8D750" box="[1249,1350,1795,1821]" italics="true" pageId="2" pageNumber="59">flavicans</emphasis>
</taxonomicName>
and
<taxonomicName id="11C24D67FFB5FFAE2E8AF8CDFF16D70C" baseAuthorityName="Johannsen" baseAuthorityYear="1903" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="E4B6EAF6FFB5FFAE2E8AF8CDFA68D753" box="[1411,1430,1796,1822]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2B9EF8EEFF16D70C" box="[151,232,1831,1857]" italics="true" pageId="2" pageNumber="59">albipes</emphasis>
</taxonomicName>
). Paramere unicolorous, constricted and medially bent, apical claw stout and curved (bicolorous, constricted and medially bent, apical claw narrow and slightly curved in
<taxonomicName id="11C24D67FFB5FFAE2846F885FC28D728" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[847,982,1867,1894]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB5FFAE2846F885FC9CD72B" box="[847,866,1868,1894]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2878F882FC28D728" box="[881,982,1867,1893]" italics="true" pageId="2" pageNumber="59">flavicans</emphasis>
</taxonomicName>
and nearly unicolorous, slightly bent or straight, apical claw stout and curved in
<taxonomicName id="11C24D67FFB5FFAE2955F8B9FD33D7C4" baseAuthorityName="Johannsen" baseAuthorityYear="1903" box="[604,717,1903,1930]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="E4B6EAF6FFB5FFAE2955F8B9FD91D7C7" box="[604,623,1904,1930]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2975F8A6FD33D7C4" box="[636,717,1903,1929]" italics="true" pageId="2" pageNumber="59">albipes</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="D67D36E4FFB5FFAF2BCEF85AFE36D098" blockId="2.[151,1436,1687,2002]" lastBlockId="3.[151,1436,151,537]" lastPageId="3" lastPageNumber="60" pageId="2" pageNumber="59">
<emphasis id="E4B6EAF6FFB5FFAE2BCEF85AFEF3D7E0" bold="true" box="[199,269,1939,1965]" pageId="2" pageNumber="59">Pupa.</emphasis>
This species may be hardly distinguished due to intraspecific variation in respiratory organ morphology according to its aquatic habitats (pond and lake). Respiratory organs subapically constricted (absent in
<taxonomicName id="11C24D67FFB5FFAE2E24F871FA72D79C" authorityName="Salmela" authorityYear="2021" box="[1325,1420,1976,2002]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="59" phylum="Arthropoda" rank="species" species="posio">
<emphasis id="E4B6EAF6FFB5FFAE2E24F871FABED79F" box="[1325,1344,1976,2002]" italics="true" pageId="2" pageNumber="59">C</emphasis>
.
<emphasis id="E4B6EAF6FFB5FFAE2E46F871FA72D79C" box="[1359,1420,1976,2001]" italics="true" pageId="2" pageNumber="59">posio</emphasis>
</taxonomicName>
). Length of respiratory horn mostly&gt; 1000 μm and slender form (mostly &lt;1000 μm in
<taxonomicName id="11C24D67FFB4FFAF2F8CFF5EFB04D0FC" baseAuthorityName="Johannsen" baseAuthorityYear="1903" box="[1157,1274,151,177]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="E4B6EAF6FFB4FFAF2F8CFF5EFB66D0FC" box="[1157,1176,151,177]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF2FA0FF5EFB04D0FC" box="[1193,1274,151,177]" italics="true" pageId="3" pageNumber="60">albipes</emphasis>
</taxonomicName>
and
<taxonomicName id="11C24D67FFB4FFAF2E3CFF5EFA68D0FD" authorityName="Salmela" authorityYear="2021" box="[1333,1430,151,177]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="posio">
<emphasis id="E4B6EAF6FFB4FFAF2E3CFF5EFAB6D0FC" box="[1333,1352,151,177]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF2E50FF5EFA68D0FD" box="[1369,1430,151,176]" italics="true" pageId="3" pageNumber="60">posio</emphasis>
</taxonomicName>
, medially wide in
<taxonomicName id="11C24D67FFB4FFAF2A53FF72FE46D099" authorityName="Salmela" authorityYear="2021" box="[346,440,187,213]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="posio">
<emphasis id="E4B6EAF6FFB4FFAF2A53FF72FE93D098" box="[346,365,187,213]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF2A72FF72FE46D099" box="[379,440,187,212]" italics="true" pageId="3" pageNumber="60">posio</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="D67D36E4FFB4FFAF2BCEFF16FB96D10C" blockId="3.[151,1436,151,537]" pageId="3" pageNumber="60">
<emphasis id="E4B6EAF6FFB4FFAF2BCEFF16FE7ED0B4" bold="true" box="[199,384,222,249]" pageId="3" pageNumber="60">IV instar larva.</emphasis>
Mandibular lateral teeth conspicuous, first lateral tooth about as long as mandibular tooth III (short or inconspicuous in
<taxonomicName id="11C24D67FFB4FFAF2ACBFECAFDB6D150" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[450,584,259,285]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB4FFAF2ACBFECAFE2BD150" box="[450,469,259,285]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF2AEAFECAFDB6D150" box="[483,584,259,285]" italics="true" pageId="3" pageNumber="60">flavicans</emphasis>
</taxonomicName>
), number of lateral teeth almost &lt;4 (almost&gt;
<quantity id="113A9B01FFB4FFAF2F5DFECBFB81D150" box="[1108,1151,258,285]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="3" pageNumber="60" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="11C24D67FFB4FFAF2F8FFECAFB06D150" baseAuthorityName="Johannsen" baseAuthorityYear="1903" box="[1158,1272,259,285]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="E4B6EAF6FFB4FFAF2F8FFECAFB67D150" box="[1158,1177,259,285]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF2FAEFECAFB06D150" box="[1191,1272,259,285]" italics="true" pageId="3" pageNumber="60">albipes</emphasis>
</taxonomicName>
and
<taxonomicName id="11C24D67FFB4FFAF2E27FECAFA72D151" authorityName="Salmela" authorityYear="2021" box="[1326,1420,259,285]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="posio">
<emphasis id="E4B6EAF6FFB4FFAF2E27FECAFABFD150" box="[1326,1345,259,285]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF2E46FECAFA72D151" box="[1359,1420,259,284]" italics="true" pageId="3" pageNumber="60">posio</emphasis>
</taxonomicName>
), number of mandibular fan bristles usually 10 (number&gt;
<quantity id="113A9B01FFB4FFAF281AFEEEFCB2D10C" box="[787,844,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="60" unit="in" value="20.0">20 in</quantity>
<taxonomicName id="11C24D67FFB4FFAF285BFEEEFC3AD10C" baseAuthorityName="Johannsen" baseAuthorityYear="1903" box="[850,964,295,321]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="E4B6EAF6FFB4FFAF285BFEEEFC9BD10C" box="[850,869,295,321]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF287AFEEEFC3AD10C" box="[883,964,295,321]" italics="true" pageId="3" pageNumber="60">albipes</emphasis>
</taxonomicName>
and
<taxonomicName id="11C24D67FFB4FFAF28F3FEEEFBA6D10D" authorityName="Salmela" authorityYear="2021" box="[1018,1112,295,321]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="60" phylum="Arthropoda" rank="species" species="posio">
<emphasis id="E4B6EAF6FFB4FFAF28F3FEEEFBF3D10C" box="[1018,1037,295,321]" italics="true" pageId="3" pageNumber="60">C</emphasis>
.
<emphasis id="E4B6EAF6FFB4FFAF2F12FEEEFBA6D10D" box="[1051,1112,295,320]" italics="true" pageId="3" pageNumber="60">posio</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9ED8656FFFB4FFAF2BCEFE82FD4FD254" pageId="3" pageNumber="60" type="materials_examined">
<paragraph id="D67D36E4FFB4FFAF2BCEFE82FD4FD254" blockId="3.[151,1436,151,537]" pageId="3" pageNumber="60">
<emphasis id="E4B6EAF6FFB4FFAF2BCEFE82FE83D129" bold="true" box="[199,381,330,357]" pageId="3" pageNumber="60">Type materials.</emphasis>
<materialsCitation id="66AA3CB9FFB4FFAF2A88FE82FD33D1C4" ID-GBIF-Occurrence="4434745301" collectingDate="2023-02-01" collectorName="Woo Jun Bang &amp; Jeungjun Lee" country="South Korea" latitude="34.46842" location="Haenam-gun" longLatPrecision="1" longitude="126.51731" pageId="3" pageNumber="60" specimenCount="1" specimenCount-male="1" stateProvince="Jeollanam-do" typeStatus="holotype">
<emphasis id="E4B6EAF6FFB4FFAF2A88FE82FE0BD128" bold="true" box="[385,501,331,357]" pageId="3" pageNumber="60">
<typeStatus id="09798846FFB4FFAF2A88FE82FE0ED128" box="[385,496,331,357]" pageId="3" pageNumber="60" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="C0C4FD6DFFB4FFAF2AF1FE82FDCCD128" box="[504,562,331,357]" pageId="3" pageNumber="60" type="male">Male</specimenCount>
(SNUE),
<date id="A27C1024FFB4FFAF2994FE82FCEAD128" box="[669,788,331,357]" pageId="3" pageNumber="60" value="2023-02-01">
<collectingDate id="B238E9CCFFB4FFAF2994FE82FCEAD128" box="[669,788,331,357]" pageId="3" pageNumber="60" value="2023-02-01">01.II.2023</collectingDate>
</date>
,
<location id="D31D603FFFB4FFAF2815FE82FC4ED128" LSID="urn:lsid:plazi:treatment:5E6B87F2FFB5FFAA2B9EFA58FB13D66A:D31D603FFFB4FFAF2815FE82FC4ED128" box="[796,944,331,357]" country="South Korea" latitude="34.46842" longLatPrecision="1" longitude="126.51731" name="Haenam-gun" pageId="3" pageNumber="60" stateProvince="Jeollanam-do">Haenam-gun</location>
,
<collectingRegion id="1406F806FFB4FFAF28B1FE82FBACD128" box="[952,1106,331,357]" country="South Korea" name="Jeonnam" pageId="3" pageNumber="60">Jeollanam-do</collectingRegion>
,
<collectingCountry id="AED57674FFB4FFAF2F53FE82FB18D128" box="[1114,1254,331,357]" name="South Korea" pageId="3" pageNumber="60">South Korea</collectingCountry>
,
<geoCoordinate id="B3F65023FFB4FFAF2FE7FE82FA66D128" box="[1262,1432,331,357]" degrees="34" direction="north" minutes="28" orientation="latitude" pageId="3" pageNumber="60" precision="1" seconds="06.30" value="34.46842">34°28'06.30&quot;N</geoCoordinate>
,
<geoCoordinate id="B3F65023FFB4FFAF2B9EFEA6FEB5D1C4" box="[151,331,367,393]" degrees="126" direction="east" minutes="31" orientation="longitude" pageId="3" pageNumber="60" precision="1" seconds="02.31" value="126.51731">126°31'02.31&quot;E</geoCoordinate>
,
<collectorName id="7B375332FFB4FFAF2A5FFEA6FE04D1C4" box="[342,506,367,393]" pageId="3" pageNumber="60">Woo Jun Bang</collectorName>
and
<collectorName id="7B375332FFB4FFAF2939FEA6FD34D1C4" box="[560,714,367,393]" pageId="3" pageNumber="60">Jeungjun Lee</collectorName>
.
</materialsCitation>
<materialsCitation id="66AA3CB9FFB4FFAF29DAFEA6FE8CD1E0" ID-GBIF-Occurrence="4434745302" collectionCode="SNUE, Regional museum of Lapland, LMM" country="Finland" location="Rovaniemi" pageId="3" pageNumber="60" specimenCount="272" specimenCount-larva="110" specimenCount-pupa="110" stateProvince="Rovaniemi" typeStatus="paratype">
<emphasis id="E4B6EAF6FFB4FFAF29DAFEA6FCACD1C4" bold="true" box="[723,850,367,393]" pageId="3" pageNumber="60">
<typeStatus id="09798846FFB4FFAF29DAFEA6FCB3D1C4" box="[723,845,367,393]" pageId="3" pageNumber="60" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="C0C4FD6DFFB4FFAF2850FEA6FC05D1C4" box="[857,1019,367,393]" pageId="3" pageNumber="60">20 adult males</specimenCount>
and
<specimenCount id="C0C4FD6DFFB4FFAF2F3BFEA6FB13D1C4" box="[1074,1261,367,393]" pageId="3" pageNumber="60">20 adult females</specimenCount>
,
<specimenCount id="C0C4FD6DFFB4FFAF2FF1FEA6FA93D1C4" box="[1272,1389,367,393]" count="100" pageId="3" pageNumber="60" type="larva">100 larvae</specimenCount>
and
<specimenCount id="C0C4FD6DFFB4FFAF2B9EFE5AFEF4D1E0" box="[151,266,403,429]" count="100" pageId="3" pageNumber="60" type="pupa">100 pupae</specimenCount>
(
<collectionCode id="B0D3AE21FFB4FFAF2A1EFE5AFE98D1E0" box="[279,358,403,429]" collectionName="SNUE" pageId="3" pageNumber="60">SNUE</collectionCode>
);
</materialsCitation>
<materialsCitation id="66AA3CB9FFB4FFAF2A71FE5AFDA1D19C" ID-GBIF-Occurrence="4434745303" collectionCode="SNUE, Regional museum of Lapland, LMM" country="Finland" location="Rovaniemi" pageId="3" pageNumber="60" specimenCount="272" specimenCount-larva="110" specimenCount-pupa="110" stateProvince="Rovaniemi" typeStatus="paratype">
<specimenCount id="C0C4FD6DFFB4FFAF2A71FE5AFDF2D1E0" box="[376,524,403,429]" pageId="3" pageNumber="60">6 adult males</specimenCount>
,
<specimenCount id="C0C4FD6DFFB4FFAF291FFE5AFD3ED1E0" box="[534,704,403,429]" pageId="3" pageNumber="60">6 adult females</specimenCount>
, and
<specimenCount id="C0C4FD6DFFB4FFAF29FEFE5AFCA3D1E0" box="[759,861,403,429]" count="10" pageId="3" pageNumber="60" type="larva">10 larvae</specimenCount>
and
<specimenCount id="C0C4FD6DFFB4FFAF2899FE5AFC0BD1E0" box="[912,1013,403,429]" count="10" pageId="3" pageNumber="60" type="pupa">10 pupae</specimenCount>
are deposited at the
<collectionCode id="B0D3AE21FFB4FFAF2FDAFE5AFEEAD19C" pageId="3" pageNumber="60">Regional museum of Lapland</collectionCode>
,
<collectingRegion id="1406F806FFB4FFAF2A17FE7EFE62D19C" box="[286,412,439,465]" pageId="3" pageNumber="60">Rovaniemi</collectingRegion>
,
<collectingCountry id="AED57674FFB4FFAF2AAFFE7EFE02D19C" box="[422,508,439,465]" name="Finland" pageId="3" pageNumber="60">Finland</collectingCountry>
(
<collectionCode id="B0D3AE21FFB4FFAF2903FE7EFDADD19C" box="[522,595,439,465]" collectionName="LMM" pageId="3" pageNumber="60">LMM</collectionCode>
).
</materialsCitation>
all materials from same collection data as
<typeStatus id="09798846FFB4FFAF2F30FE7EFB61D19C" box="[1081,1183,439,465]" pageId="3" pageNumber="60" type="holotype">holotype</typeStatus>
.
<specimenCount id="C0C4FD6DFFB4FFAF2FA0FE7EFAFDD19C" box="[1193,1283,439,465]" count="10" pageId="3" pageNumber="60" type="adult">10 adult</specimenCount>
males and
<specimenCount id="C0C4FD6DFFB4FFAF2E89FE7EFF31D1B8" count="10" pageId="3" pageNumber="60" type="adult">10 adult</specimenCount>
females, pinned.
<specimenCount id="C0C4FD6DFFB4FFAF2A9AFE12FE10D1B8" box="[403,494,475,501]" count="10" pageId="3" pageNumber="60" type="adult">10 adult</specimenCount>
males and
<specimenCount id="C0C4FD6DFFB4FFAF2963FE12FD3AD1B8" box="[618,708,475,501]" count="10" pageId="3" pageNumber="60" type="adult">10 adult</specimenCount>
females in 70 % EtOH. All larval and pupal specimens preserved in 70 % EtOH.
<specimenCount id="C0C4FD6DFFB4FFAF2A4DFE36FE3CD254" box="[324,450,511,537]" count="3" pageId="3" pageNumber="60" type="male">Three male</specimenCount>
hypopygia on slides.
</paragraph>
</subSubSection>
<subSubSection id="9ED8656FFFB4FFA82B9EFDA3FB47D709" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="description">
<paragraph id="D67D36E4FFB4FFAF2B9EFDA3FED6D2C9" blockId="3.[151,296,618,644]" box="[151,296,618,644]" pageId="3" pageNumber="60">
<heading id="8D358188FFB4FFAF2B9EFDA3FED6D2C9" bold="true" box="[151,296,618,644]" fontSize="11" level="1" pageId="3" pageNumber="60" reason="1">
<emphasis id="E4B6EAF6FFB4FFAF2B9EFDA3FED6D2C9" bold="true" box="[151,296,618,644]" pageId="3" pageNumber="60">Description</emphasis>
</heading>
</paragraph>
<paragraph id="D67D36E4FFB4FFAF2B9EFD7BFC65D644" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="60">
<emphasis id="E4B6EAF6FFB4FFAF2B9EFD7BFE85D280" bold="true" box="[151,379,690,717]" pageId="3" pageNumber="60">Adult male. Head.</emphasis>
Dark brown with pale setae; Antennae 14-segmented, bearing whorl pale setae; antennal flagellomeres circular form, basal segments dark brown, but distal segments pale; apical flagellomere dark brown, length of penultimate flagellomere 295 (268320), apical flagellomere 224 (183262), penultimate/apical 1.32 (1.03 1.74, n=7); maxillary palpi 5-segmented, all segments greyish brown with dark brown setae. Length of palpomeres 14 (n=5): 1st 76 (6980), 2nd 110 (99120), 3rd 249 (204289), 4th 231 (192256) (
<figureCitation id="4EF92A61FFB4FFAF2F57FC8AFB33D310" box="[1118,1229,835,861]" captionStart="FIGURE 2" captionStartId="4.[152,255,807,831]" captionTargetBox="[168,1436,196,783]" captionTargetId="figure-27@4.[151,1436,181,783]" captionTargetPageId="4" captionText="FIGURE 2. Chaoborus pseudoflavicans sp. nov., adult male habitus in the (a) lateral and (b) dorsal views (scale bar 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.10084847" httpUri="https://zenodo.org/record/10084847/files/figure.png" pageId="3" pageNumber="60">Fig. 2ab</figureCitation>
).
<emphasis id="E4B6EAF6FFB4FFAF2FE9FC8AFABED310" bold="true" box="[1248,1344,835,861]" pageId="3" pageNumber="60">Thorax.</emphasis>
Scutum with trident light brown stripes and some black specks scattered, ground color brown with pale setae; scutellum and mediotergite brown, most pleuron colors composed pale to brown areas: antepronotum, postpronotum, anepimeron, anepisternum, metanepsiternum and katepisternum brown (antepronotum slightly darkened than postpronotum), halteres pale, a number of thorax setae (n=12): antepronotum 26 (2229), postpronotum 5 (36), proepisternum 5 (46), katepisternum 7 (49), anepisternum 11 (812) and anepimeron 5 (59) (
<figureCitation id="4EF92A61FFB4FFAF2F35FC3EFB6ED45C" box="[1084,1168,1015,1041]" captionStart="FIGURE 2" captionStartId="4.[152,255,807,831]" captionTargetBox="[168,1436,196,783]" captionTargetId="figure-27@4.[151,1436,181,783]" captionTargetPageId="4" captionText="FIGURE 2. Chaoborus pseudoflavicans sp. nov., adult male habitus in the (a) lateral and (b) dorsal views (scale bar 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.10084847" httpUri="https://zenodo.org/record/10084847/files/figure.png" pageId="3" pageNumber="60">Fig. 2a</figureCitation>
).
<emphasis id="E4B6EAF6FFB4FFAF2FAFFC3EFB24D45C" bold="true" box="[1190,1242,1015,1041]" pageId="3" pageNumber="60">Leg.</emphasis>
Mostly pale, but fourth-fifth tarsomeres slightly darkened than first to third tarsomeres; foreleg, lengths of femur, tibia and five tarsomeres (n=12): femur 2069 (19922226), tibia 2259 (21232357), tarsomeres: 1st 931 (861992), 2nd 567 (510628), 3rd 483 (440544), 4th 321 (273385), 5th 218 (161248); midleg, lengths of femur, tibia and five tarsomeres (n=12): femur 1714 (16481775), tibia 1716 (16491815), tarsomeres: 1st 735 (701792), 2nd 443 (415471), 3rd 355 (334390), 4th 269 (251298), 5th 235 (220242); hindleg, lengths of femur, tibia and five tarsomeres (n=12): femur 2189 (19702400), tibia 2170 (18062373), tarsomeres: 1st 1250 (11971291), 2nd 730 (686756), 3rd 494 (472524), 4th 308 (295330), 5th 260 (235288) (
<figureCitation id="4EF92A61FFB4FFAF28C4FB3AFBDFD540" box="[973,1057,1267,1293]" captionStart="FIGURE 2" captionStartId="4.[152,255,807,831]" captionTargetBox="[168,1436,196,783]" captionTargetId="figure-27@4.[151,1436,181,783]" captionTargetPageId="4" captionText="FIGURE 2. Chaoborus pseudoflavicans sp. nov., adult male habitus in the (a) lateral and (b) dorsal views (scale bar 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.10084847" httpUri="https://zenodo.org/record/10084847/files/figure.png" pageId="3" pageNumber="60">Fig. 2a</figureCitation>
).
<emphasis id="E4B6EAF6FFB4FFAF2F3EFB3BFB81D541" bold="true" box="[1079,1151,1266,1292]" pageId="3" pageNumber="60">Wing.</emphasis>
mostly light brown with yellowish scales around the margin, length 4238 (41024334), width 881 (867906), length/width 4.8 (4.94.5), fork of R
<subScript id="4A4634A1FFB4FFAF2BF7FA84FEE6D516" attach="left" box="[254,280,1357,1371]" fontSize="6" pageId="3" pageNumber="60">2+3</subScript>
517 (395654), fork of M
<subScript id="4A4634A1FFB4FFAF2948FA84FDA4D516" attach="left" box="[577,602,1357,1371]" fontSize="6" pageId="3" pageNumber="60">1+2</subScript>
461 (400490), R
<subScript id="4A4634A1FFB4FFAF2820FA84FCCFD516" attach="left" box="[809,817,1357,1371]" fontSize="6" pageId="3" pageNumber="60">3</subScript>
1288 (11201385), M
<subScript id="4A4634A1FFB4FFAF2F27FA84FBC8D516" attach="left" box="[1070,1078,1357,1371]" fontSize="6" pageId="3" pageNumber="60">1</subScript>
1136 (10591184) (n=10) (
<figureCitation id="4EF92A61FFB4FFAF2E66FAF2FF49D534" captionStart="FIGURE 2" captionStartId="4.[152,255,807,831]" captionTargetBox="[168,1436,196,783]" captionTargetId="figure-27@4.[151,1436,181,783]" captionTargetPageId="4" captionText="FIGURE 2. Chaoborus pseudoflavicans sp. nov., adult male habitus in the (a) lateral and (b) dorsal views (scale bar 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.10084847" httpUri="https://zenodo.org/record/10084847/files/figure.png" pageId="3" pageNumber="60">Fig. 2b</figureCitation>
).
<emphasis id="E4B6EAF6FFB4FFAF2BC2FA97FEB9D535" bold="true" box="[203,327,1374,1400]" pageId="3" pageNumber="60">Abdomen.</emphasis>
Ground color light brown with black lateral specks but not connected, tergal bands absent, fifth to distal tergal segments slightly darkened (
<figureCitation id="4EF92A61FFB4FFAF2963FA4AFD41D5D0" box="[618,703,1411,1437]" captionStart="FIGURE 2" captionStartId="4.[152,255,807,831]" captionTargetBox="[168,1436,196,783]" captionTargetId="figure-27@4.[151,1436,181,783]" captionTargetPageId="4" captionText="FIGURE 2. Chaoborus pseudoflavicans sp. nov., adult male habitus in the (a) lateral and (b) dorsal views (scale bar 1.0 mm)." figureDoi="http://doi.org/10.5281/zenodo.10084847" httpUri="https://zenodo.org/record/10084847/files/figure.png" pageId="3" pageNumber="60">Fig. 2b</figureCitation>
).
<emphasis id="E4B6EAF6FFB4FFAF29DCFA4BFC86D5D1" bold="true" box="[725,888,1410,1436]" pageId="3" pageNumber="60">Hypopygium.</emphasis>
Gonocoxite light brown, length 538 (512547), width 186 (167206), length/width 2.9 (2.53.3, n=7); gonostylus mostly dark brown, length 485 (449518), width 36 (3239), length/width 16.1911.51 (n=7) (
<figureCitation id="4EF92A61FFB4FFAF299BFA02FD1DD5A8" box="[658,739,1483,1509]" captionStart="FIGURE 3" captionStartId="4.[152,255,1402,1426]" captionTargetBox="[158,1429,898,1372]" captionTargetId="figure-55@4.[151,1436,891,1379]" captionTargetPageId="4" captionText="FIGURE 3. Chaoborus pseudoflavicans sp. nov., hypopygium, (a) dorsal view (scale bar 200 μm), (b) parameres viewed in different positions (scale bar 50 μm). gst = gonostylus, pm = paramere, gx = gonocoxite, ac = apical claw." figureDoi="http://doi.org/10.5281/zenodo.10084851" httpUri="https://zenodo.org/record/10084851/files/figure.png" pageId="3" pageNumber="60">Fig. 3a</figureCitation>
); paramere unicolorous, brown, medially bent and constricted, apical claw stout and curved, length 149 (133160, n=5) (
<figureCitation id="4EF92A61FFB4FFAF2829FA26FC71D644" box="[800,911,1519,1545]" captionStart="FIGURE 3" captionStartId="4.[152,255,1402,1426]" captionTargetBox="[158,1429,898,1372]" captionTargetId="figure-55@4.[151,1436,891,1379]" captionTargetPageId="4" captionText="FIGURE 3. Chaoborus pseudoflavicans sp. nov., hypopygium, (a) dorsal view (scale bar 200 μm), (b) parameres viewed in different positions (scale bar 50 μm). gst = gonostylus, pm = paramere, gx = gonocoxite, ac = apical claw." figureDoi="http://doi.org/10.5281/zenodo.10084851" httpUri="https://zenodo.org/record/10084851/files/figure.png" pageId="3" pageNumber="60">Fig. 3bc</figureCitation>
).
</paragraph>
<paragraph id="D67D36E4FFB4FFAF2BCEF9DBFE98D790" blockId="3.[151,1437,690,2013]" pageId="3" pageNumber="60">
<emphasis id="E4B6EAF6FFB4FFAF2BCEF9DBFE93D661" bold="true" box="[199,365,1554,1580]" pageId="3" pageNumber="60">Adult female.</emphasis>
Most are similar with male.
<emphasis id="E4B6EAF6FFB4FFAF29CFF9DAFCF3D660" bold="true" box="[710,781,1555,1581]" pageId="3" pageNumber="60">Head.</emphasis>
Length of penultimate flagellomere 95 (84115), apical flagellomere 104 (94134). penultimate/apical 0.92 (0.631.30, n=4); maxillary palpi 5-segmented, dark brown, length of palpomeres 14 (n=6): 1st 55 (5163), 2nd 65 (5869), 3rd 174 (149209), 4th 149 (131167).
<emphasis id="E4B6EAF6FFB4FFAF2E34F992FA62D638" bold="true" box="[1341,1436,1627,1653]" pageId="3" pageNumber="60">Thorax.</emphasis>
Number of setae on thorax (n=8): antepronotum 20 (1722), postpronotum 5 (56), proepisternum 4 (35), katepisternum 5 (36), anepisternum 11 (1013), anepimerum 11 (611).
<emphasis id="E4B6EAF6FFB4FFAF28DDF96AFBF6D6F0" bold="true" box="[980,1032,1699,1725]" pageId="3" pageNumber="60">Leg.</emphasis>
Foreleg, lengths of femur, tibia and five tarsomeres (n=6): femur 1591 (15431663), tibia 1846 (17761902), tarsomeres: 1st 800 (736874), 2nd 484 (437511), 3rd 385 (337423), 4th 255 (227282), 5th 180 (162209); midleg, lengths of femur, tibia and five tarsomeres (n=4): femur 1506 (14421596), tibia 1456 (14201495), tarsomeres: 1st 631 (600652), 2nd 354 (322 394), 3rd 286 (281296), 4th 199 (182213), 5th 168 (153180); hindleg, lengths of femur, tibia and five tarsomeres (n=5): femur 1814 (17251861), tibia 1836 (17731916), tarsomeres: 1st 1098 (10301166), 2nd 609 (562644), 3rd 400 (349445), 4th 236 (212263), 5th 175 (161194).
<emphasis id="E4B6EAF6FFB4FFAF2838F8B3FC8CD7D9" bold="true" box="[817,882,1914,1940]" pageId="3" pageNumber="60">Wing</emphasis>
. length 3441 (33833517), width 945 (879991), length/width 3.6 (3.44.0), fork of R
<subScript id="4A4634A1FFB4FFAF293AF878FDB2D7F2" attach="left" box="[563,588,1969,1983]" fontSize="6" pageId="3" pageNumber="60">2+3</subScript>
358 (290406), fork of M
<subScript id="4A4634A1FFB4FFAF287CF878FC71D7F2" attach="left" box="[885,911,1969,1983]" fontSize="6" pageId="3" pageNumber="60">1+2</subScript>
303 (273320), R
<subScript id="4A4634A1FFB4FFAF2F54F878FB9BD7F2" attach="left" box="[1117,1125,1969,1983]" fontSize="6" pageId="3" pageNumber="60">3</subScript>
1207 (9341296), M
<subScript id="4A4634A1FFB4FFAF2E5FF878FAA0D7F2" attach="left" box="[1366,1374,1969,1983]" fontSize="6" pageId="3" pageNumber="60">1</subScript>
1105 (8321254) (n=6).
</paragraph>
<caption id="82BD666CFFB3FFA82B91FCEEFF30D32E" ID-DOI="http://doi.org/10.5281/zenodo.10084847" ID-Zenodo-Dep="10084847" httpUri="https://zenodo.org/record/10084847/files/figure.png" pageId="4" pageNumber="61" startId="4.[152,255,807,831]" targetBox="[168,1436,196,783]" targetPageId="4" targetType="figure">
<paragraph id="D67D36E4FFB3FFA82B91FCEEFF30D32E" blockId="4.[151,1437,807,867]" pageId="4" pageNumber="61">
<emphasis id="E4B6EAF6FFB3FFA82B91FCEEFEE5D372" bold="true" box="[152,283,807,831]" pageId="4" pageNumber="61">FIGURE 2.</emphasis>
<taxonomicName id="11C24D67FFB3FFA82A2CFCEEFDBCD372" authority="Bang &amp; Shin, 2023" authorityName="Bang &amp; Shin" authorityYear="2023" box="[293,578,807,831]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="pseudoflavicans" status="sp. nov.">
<emphasis id="E4B6EAF6FFB3FFA82A2CFCEEFDBCD372" box="[293,578,807,831]" italics="true" pageId="4" pageNumber="61">Chaoborus pseudoflavicans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF85578DFFB3FFA82945FCEEFD5FD373" box="[588,673,807,831]" pageId="4" pageNumber="61" rank="species">
<emphasis id="E4B6EAF6FFB3FFA82945FCEEFD9AD372" bold="true" box="[588,612,807,831]" pageId="4" pageNumber="61">sp</emphasis>
.
<emphasis id="E4B6EAF6FFB3FFA8297AFCEEFD64D372" bold="true" box="[627,666,807,831]" pageId="4" pageNumber="61">nov</emphasis>
.
</taxonomicNameLabel>
, adult male habitus in the (a) lateral and (b) dorsal views (scale bar 1.0 mm).
</paragraph>
</caption>
<caption id="82BD666CFFB3FFA82B91FAB3FB37D5FA" ID-DOI="http://doi.org/10.5281/zenodo.10084851" ID-Zenodo-Dep="10084851" httpUri="https://zenodo.org/record/10084851/files/figure.png" pageId="4" pageNumber="61" startId="4.[152,255,1402,1426]" targetBox="[158,1429,898,1372]" targetPageId="4" targetType="figure">
<paragraph id="D67D36E4FFB3FFA82B91FAB3FB37D5FA" blockId="4.[151,1437,1402,1463]" pageId="4" pageNumber="61">
<emphasis id="E4B6EAF6FFB3FFA82B91FAB3FEE7D5DF" bold="true" box="[152,281,1402,1426]" pageId="4" pageNumber="61">FIGURE 3.</emphasis>
<taxonomicName id="11C24D67FFB3FFA82A28FAB2FDC2D5DE" authority="Bang &amp; Shin, 2023" authorityName="Bang &amp; Shin" authorityYear="2023" box="[289,572,1403,1427]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="61" phylum="Arthropoda" rank="species" species="pseudoflavicans" status="sp. nov.">
<emphasis id="E4B6EAF6FFB3FFA82A28FAB2FDC2D5DE" box="[289,572,1403,1427]" italics="true" pageId="4" pageNumber="61">Chaoborus pseudoflavicans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF85578DFFB3FFA8294DFAB2FD69D5DF" box="[580,663,1403,1427]" pageId="4" pageNumber="61" rank="species">
<emphasis id="E4B6EAF6FFB3FFA8294DFAB2FDA2D5DE" bold="true" box="[580,604,1403,1427]" pageId="4" pageNumber="61">sp</emphasis>
.
<emphasis id="E4B6EAF6FFB3FFA82963FAB2FD6FD5DE" bold="true" box="[618,657,1403,1427]" pageId="4" pageNumber="61">nov</emphasis>
.
</taxonomicNameLabel>
, hypopygium, (a) dorsal view (scale bar 200 μm), (b) parameres viewed in different positions (scale bar 50 μm). gst = gonostylus, pm = paramere, gx = gonocoxite, ac = apical claw.
</paragraph>
</caption>
<paragraph id="D67D36E4FFB3FFA82BCEF9C0FBFDD6C2" blockId="4.[151,1437,1545,1860]" pageId="4" pageNumber="61">
<emphasis id="E4B6EAF6FFB3FFA82BCEF9C0FEFBD66E" bold="true" box="[199,261,1545,1571]" pageId="4" pageNumber="61">Pupa</emphasis>
(
<figureCitation id="4EF92A61FFB3FFA82A1CF9C0FE96D66E" box="[277,360,1545,1571]" captionStart="FIGURE 4" captionStartId="5.[152,255,1195,1219]" captionTargetBox="[151,1408,181,1171]" captionTargetId="figure-19@5.[151,1435,181,1171]" captionTargetPageId="5" captionText="FIGURE 4. Chaoborus pseudoflavicans sp. nov., immature stages, (a) larva, habitus in lateral view (scale bar 1.0 mm), (b) pupa, habitus in lateral view (scale bar 1.0 mm), (c) larval mandibles (scale bar 50 μm), (d) larval blades (scale bar 50 μm), (e) respiratory organ (scale bar 500 μm). lb = larval blade, ant = antenna, md = mandible, ans = anal fan setae, ro = respiratory organ, tp = terminal process, Lt = lateral teeth, IIV = mandibular teeth, III being the subordinate tooth." figureDoi="http://doi.org/10.5281/zenodo.10084855" httpUri="https://zenodo.org/record/10084855/files/figure.png" pageId="4" pageNumber="61">Fig. 4b</figureCitation>
). Mib rib of terminal process incomplete, not reaching the distal margin of paddles, and curved apically; outer rib of terminal process curved shape and smooth with no spines; mid rib and lateral rib almost pale, but mid rib slightly darkened than lateral rib; respiratory organ, slender form, subapically constricted, length 1141 (10161187), width 225 (205309), length/width 4.6 (3.35.9, n=10) (
<figureCitation id="4EF92A61FFB3FFA828A5F9BCFC02D6C2" box="[940,1020,1653,1679]" captionStart="FIGURE 4" captionStartId="5.[152,255,1195,1219]" captionTargetBox="[151,1408,181,1171]" captionTargetId="figure-19@5.[151,1435,181,1171]" captionTargetPageId="5" captionText="FIGURE 4. Chaoborus pseudoflavicans sp. nov., immature stages, (a) larva, habitus in lateral view (scale bar 1.0 mm), (b) pupa, habitus in lateral view (scale bar 1.0 mm), (c) larval mandibles (scale bar 50 μm), (d) larval blades (scale bar 50 μm), (e) respiratory organ (scale bar 500 μm). lb = larval blade, ant = antenna, md = mandible, ans = anal fan setae, ro = respiratory organ, tp = terminal process, Lt = lateral teeth, IIV = mandibular teeth, III being the subordinate tooth." figureDoi="http://doi.org/10.5281/zenodo.10084855" httpUri="https://zenodo.org/record/10084855/files/figure.png" pageId="4" pageNumber="61">Fig. 4e</figureCitation>
)
</paragraph>
<paragraph id="D67D36E4FFB3FFA82BCEF950FB47D709" blockId="4.[151,1437,1545,1860]" pageId="4" pageNumber="61">
<emphasis id="E4B6EAF6FFB3FFA82BCEF950FE74D6FE" bold="true" box="[199,394,1689,1715]" pageId="4" pageNumber="61">IV instar larvae</emphasis>
(
<figureCitation id="4EF92A61FFB3FFA82A95F950FE11D6FE" box="[412,495,1689,1715]" captionStart="FIGURE 4" captionStartId="5.[152,255,1195,1219]" captionTargetBox="[151,1408,181,1171]" captionTargetId="figure-19@5.[151,1435,181,1171]" captionTargetPageId="5" captionText="FIGURE 4. Chaoborus pseudoflavicans sp. nov., immature stages, (a) larva, habitus in lateral view (scale bar 1.0 mm), (b) pupa, habitus in lateral view (scale bar 1.0 mm), (c) larval mandibles (scale bar 50 μm), (d) larval blades (scale bar 50 μm), (e) respiratory organ (scale bar 500 μm). lb = larval blade, ant = antenna, md = mandible, ans = anal fan setae, ro = respiratory organ, tp = terminal process, Lt = lateral teeth, IIV = mandibular teeth, III being the subordinate tooth." figureDoi="http://doi.org/10.5281/zenodo.10084855" httpUri="https://zenodo.org/record/10084855/files/figure.png" pageId="4" pageNumber="61">Fig. 4a</figureCitation>
). Length of antennae 476 (401530, n=19); labral blade transparent and serrated, length 229 (201279), width 48 (3158), length/width 4.8 (3.59.0, n=14) (
<figureCitation id="4EF92A61FFB3FFA828FAF974FBB6D69A" box="[1011,1096,1725,1751]" captionStart="FIGURE 4" captionStartId="5.[152,255,1195,1219]" captionTargetBox="[151,1408,181,1171]" captionTargetId="figure-19@5.[151,1435,181,1171]" captionTargetPageId="5" captionText="FIGURE 4. Chaoborus pseudoflavicans sp. nov., immature stages, (a) larva, habitus in lateral view (scale bar 1.0 mm), (b) pupa, habitus in lateral view (scale bar 1.0 mm), (c) larval mandibles (scale bar 50 μm), (d) larval blades (scale bar 50 μm), (e) respiratory organ (scale bar 500 μm). lb = larval blade, ant = antenna, md = mandible, ans = anal fan setae, ro = respiratory organ, tp = terminal process, Lt = lateral teeth, IIV = mandibular teeth, III being the subordinate tooth." figureDoi="http://doi.org/10.5281/zenodo.10084855" httpUri="https://zenodo.org/record/10084855/files/figure.png" pageId="4" pageNumber="61">Fig. 4d</figureCitation>
); a number of mandibular fan bristles 10.8 (1013, n=19); mandibular teeth, mandibular tooth 1,2 and 4 darkened apically, mandibular tooth 3 dark; lateral teeth conspicuous, first lateral tooth about as long as mandibular tooth 3 (subordinate tooth), average number of lateral teeth 3.4 (34, n=21) (
<figureCitation id="4EF92A61FFB3FFA82953F8E0FD53D70E" box="[602,685,1833,1859]" captionStart="FIGURE 4" captionStartId="5.[152,255,1195,1219]" captionTargetBox="[151,1408,181,1171]" captionTargetId="figure-19@5.[151,1435,181,1171]" captionTargetPageId="5" captionText="FIGURE 4. Chaoborus pseudoflavicans sp. nov., immature stages, (a) larva, habitus in lateral view (scale bar 1.0 mm), (b) pupa, habitus in lateral view (scale bar 1.0 mm), (c) larval mandibles (scale bar 50 μm), (d) larval blades (scale bar 50 μm), (e) respiratory organ (scale bar 500 μm). lb = larval blade, ant = antenna, md = mandible, ans = anal fan setae, ro = respiratory organ, tp = terminal process, Lt = lateral teeth, IIV = mandibular teeth, III being the subordinate tooth." figureDoi="http://doi.org/10.5281/zenodo.10084855" httpUri="https://zenodo.org/record/10084855/files/figure.png" pageId="4" pageNumber="61">Fig. 4c</figureCitation>
); a number of anal fan setae 23 (2126, n=15).
</paragraph>
</subSubSection>
<caption id="82BD666CFFB2FFA92B91FB62FB92D562" ID-DOI="http://doi.org/10.5281/zenodo.10084855" ID-Zenodo-Dep="10084855" httpUri="https://zenodo.org/record/10084855/files/figure.png" pageId="5" pageNumber="62" startId="5.[152,255,1195,1219]" targetBox="[151,1408,181,1171]" targetPageId="5" targetType="figure">
<paragraph id="D67D36E4FFB2FFA92B91FB62FB92D562" blockId="5.[151,1437,1195,1327]" pageId="5" pageNumber="62">
<emphasis id="E4B6EAF6FFB2FFA92B91FB62FEE4D48E" bold="true" box="[152,282,1195,1219]" pageId="5" pageNumber="62">FIGURE 4.</emphasis>
<taxonomicName id="11C24D67FFB2FFA92A2BFB62FDC3D48E" authority="Bang &amp; Shin, 2023" authorityName="Bang &amp; Shin" authorityYear="2023" box="[290,573,1195,1219]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="pseudoflavicans" status="sp. nov.">
<emphasis id="E4B6EAF6FFB2FFA92A2BFB62FDC3D48E" box="[290,573,1195,1219]" italics="true" pageId="5" pageNumber="62">Chaoborus pseudoflavicans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF85578DFFB2FFA9294CFB65FD67D48E" box="[581,665,1196,1220]" pageId="5" pageNumber="62" rank="species">
<emphasis id="E4B6EAF6FFB2FFA9294CFB65FDA3D489" bold="true" box="[581,605,1196,1220]" pageId="5" pageNumber="62">sp</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA92962FB65FD6CD489" bold="true" box="[619,658,1196,1220]" pageId="5" pageNumber="62">nov</emphasis>
.
</taxonomicNameLabel>
, immature stages, (a) larva, habitus in lateral view (scale bar 1.0 mm), (b) pupa, habitus in lateral view (scale bar 1.0 mm), (c) larval mandibles (scale bar 50 μm), (d) larval blades (scale bar 50 μm), (e) respiratory organ (scale bar 500 μm). lb = larval blade, ant = antenna, md = mandible, ans = anal fan setae, ro = respiratory organ, tp = terminal process, Lt = lateral teeth, IIV = mandibular teeth, III being the subordinate tooth.
</paragraph>
</caption>
<subSubSection id="9ED8656FFFB2FFA92BCEFAB2FBA5D5F4" pageId="5" pageNumber="62" type="etymology">
<paragraph id="D67D36E4FFB2FFA92BCEFAB2FBA5D5F4" blockId="5.[151,1437,1403,2041]" pageId="5" pageNumber="62">
<emphasis id="E4B6EAF6FFB2FFA92BCEFAB2FEB0D5D8" bold="true" box="[199,334,1403,1429]" pageId="5" pageNumber="62">Etymology.</emphasis>
The name of this species reflects its taxonomic history, as it was often identified as
<taxonomicName id="11C24D67FFB2FFA92FE0FAB2FA90D5D8" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[1257,1390,1403,1429]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB2FFA92FE0FAB2FB02D5D8" box="[1257,1276,1403,1429]" italics="true" pageId="5" pageNumber="62">C</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA92E00FAB2FA90D5D8" box="[1289,1390,1403,1429]" italics="true" pageId="5" pageNumber="62">flavicans</emphasis>
</taxonomicName>
and remained undescribed for a prolonged period within the
<taxonomicName id="11C24D67FFB2FFA92806FA56FC6AD5F4" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[783,916,1439,1465]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB2FFA92806FA56FCDCD5F4" box="[783,802,1439,1465]" italics="true" pageId="5" pageNumber="62">C</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA92826FA56FC6AD5F4" box="[815,916,1439,1465]" italics="true" pageId="5" pageNumber="62">flavicans</emphasis>
</taxonomicName>
species complex.
</paragraph>
</subSubSection>
<subSubSection id="9ED8656FFFB2FFA92BCEFA0AFA6DD694" pageId="5" pageNumber="62" type="description">
<paragraph id="D67D36E4FFB2FFA92BCEFA0AFA6DD694" blockId="5.[151,1437,1403,2041]" pageId="5" pageNumber="62">
<emphasis id="E4B6EAF6FFB2FFA92BCEFA0AFD64D590" bold="true" box="[199,666,1475,1501]" pageId="5" pageNumber="62">DNA barcodes &amp; neighbor-joining tree.</emphasis>
Partial COI sequences from larvae to adults were obtained. Based on the pairwise distance histogram, the intraspecific variation within the species complex ranged up to 4.9% (with a mode of 1.3%), while the minimum interspecific variation was 11.1% (with a mode of 15.4%). These results suggest that the genetic distances in the
<taxonomicName id="11C24D67FFB2FFA92908F9E6FD79D604" baseAuthorityName="Meigen" baseAuthorityYear="1830" box="[513,647,1583,1609]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="flavicans">
<emphasis id="E4B6EAF6FFB2FFA92908F9E6FDEAD604" box="[513,532,1583,1609]" italics="true" pageId="5" pageNumber="62">C</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA9292BF9E6FD79D604" box="[546,647,1583,1609]" italics="true" pageId="5" pageNumber="62">flavicans</emphasis>
</taxonomicName>
species complex are sufficient to delimitate and identify species (
<figureCitation id="4EF92A61FFB2FFA92E66F9E6FF59D623" captionStart="FIGURE 5" captionStartId="6.[152,255,1299,1323]" captionTargetBox="[159,1428,189,1267]" captionTargetId="figure-27@6.[151,1436,181,1275]" captionTargetPageId="6" captionText="FIGURE 5. The distribution of pairwise genetic distances based on the pairwise distance model using the DNA barcoding regions (COI) of four species in the complex. The intraspecific variation is in black, and the interspecific variation in light blue." figureDoi="http://doi.org/10.5281/zenodo.10084857" httpUri="https://zenodo.org/record/10084857/files/figure.png" pageId="5" pageNumber="62">Fig. 5</figureCitation>
). In addition, it showed that the collected chaoborids are identical to
<taxonomicName id="11C24D67FFB2FFA928AAF99AFB85D620" box="[931,1147,1619,1645]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E4B6EAF6FFB2FFA928AAF99AFBE1D620" box="[931,1055,1619,1645]" italics="true" pageId="5" pageNumber="62">Chaoborus</emphasis>
JPN sp.
</taxonomicName>
sequenced in
<bibRefCitation id="B2534B15FFB2FFA92E1EF99AFF1AD6DC" author="Zhang, X. &amp; Kang, Z. &amp; Ding, S. &amp; Wang, Y. &amp; Borkent, C. &amp; Saigusa, T. &amp; Yang, D." pageId="5" pageNumber="62" pagination="747" refId="ref7482" refString="Zhang, X., Kang, Z., Ding, S., Wang, Y., Borkent, C., Saigusa, T. &amp; Yang, D. (2019) Mitochondrial Genomes Provide Insights into the Phylogeny of Culicomorpha (Insecta: Diptera). International journal of molecular sciences, 20 (3), 747. https: // doi. org / 10.3390 / ijms 20030747" type="journal article" year="2019">
Zhang
<emphasis id="E4B6EAF6FFB2FFA92E6CF99DFA6BD620" box="[1381,1429,1619,1645]" italics="true" pageId="5" pageNumber="62">et al</emphasis>
. (2019)
</bibRefCitation>
, and supported by the NJ tree, which clustered these sequences into a single group (&gt; 99% support), similar to the COI-based tree of
<bibRefCitation id="B2534B15FFB2FFA92AA1F952FD5CD6F8" author="Salmela, J. &amp; Harma, O. &amp; Taylor, D. J." box="[424,674,1691,1717]" pageId="5" pageNumber="62" pagination="151 - 196" refId="ref7342" refString="Salmela, J., Harma, O. &amp; Taylor, D. J. (2021 b) Chaoborus flavicans Meigen (Diptera, Chaoboridae) is a complex of lake and pond dwelling species: a revision. Zootaxa, 4927 (2), 151 - 196. https: // doi. org / 10.11646 / zootaxa. 4927.2.1" type="journal article" year="2021">
Salmela
<emphasis id="E4B6EAF6FFB2FFA92900F955FDC7D6F8" box="[521,569,1691,1717]" italics="true" pageId="5" pageNumber="62">et al</emphasis>
. (2021b)
</bibRefCitation>
. In the tree,
<taxonomicName id="11C24D67FFB2FFA92820F952FC02D6F8" authority="Bang &amp; Shin, 2023" authorityName="Bang &amp; Shin" authorityYear="2023" box="[809,1020,1691,1717]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="pseudoflavicans" status="sp. nov.">
C.
<emphasis id="E4B6EAF6FFB2FFA92841F952FC02D6F8" box="[840,1020,1691,1717]" italics="true" pageId="5" pageNumber="62">pseudoflavicans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF85578DFFB2FFA92F0AF952FBA3D6F8" box="[1027,1117,1691,1717]" pageId="5" pageNumber="62" rank="species">
<emphasis id="E4B6EAF6FFB2FFA92F0AF952FBE3D6F8" bold="true" box="[1027,1053,1691,1717]" pageId="5" pageNumber="62">sp</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA92F22F952FBA9D6F8" bold="true" box="[1067,1111,1691,1717]" pageId="5" pageNumber="62">nov</emphasis>
.
</taxonomicNameLabel>
formed a single cluster (
<figureCitation id="4EF92A61FFB2FFA92E66F952FF59D694" captionStart="FIGURE 6" captionStartId="7.[152,255,1193,1217]" captionTargetBox="[151,1428,181,1162]" captionTargetId="figure-19@7.[151,1436,181,1169]" captionTargetPageId="7" captionText="FIGURE 6. A neighbor-joining tree of the Chaoboridae species based on the pairwise distance model using the mitochondrial DNA barcoding region (COI). In all, 552 COI sequences (C. pseudoflavicans sp. nov., eight specimens sequenced in this study) were used to construct the NJ tree using MEGA 11. The red asterisk indicates that the collected Chaoborids clustered with Chaoborus JPN sp. (MK281356) with&gt; 99% support, and black asterisks indicate the species of the C. flavicans species complex. The light blue shading indicates the recorded species of Chaoboridae in South Korea. Three species of Culicidae are used as outgroups. Triangles represent collapsed clades to save space. F = female, M = male, P = pupa, L = larva." figureDoi="http://doi.org/10.5281/zenodo.10084859" httpUri="https://zenodo.org/record/10084859/files/figure.png" pageId="5" pageNumber="62">Fig. 6</figureCitation>
). All results confirmed that
<taxonomicName id="11C24D67FFB2FFA92AE8F976FD48D694" authority="Bang &amp; Shin, 2023" authorityName="Bang &amp; Shin" authorityYear="2023" box="[481,694,1727,1753]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="67" phylum="Arthropoda" rank="species" species="pseudoflavicans" status="sp. nov.">
C.
<emphasis id="E4B6EAF6FFB2FFA9290BF976FD48D694" box="[514,694,1727,1753]" italics="true" pageId="5" pageNumber="62">pseudoflavicans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF85578DFFB2FFA929B4F976FCE6D694" box="[701,792,1727,1753]" pageId="5" pageNumber="62" rank="species">
<emphasis id="E4B6EAF6FFB2FFA929B4F976FD29D694" bold="true" box="[701,727,1727,1753]" pageId="5" pageNumber="62">sp</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA929ECF976FCEFD694" bold="true" box="[741,785,1727,1753]" pageId="5" pageNumber="62">nov</emphasis>
.
</taxonomicNameLabel>
is a new distinct species that has not yet been identified.
</paragraph>
</subSubSection>
<subSubSection id="9ED8656FFFB2FFA92BCEF92AFD3ED6B0" box="[199,704,1763,1789]" pageId="5" pageNumber="62" type="distribution">
<paragraph id="D67D36E4FFB2FFA92BCEF92AFD3ED6B0" blockId="5.[151,1437,1403,2041]" box="[199,704,1763,1789]" pageId="5" pageNumber="62">
<emphasis id="E4B6EAF6FFB2FFA92BCEF92AFE9FD6B0" bold="true" box="[199,353,1763,1789]" pageId="5" pageNumber="62">Distribution.</emphasis>
<collectingCountry id="AED57674FFB2FFA92A6EF92AFE55D6B0" box="[359,427,1763,1789]" name="South Korea" pageId="5" pageNumber="62">Korea</collectingCountry>
(new record) and
<collectingCountry id="AED57674FFB2FFA92970F92AFD42D6B0" box="[633,700,1763,1789]" name="Japan" pageId="5" pageNumber="62">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9ED8656FFFB2FFA92BCEF8CEFC29D7B4" pageId="5" pageNumber="62" type="discussion">
<paragraph id="D67D36E4FFB2FFA92BCEF8CEFC29D7B4" blockId="5.[151,1437,1403,2041]" pageId="5" pageNumber="62">
<emphasis id="E4B6EAF6FFB2FFA92BCEF8CEFEC5D76C" bold="true" box="[199,315,1799,1825]" pageId="5" pageNumber="62">Remarks.</emphasis>
The new species has been mentioned in some taxonomic works (e.g.
<bibRefCitation id="B2534B15FFB2FFA92F52F8CEFB0DD76F" author="Komyo, E." box="[1115,1267,1799,1826]" pageId="5" pageNumber="62" pagination="11 - 15" refId="ref6831" refString="Komyo, E. (1954) Japanese Chaoborin Mosquitoes. The Scientific Reports of the Saikyo University Agriculture, 6, 11 - 15." type="journal article" year="1954">Komyo 1954</bibRefCitation>
,
<bibRefCitation id="B2534B15FFB2FFA92E09F8CEFF1FD708" author="Salmela, J. &amp; Harma, O. &amp; Taylor, D. J." pageId="5" pageNumber="62" pagination="151 - 196" refId="ref7342" refString="Salmela, J., Harma, O. &amp; Taylor, D. J. (2021 b) Chaoborus flavicans Meigen (Diptera, Chaoboridae) is a complex of lake and pond dwelling species: a revision. Zootaxa, 4927 (2), 151 - 196. https: // doi. org / 10.11646 / zootaxa. 4927.2.1" type="journal article" year="2021">
Salmela
<emphasis id="E4B6EAF6FFB2FFA92E6AF8C1FA6BD76C" box="[1379,1429,1799,1825]" italics="true" pageId="5" pageNumber="62">et al</emphasis>
. 2021b
</bibRefCitation>
), and included in the molecular phylogeny of
<bibRefCitation id="B2534B15FFB2FFA929F2F8E2FC21D708" author="Zhang, X. &amp; Kang, Z. &amp; Ding, S. &amp; Wang, Y. &amp; Borkent, C. &amp; Saigusa, T. &amp; Yang, D." box="[763,991,1835,1861]" pageId="5" pageNumber="62" pagination="747" refId="ref7482" refString="Zhang, X., Kang, Z., Ding, S., Wang, Y., Borkent, C., Saigusa, T. &amp; Yang, D. (2019) Mitochondrial Genomes Provide Insights into the Phylogeny of Culicomorpha (Insecta: Diptera). International journal of molecular sciences, 20 (3), 747. https: // doi. org / 10.3390 / ijms 20030747" type="journal article" year="2019">
Zhang
<emphasis id="E4B6EAF6FFB2FFA92845F8E5FC7ED708" box="[844,896,1835,1861]" italics="true" pageId="5" pageNumber="62">et al</emphasis>
. (2019)
</bibRefCitation>
. While it was previously believed that this taxon was endemic to
<collectingCountry id="AED57674FFB2FFA92AC4F886FDF3D724" box="[461,525,1871,1897]" name="Japan" pageId="5" pageNumber="62">Japan</collectingCountry>
(
<bibRefCitation id="B2534B15FFB2FFA92917F886FCEFD724" author="Salmela, J. &amp; Harma, O. &amp; Taylor, D. J." box="[542,785,1871,1897]" pageId="5" pageNumber="62" pagination="151 - 196" refId="ref7342" refString="Salmela, J., Harma, O. &amp; Taylor, D. J. (2021 b) Chaoborus flavicans Meigen (Diptera, Chaoboridae) is a complex of lake and pond dwelling species: a revision. Zootaxa, 4927 (2), 151 - 196. https: // doi. org / 10.11646 / zootaxa. 4927.2.1" type="journal article" year="2021">
Salmela
<emphasis id="E4B6EAF6FFB2FFA9298DF899FD49D724" box="[644,695,1871,1897]" italics="true" pageId="5" pageNumber="62">et al</emphasis>
. 2021b
</bibRefCitation>
), the morphological and molecular analyses showed that this species also occurs in the Korean Peninsula. In conclusion, the paramere of
<taxonomicName id="11C24D67FFB2FFA92F24F8BAFAFDD7C0" authority="Bang &amp; Shin, 2023" authorityName="Bang &amp; Shin" authorityYear="2023" box="[1069,1283,1907,1933]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="pseudoflavicans" status="sp. nov.">
<emphasis id="E4B6EAF6FFB2FFA92F24F8BAFBBED7C0" box="[1069,1088,1907,1933]" italics="true" pageId="5" pageNumber="62">C</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA92F46F8BAFAFDD7C0" box="[1103,1283,1907,1933]" italics="true" pageId="5" pageNumber="62">pseudoflavicans</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FF85578DFFB2FFA92E05F8BAFA96D7C0" box="[1292,1384,1907,1933]" pageId="5" pageNumber="62" rank="species">
<emphasis id="E4B6EAF6FFB2FFA92E05F8BAFAD8D7C0" bold="true" box="[1292,1318,1907,1933]" pageId="5" pageNumber="62">sp</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA92E3FF8BAFA9CD7C0" bold="true" box="[1334,1378,1907,1933]" pageId="5" pageNumber="62">nov</emphasis>
.
</taxonomicNameLabel>
was practically identical to that of
<taxonomicName id="11C24D67FFB2FFA92AE9F85EFDAFD7FC" baseAuthorityName="Johannsen" baseAuthorityYear="1903" box="[480,593,1943,1969]" class="Insecta" family="Chaoboridae" genus="Chaoborus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="62" phylum="Arthropoda" rank="species" species="albipes">
<emphasis id="E4B6EAF6FFB2FFA92AE9F85EFE0DD7FC" box="[480,499,1943,1969]" italics="true" pageId="5" pageNumber="62">C</emphasis>
.
<emphasis id="E4B6EAF6FFB2FFA92909F85EFDAFD7FC" box="[512,593,1943,1969]" italics="true" pageId="5" pageNumber="62">albipes</emphasis>
</taxonomicName>
, except for slightly greater curvature in the median position of the paramere. Additionally, the presence or absence of the tergal band in adults, the differences in mandibular teeth of larvae, and the DNA barcode confirmed it is a distinct species in the species complex.
</paragraph>
</subSubSection>
<caption id="82BD666CFFB1FFAA2B91FADAFF34D53E" ID-DOI="http://doi.org/10.5281/zenodo.10084857" ID-Zenodo-Dep="10084857" httpUri="https://zenodo.org/record/10084857/files/figure.png" pageId="6" pageNumber="63" startId="6.[152,255,1299,1323]" targetBox="[159,1428,189,1267]" targetPageId="6" targetType="figure">
<paragraph id="D67D36E4FFB1FFAA2B91FADAFF34D53E" blockId="6.[151,1436,1299,1395]" pageId="6" pageNumber="63">
<emphasis id="E4B6EAF6FFB1FFAA2B91FADAFEE4D566" bold="true" box="[152,282,1299,1323]" pageId="6" pageNumber="63">FIGURE 5.</emphasis>
The distribution of pairwise genetic distances based on the pairwise distance model using the DNA barcoding regions (COI) of four species in the complex. The intraspecific variation is in black, and the interspecific variation in light blue.
</paragraph>
</caption>
<subSubSection id="9ED8656FFFB1FFAA2BCEFA0CFB13D66A" pageId="6" pageNumber="63" type="description">
<paragraph id="D67D36E4FFB1FFAA2BCEFA0CFB13D66A" blockId="6.[151,1436,1477,1576]" pageId="6" pageNumber="63">
<emphasis id="E4B6EAF6FFB1FFAA2BCEFA0CFEB5D592" bold="true" box="[199,331,1477,1503]" pageId="6" pageNumber="63">Bionomics.</emphasis>
The immature specimens of this new species were collected from a small, lightly iced, fishless pond (10 ×
<quantity id="113A9B01FFB1FFAA2BD5FA20FEE8D64E" box="[220,278,1513,1539]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="63" unit="m" value="10.0">10 m</quantity>
) in
<date id="A27C1024FFB1FFAA2A4DFA20FD93D64E" box="[324,621,1513,1539]" pageId="6" pageNumber="63" value="2023-02" valueMax="2023-03-31" valueMin="2023-02-00">February and March 2023</date>
. The water was eutrophic, and colored light brown with
<taxonomicName id="11C24D67FFB1FFAA2FE5FA20FA96D649" box="[1260,1384,1513,1540]" class="Liliopsida" family="Typhaceae" genus="Typha" kingdom="Plantae" order="Poales" pageId="6" pageNumber="63" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="E4B6EAF6FFB1FFAA2FE5FA20FACED64E" box="[1260,1328,1513,1539]" italics="true" pageId="6" pageNumber="63">Typha</emphasis>
spp.
</taxonomicName>
, and fallen leaves. Chironomid larvae and
<taxonomicName id="11C24D67FFB1FFAA2932F9C7FD1DD665" box="[571,739,1550,1576]" class="Insecta" family="Notonectidae" genus="Notonecta" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="63" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E4B6EAF6FFB1FFAA2932F9C7FD50D66A" box="[571,686,1550,1575]" italics="true" pageId="6" pageNumber="63">Notonecta</emphasis>
spp.
</taxonomicName>
were also collected with the chaoborid larvae.
</paragraph>
</subSubSection>
</treatment>
</document>