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<document id="DC844EDBE51A3482EF530881BFE992F7" ID-CLB-Dataset="22147" ID-DOI="10.1093/isd/ixaa006" ID-GBIF-Dataset="6c2ad151-a6e0-4c1b-be84-d1950abe22dd" ID-ISSN="2399-3421" ID-Zenodo-Dep="3826921" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589464781686" checkinUser="felipe" docAuthor="Smith-Pardo, Allan H., Carpenter, James M. &amp; Kimsey, Lynn" docDate="2020" docId="514381547E4EFF89FF1D11773B58F9C8" docLanguage="en" docName="InsectSystDivers.4.3.2.1-27.pdf.imf" docOrigin="Insect Systematics and Diversity 4 (3), No. 2" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Vespa multimaculata Perez 1910" docType="treatment" docVersion="7" lastPageNumber="17" masterDocId="AD7AF92C7E5EFF99FF8F16743F73FFC9" masterDocTitle="The Diversity of Hornets in the Genus Vespa (Hymenoptera: Vespidae; Vespinae), Their Importance and Interceptions in the United States" masterLastPageNumber="27" masterPageNumber="1" pageNumber="17" updateTime="1698839653073" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="6B1CD140AD4AFF7B6F81B0D362E19D31" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="86CAC4795253B7669F73C227E26755B1">
<mods:title id="C492A1E7E8A8EE5BE56F3B3C5FEE773E">The Diversity of Hornets in the Genus Vespa (Hymenoptera: Vespidae; Vespinae), Their Importance and Interceptions in the United States</mods:title>
</mods:titleInfo>
<mods:name id="F7816F286465202B0C986B22340AA1E4" type="personal">
<mods:role id="82C27ED47E0D2B857982796E4B61E4BC">
<mods:roleTerm id="098F24B2928AA70DFED217325F31D26D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C0CB1D17D6DCF36BB98203B43E72B193">Smith-Pardo, Allan H.</mods:namePart>
<mods:affiliation id="8777CF991B172DD51173BC1233D380A1">USDA-APHIS-PPQ, Science and Technology (S &amp; T), Sacramento, CA,</mods:affiliation>
<mods:nameIdentifier id="B443C55F63A77092184BA13B142632EF" type="email">allan.h.smith-pardo@usda.gov</mods:nameIdentifier>
</mods:name>
<mods:name id="37E7A39AD2FC217B05DB1C03A808C30B" type="personal">
<mods:role id="4D13BB07B90968847E85CB30F2C87475">
<mods:roleTerm id="701AC57D21424AB0A55B93C64FCEE55C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C1A7BAE5E2FE502CB8F47FEC7F2A5F1D">Carpenter, James M.</mods:namePart>
<mods:affiliation id="4919BF250B740E461F2CB53180F22A72">Department of Invertebrate Zoology, American Museum of Natural History, New York, NY,</mods:affiliation>
</mods:name>
<mods:name id="C46E0ADA73A8689251516F1C8E8F8AB2" type="personal">
<mods:role id="D4BE58E47806DE7D0F72D734983E6988">
<mods:roleTerm id="5FB6D32B128AC4FC9EC4C7F56C49AAF4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EF9DB1150BA2DE3E9C65046D14CD978E">Kimsey, Lynn</mods:namePart>
<mods:affiliation id="B4D0BA9B91375B741AE93263080EC95A">Bohart Museum of Entomology, University of California, Davis, Davis, CA, and</mods:affiliation>
</mods:name>
<mods:typeOfResource id="8CB4AAF6467BF7295678F1427EA18D9C">text</mods:typeOfResource>
<mods:relatedItem id="1B94FE49CA7980AE486ABC43A204BC81" type="host">
<mods:titleInfo id="97F30DCCC682E397C8A5E2734D5E9D92">
<mods:title id="5F327E53D0E37058B833C39EC9798E39">Insect Systematics and Diversity</mods:title>
</mods:titleInfo>
<mods:part id="CCA0ED43D39BED74E5C7AD3D4442C5D7">
<mods:date id="1558783C31E1C2CCFA3326EAC353C134">2020</mods:date>
<mods:detail id="1C7D341B9AC46307C00339FBF854953C" type="pubDate">
<mods:number id="4CAFE8165233F12186B6444AAD3BA75F">2020-05-31</mods:number>
</mods:detail>
<mods:detail id="84D76A12BD2B02933750324875BC1003" type="volume">
<mods:number id="0FA7C97528E548A7ED4C90DAF26B39DB">4</mods:number>
</mods:detail>
<mods:detail id="6E590E17183DDDE88C08C3AD395A2711" type="issue">
<mods:number id="789DB11ADDB1EEC4D64ED0F6E743239C">3</mods:number>
</mods:detail>
<mods:detail id="F4BA1DBBEBA676CE02939FCCF3889235" type="numero">
<mods:number id="544E78710FDFD634A3F23A8FA086EFAB">2</mods:number>
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<mods:start id="8FA7F3C5CA92C6B0BFEE0A785B4E8C0A">1</mods:start>
<mods:end id="F30960F0C32799277097CAEE0786C291">27</mods:end>
</mods:extent>
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<mods:classification id="EDDDEC456B8CD7F3E2B655C40AB39240">journal article</mods:classification>
<mods:identifier id="584EF9A94F3C4669F93A6488814C06DD" type="CLB-Dataset">22147</mods:identifier>
<mods:identifier id="C3E7D6C00166012BE150201DE02B0687" type="DOI">10.1093/isd/ixaa006</mods:identifier>
<mods:identifier id="6B45241611524FF8D4E69D0007879808" type="GBIF-Dataset">6c2ad151-a6e0-4c1b-be84-d1950abe22dd</mods:identifier>
<mods:identifier id="B63F0033E6379DCBCEB10026E3997E9D" type="ISSN">2399-3421</mods:identifier>
<mods:identifier id="99FECD1AE4C51F2816FBDC30B5BD0F2B" type="Zenodo-Dep">3826921</mods:identifier>
</mods:mods>
<treatment id="514381547E4EFF89FF1D11773B58F9C8" ID-DOI="http://doi.org/10.5281/zenodo.3847042" ID-GBIF-Taxon="164133884" ID-Zenodo-Dep="3847042" LSID="urn:lsid:plazi:treatment:514381547E4EFF89FF1D11773B58F9C8" httpUri="http://treatment.plazi.org/id/514381547E4EFF89FF1D11773B58F9C8" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="91F063C97E4EFF89FF1D11773EB1F8D3" box="[146,450,1795,1818]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="D95530427E4EFF89FF1D11773EB1F8D3" blockId="16.[146,450,1795,1849]" box="[146,450,1795,1818]" pageId="16" pageNumber="17">
<heading id="821D872E7E4EFF89FF1D11773EB1F8D3" box="[146,450,1795,1818]" fontSize="9" level="2" pageId="16" pageNumber="17" reason="4">
<taxonomicName id="1EEA4BC17E4EFF89FF1D11773EB1F8D3" ID-CoL="5B9XH" authority="Perez" authorityName="Perez" authorityYear="1910" box="[146,450,1795,1818]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="multimaculata">
<emphasis id="EB9EEC507E4EFF89FF1D11773E0CF8D3" box="[146,383,1795,1818]" italics="true" pageId="16" pageNumber="17">Vespa multimaculata</emphasis>
Pérez
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="91F063C97E4EFF89FF1D11523ED3F8F0" box="[146,416,1830,1849]" pageId="16" pageNumber="17" type="description">
<paragraph id="D95530427E4EFF89FF1D11523ED3F8F0" blockId="16.[146,450,1795,1849]" box="[146,416,1830,1849]" pageId="16" pageNumber="17">
<figureCitation id="41D12CC77E4EFF89FF1D11523FA9F8F0" box="[146,218,1830,1849]" captionStart="Figs" captionStartId="2.[146,181,1617,1634]" captionTargetBox="[153,1435,148,1581]" captionTargetId="figure@2.[152,1435,147,1582]" captionTargetPageId="2" captionText="Figs. 1–4. Vespa morphology. Fig. 1 (a) Hindwing. (b) Forewing apex.(c) Lateral view of pronotal lobe.(e) Hindtarsal claws.(f)Thoracic dorsum. (g) Lateral view of metasomal segment 1. Fig. 2. (a) Lateral view of pronotal lobe. (b) Dorsal view of vertex, i =distance from hind ocellus to posterior head margin, ii=distance between lateral ocellus and eye. (c) Lateral view of metasomal segment 1. (d) Costal margin of forewing apex.(e) Dorsal view of vertex.(c) Front view of clypeus and mandibles. Fig. 3.(a, b) Lateral view of head. (c, d) Lateral view of metasoma.(3a, 3c)V.soror. Fig. 4. (a, b) Lateral view of metasomal tergum 2. (c, d) Lateral view of pronotal carina. (e, f) Front view of clypeus. Figs. 1a-g, 2a, 2b, 3b, 3d, V. crabro. Fig. 2c, V. binghami. Figs. 2d, 2e, V. analis. Fig. 2f, V. bicolor. Figs. 4a, 4e, V. fervida. Fig. 4b, V. multimaculata. Figs. 4c, 4f, V. luctuosa. Fig. 4d, V. affinis." figureDoi="http://doi.org/10.5281/zenodo.3826923" httpUri="https://zenodo.org/record/3826923/files/figure.png" pageId="16" pageNumber="17">Figs. 4b</figureCitation>
,
<figureCitation id="41D12CC77E4EFF89FF6A11523E42F8F0" box="[229,305,1830,1849]" captionStart="Figs" captionStartId="5.[146,181,1785,1802]" captionTargetBox="[187,1402,148,1736]" captionTargetId="figure@5.[186,1402,147,1750]" captionTargetPageId="5" captionText="Figs. 9–12. Vespa morphology. (a, b) Dorsal view of metasoma. (c, d) Dorsal view of basal metasomal terga. Fig. 10. (a, b) Dorsal view of scutum. (c, d) Dorsal view of metasoma. Fig. 11. (a, b) Lateral view of head. (c, d) Dorsal view of metasoma. Fig. 12. (a, b) Front view of clypeus.(c-f) Dorsal view of metanotum. Figs. 9a, 9b,V. orientalis. Figs. 9c, 10b, d, V. affinis. Fig. 9d, V. crabro. Figs. 10a, 10c, 11b, 11c, V. mocsaryana. Fig. 11a, V. fumida. Fig. 11d. V. dybowskii. Figs. 12a, 12f, V. multimaculata. Figs. 12b, d, e, V. bicolor. Fig. 12c, V. luctuosa." figureDoi="http://doi.org/10.5281/zenodo.3826927" httpUri="https://zenodo.org/record/3826927/files/figure.png" pageId="16" pageNumber="17">12a, 12f</figureCitation>
,
<figureCitation id="41D12CC77E4EFF89FEB311523E27F8F0" box="[316,340,1830,1849]" captionStart="Figs" captionStartId="18.[146,181,1354,1371]" captionTargetBox="[147,1429,148,1341]" captionTargetId="figure@18.[146,1429,147,1342]" captionTargetPageId="18" captionText="Figs. 38 and 39. Vespa multimaculata color forms. (a) Lateral view. (b) Dorsal view. (c) Front view of face." figureDoi="http://doi.org/10.5281/zenodo.3826957" httpUri="https://zenodo.org/record/3826957/files/figure.png" pageId="16" pageNumber="17">38</figureCitation>
, and
<figureCitation id="41D12CC77E4EFF89FE0711523ED3F8F0" box="[392,416,1830,1849]" captionStart="Figs" captionStartId="18.[146,181,1354,1371]" captionTargetBox="[147,1429,148,1341]" captionTargetId="figure@18.[146,1429,147,1342]" captionTargetPageId="18" captionText="Figs. 38 and 39. Vespa multimaculata color forms. (a) Lateral view. (b) Dorsal view. (c) Front view of face." figureDoi="http://doi.org/10.5281/zenodo.3826957" httpUri="https://zenodo.org/record/3826957/files/figure.png" pageId="16" pageNumber="17">39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="91F063C97E4EFF89FF1D11223A79FB3C" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="D95530427E4EFF89FF1D11223D20F86D" blockId="16.[146,769,1878,1956]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="F9FA176C7E4EFF89FF1D11223D20F86D" pageId="16" pageNumber="17">
<treatmentCitation id="584B16537E4EFF89FF1D11223EAEF8A0" box="[146,477,1878,1897]" pageId="16" pageNumber="17">
<taxonomicName id="1EEA4BC17E4EFF89FF1D11223EAEF8A0" ID-CoL="5B9QV" authority="Smith, 1858: 116" authorityName="Smith" authorityPageNumber="116" authorityYear="1858" box="[146,477,1878,1897]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="EB9EEC507E4EFF89FF1D11223E56F8A0" box="[146,293,1878,1897]" italics="true" pageId="16" pageNumber="17">Vespa annulata</emphasis>
Smith, 1858: 116
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="06518EE07E4EFF89FE6311223D35F8A0" box="[492,582,1878,1897]" pageId="16" pageNumber="17" type="lectotype">Lectotype</typeStatus>
female (designated by Carpenter and Kojima 1997);
<collectingCountry id="A1FD70D27E4EFF89FE7611073D23F84F" box="[505,592,1907,1926]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
:
<location id="DC3566997E4EFF89FDD111003DD7F84E" LSID="urn:lsid:plazi:treatment:514381547E4EFF89FF1D11773B58F9C8:DC3566997E4EFF89FDD111003DD7F84E" box="[606,676,1908,1927]" country="Malaysia" name="Borneo" pageId="16" pageNumber="17">Borneo</location>
: Sarawak (OXFORD). Nec
<taxonomicName id="1EEA4BC17E4EFF89FEC211E53D3DF86D" ID-CoL="5B9QV" authority="Rossi, 1790" authorityName="Rossi" authorityYear="1790" box="[333,590,1936,1956]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="EB9EEC507E4EFF89FEC211E53EA9F86A" box="[333,474,1936,1955]" italics="true" pageId="16" pageNumber="17">Vespa annulata</emphasis>
Rossi, 1790
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D95530427E4EFF89FCBD123A3BCCFBB6" blockId="16.[818,1442,1102,1269]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="F9FA176C7E4EFF89FCBD123A3BCCFBB6" pageId="16" pageNumber="17">
<treatmentCitation id="584B16537E4EFF89FCBD123A3BFFFBA8" box="[818,1164,1102,1121]" pageId="16" pageNumber="17">
<taxonomicName id="1EEA4BC17E4EFF89FCBD123A3BFFFBA8" ID-CoL="5B9XH" authority="Perez, 1910: 14" authorityName="Perez" authorityPageNumber="14" authorityYear="1910" box="[818,1164,1102,1121]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="multimaculata">
<emphasis id="EB9EEC507E4EFF89FCBD123A3C80FBA8" box="[818,1011,1102,1121]" italics="true" pageId="16" pageNumber="17">Vespa multimaculata</emphasis>
Pérez, 1910: 14
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="06518EE07E4EFF89FB19123A3B83FBA8" box="[1174,1264,1102,1121]" pageId="16" pageNumber="17" type="lectotype">Lectotype</typeStatus>
female (designated by
<collectorName id="741F55947E4EFF89FCE912183C97FBB7" box="[870,996,1131,1151]" pageId="16" pageNumber="17">van der Vecht</collectorName>
1957);
<collectingCountry id="A1FD70D27E4EFF89FBA3121F3B1BFBB7" box="[1068,1128,1131,1150]" name="Brunei" pageId="16" pageNumber="17">Brunei</collectingCountry>
(PARIS).
</treatmentCitationGroup>
</paragraph>
<paragraph id="D95530427E4EFF89FCBD12FE3B49FB73" blockId="16.[818,1442,1102,1269]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="F9FA176C7E4EFF89FCBD12FE3B49FB73" pageId="16" pageNumber="17">
<treatmentCitation id="584B16537E4EFF89FCBD12FE3B99FB55" box="[818,1258,1161,1180]" pageId="16" pageNumber="17">
<taxonomicName id="1EEA4BC17E4EFF89FCBD12FE3B99FB55" ID-CoL="5B9WS" authority="Bequaert, 1934: 4" authorityName="Bequaert" authorityPageNumber="4" authorityYear="1934" box="[818,1258,1161,1180]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="variety" species="luctuosa" variety="malayana">
<emphasis id="EB9EEC507E4EFF89FCBD12FE3CC8FB55" box="[818,955,1161,1180]" italics="true" pageId="16" pageNumber="17">Vespa luctuosa</emphasis>
var.
<emphasis id="EB9EEC507E4EFF89FC6812FD3B4CFB55" box="[999,1087,1161,1180]" italics="true" pageId="16" pageNumber="17">malayana</emphasis>
Bequaert, 1934: 4
</taxonomicName>
</treatmentCitation>
. Replacement name for
<taxonomicName id="1EEA4BC17E4EFF89FCE412D33B49FB73" ID-CoL="5B9QV" authority="Smith." authorityName="Smith." box="[875,1082,1191,1210]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="annulata">
<emphasis id="EB9EEC507E4EFF89FCE412D33C8BFB73" box="[875,1016,1191,1210]" italics="true" pageId="16" pageNumber="17">Vespa annulata</emphasis>
Smith.
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="D95530427E4EFF89FCBD12B13A79FB3C" blockId="16.[818,1442,1102,1269]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="F9FA176C7E4EFF89FCBD12B13A79FB3C" pageId="16" pageNumber="17">
<treatmentCitation id="584B16537E4EFF89FCBD12B13A30FB1E" box="[818,1347,1220,1240]" pageId="16" pageNumber="17">
<taxonomicName id="1EEA4BC17E4EFF89FCBD12B13A30FB1E" ID-CoL="5LR6P" authority="van der Vecht, 1957: 43" authorityName="van der Vecht" authorityPageNumber="43" authorityYear="1957" box="[818,1347,1220,1240]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="multimaculata" subSpecies="pendleburyi">
<emphasis id="EB9EEC507E4EFF89FCBD12B13B10FB1E" box="[818,1123,1220,1239]" italics="true" pageId="16" pageNumber="17">Vespa multimaculata pendleburyi</emphasis>
van der Vecht, 1957: 43
</taxonomicName>
</treatmentCitation>
;
<typeStatus id="06518EE07E4EFF89FAC412B03AD1FB1E" box="[1355,1442,1220,1239]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
female;
<collectingCountry id="A1FD70D27E4EFF89FC1D12963C9AFB3C" box="[914,1001,1250,1269]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
: ‘near
<location id="DC3566997E4EFF89FBAA12963B3CFB3C" LSID="urn:lsid:plazi:treatment:514381547E4EFF89FF1D11773B58F9C8:DC3566997E4EFF89FBAA12963B3CFB3C" box="[1061,1103,1250,1269]" country="Malaysia" municipality="Kedah'" name="Jitra" pageId="16" pageNumber="17">Jitra</location>
,
<collectingMunicipality id="3931AA387E4EFF89FBD712963BE8FB3C" box="[1112,1179,1250,1269]" pageId="16" pageNumber="17">Kedah’</collectingMunicipality>
, (LEIDEN).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="91F063C97E4EFF89FCBD13693B54FAA2" pageId="16" pageNumber="17" type="distribution">
<paragraph id="D95530427E4EFF89FCBD13693B54FAA2" blockId="16.[818,1442,1309,1387]" pageId="16" pageNumber="17">
<emphasis id="EB9EEC507E4EFF89FCBD13693CD0FAF9" bold="true" box="[818,931,1309,1328]" pageId="16" pageNumber="17">Distribution</emphasis>
.
<collectingCountry id="A1FD70D27E4EFF89FC3613693B7CFAF9" box="[953,1039,1309,1328]" name="Thailand" pageId="16" pageNumber="17">Thailand</collectingCountry>
;
<collectingCountry id="A1FD70D27E4EFF89FBAA13693B20FAF9" box="[1061,1107,1309,1328]" name="Laos" pageId="16" pageNumber="17">Laos</collectingCountry>
;
<collectingCountry id="A1FD70D27E4EFF89FBE613693BB3FAF9" box="[1129,1216,1309,1328]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
: Peninsular
<collectingCountry id="A1FD70D27E4EFF89FAC713693AEDFAF9" box="[1352,1438,1309,1328]" name="Malaysia" pageId="16" pageNumber="17">Malaysia</collectingCountry>
,
<collectingCountry id="A1FD70D27E4EFF89FCBD134E3CF0FA84" box="[818,899,1338,1357]" name="Malaysia" pageId="16" pageNumber="17">Sarawak</collectingCountry>
,
<collectingRegion id="1B2EFEA07E4EFF89FC2D134E3CAFFA84" box="[930,988,1338,1357]" country="Malaysia" name="Sabah" pageId="16" pageNumber="17">Sabah</collectingRegion>
;
<collectingCountry id="A1FD70D27E4EFF89FC74134E3B2AFA84" box="[1019,1113,1338,1357]" name="Singapore" pageId="16" pageNumber="17">Singapore</collectingCountry>
;
<collectingCountry id="A1FD70D27E4EFF89FBF8134E3BA0FA84" box="[1143,1235,1338,1357]" name="Indonesia" pageId="16" pageNumber="17">Indonesia</collectingCountry>
:
<collectingRegion id="1B2EFEA07E4EFF89FB7D134E3A32FA84" box="[1266,1345,1338,1357]" country="Indonesia" name="Sumatera Utara" pageId="16" pageNumber="17">Sumatra</collectingRegion>
, Borneo (
<collectingRegion id="1B2EFEA07E4EFF89FCB7132C3CA7FAA2" box="[824,980,1368,1387]" country="Indonesia" name="Kalimantan Utara" pageId="16" pageNumber="17">KALIMANTAN</collectingRegion>
);
<collectingCountry id="A1FD70D27E4EFF89FC6C132C3B50FAA2" box="[995,1059,1368,1387]" name="Brunei" pageId="16" pageNumber="17">Brunei</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="91F063C97E4EFF89FCBD13E13B58F9C8" pageId="16" pageNumber="17" type="discussion">
<paragraph id="D95530427E4EFF89FCBD13E13B58F9C8" blockId="16.[818,1442,1429,1537]" pageId="16" pageNumber="17">
<emphasis id="EB9EEC507E4EFF89FCBD13E13CE7FA61" bold="true" box="[818,916,1429,1448]" pageId="16" pageNumber="17">Discussion</emphasis>
.
<taxonomicName id="1EEA4BC17E4EFF89FC2F13E23B10FA61" authorityName="Perez" authorityYear="1910" box="[928,1123,1429,1448]" class="Insecta" family="Vespidae" genus="Vespa" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="multimaculata">
<emphasis id="EB9EEC507E4EFF89FC2F13E23B10FA61" box="[928,1123,1429,1448]" italics="true" pageId="16" pageNumber="17">Vespa multimaculata</emphasis>
</taxonomicName>
is found in lowland and montane forests in Southeast Asia (
<bibRefCitation id="BD7B4DB37E4EFF89FB9113C73BE2FA0F" author="Archer" box="[1054,1169,1459,1478]" firstAuthor="Archer" pageId="16" pageNumber="17" pagination="43 - 50" refId="ref11948" refString="Archer, M. E. 1999. Taxonomy, distribution and nesting biology of Vespa binghami, V. basalis, V. variabilis, V. fervida, V. luctuosa, V. multimaculata and V. bellicosa. Entomol. Monthly Mag. 135: 43 - 50." type="journal article" year="1999">Archer 1999</bibRefCitation>
,
<bibRefCitation id="BD7B4DB37E4EFF89FB1313C73BB8FA0F" author="Archer" box="[1180,1227,1459,1478]" firstAuthor="Archer" pageId="16" pageNumber="17" pagination="561 - 567" refId="ref12041" refString="Archer, M. E. 2011. Analysis of a nest of Vespa multimaculata Perez, 1910. Malay. Nat. J. 63: 561 - 567." type="journal article" year="2011">2011</bibRefCitation>
), and is unlikely to disperse to temperate regions.
<bibRefCitation id="BD7B4DB37E4EFF89FBA213A43BC1FA2A" author="Martin" box="[1069,1202,1488,1507]" firstAuthor="Martin" pageId="16" pageNumber="17" pagination="71 - 78" refId="ref13041" refString="Martin, S. J. 1995. Hornets (Hymenoptera: Vespidae) of Malaysia. Malay. Nat. J. 49: 71 - 78." type="journal article" year="1995">Martin (1995)</bibRefCitation>
found nests in the ground beneath the roots of a tree.
</paragraph>
</subSubSection>
</treatment>
</document>
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