94 lines
14 KiB
XML
94 lines
14 KiB
XML
<document id="091F1F9AE1124333C1816F9D75E7401C" ENCODING="UTF-8" ID-GBIF-Dataset="d5fd4c73-69cc-419e-ae2e-811f29efd1bc" ModsDocID="z00791p001" checkinTime="1246965505869" checkinUser="thomas" docAuthor="Victor G. Springer & Gerald R. Allen" docDate="2004" docId="4CCD2173165783E13C238D3E3F8978DE" docLanguage="en" docName="2004_Springer_Allen_gg1.xml" docOrigin="Zootaxa 791" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:87531ECD-559C-4BFB-98B8-29828E58A3A6" docTitle="Ecsenius" docType="treatment" docVersion="13" lastPageNumber="2" masterDocId="EF1D1312A59C6883FD90C25C51F795A2" masterDocTitle="Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius." masterLastPageNumber="12" masterPageNumber="1" pageNumber="1" updateTime="1701314078103" updateUser="plazi">
|
|
<mods:mods id="AAB91A0511BBA774E6148A68F4A77FB7" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="2513E13967341049C0FC8BCE8DBA8CC9">
|
|
<mods:title id="461D369FE230626E8482DC4D045D8E4D">Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius.</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="FBFC959E1A892C61FB16F9406A0E468E" type="personal">
|
|
<mods:role id="DBC82D6CADA8E1A14651DC5631C6420C">
|
|
<mods:roleTerm id="66288E2EFB375BF9273E3B8B6CC9A749">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="46B0688EA34901E5AA00A745566B9E3E">Victor G. Springer</mods:namePart>
|
|
</mods:name>
|
|
<mods:name id="B73ED5FB9F8CFA6C7F23D77CDE7AA364" type="personal">
|
|
<mods:role id="64CB5F29BF3AC80F852EEC28741EDF9D">
|
|
<mods:roleTerm id="AF4E5D9BC3D37DA005480042ED5CCB62">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="E2C264375AE38120EDFA32240999F905">Gerald R. Allen</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource id="1363123BB74B79FEE76A2F12B7C1DEF5">text</mods:typeOfResource>
|
|
<mods:relatedItem id="9BA913129ADD994D749F6331FAF5E03B" type="host">
|
|
<mods:titleInfo id="D6A514DE3744E3CD759B2A4002DE9EB5">
|
|
<mods:title id="95BB2EB16B7C86E2465FC0CCCF0BD7C3">Zootaxa</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="FCEC86D4895D8AD3672F40C67EFBA95C">
|
|
<mods:date id="E5160894FD0065EC12A6A79C42B422D0">2004</mods:date>
|
|
<mods:detail id="9FEBA065E5EC0D536664BF4BBA0A71A3" type="volume">
|
|
<mods:number id="FD84C46CF07021C0CB7AC70C72F6D047">791</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="BFD006AE8383BF581BCFE38C21A199AC" unit="page">
|
|
<mods:start id="3C3F0D75A61049579A29B1D51F0989D5">1</mods:start>
|
|
<mods:end id="C86E95B6D49B435C3292D34DB5544102">12</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location id="B2BB127D5E82EB76A141EC85C3489415">
|
|
<mods:url id="F3EE13AFC868D13E0E4F5BAE4A76122F">http://www.zoobank.org/urn:lsid:zoobank.org:pub:87531ECD-559C-4BFB-98B8-29828E58A3A6</mods:url>
|
|
</mods:location>
|
|
<mods:classification id="7B7CDDA71F8317B264C5DEF08E076CD1">journal article</mods:classification>
|
|
<mods:identifier id="F3FD82781FCA688B2BE57DFE82441CDF" type="Plazi-Custom">z00791p001</mods:identifier>
|
|
<mods:identifier id="21E25CE28D57E145F1AB4D9B6F45B455" type="ZooBank">87531ECD-559C-4BFB-98B8-29828E58A3A6</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="4CCD2173165783E13C238D3E3F8978DE" ID-DOI="http://doi.org/10.5281/zenodo.6271242" ID-GBIF-Taxon="100122813" ID-Zenodo-Dep="6271242" LSID="urn:lsid:plazi:treatment:4CCD2173165783E13C238D3E3F8978DE" httpUri="http://treatment.plazi.org/id/4CCD2173165783E13C238D3E3F8978DE" lastPageNumber="2" pageNumber="1">
|
|
<subSubSection id="391FD9FCFAD390C23F9C074D7DDFDA8F" type="nomenclature">
|
|
<paragraph id="00F9EFD7D00DAE38CFEC2879A22D118A" pageNumber="1">
|
|
<taxonomicName id="6075484746C020962EA5A51ADE1346F8" ID-CoL="632JT" ID-ENA="57791" LSID-ZBK="urn:lsid:zoobank.org:act:B3F80107-BABB-495D-BFB7-721DCA10C4F8" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius McCulloch 1923:121" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">[[ Ecsenius ]]</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4A198D9928CAA89D57556F097A8C91D5" type="discussion">
|
|
<paragraph id="D3702252F5F027D809B1D0F491E9DC11" pageNumber="1">
|
|
<taxonomicName id="B6127FF0684455CA02C153CE73F69F11" LSID-ZBK="urn:lsid:zoobank.org:act:B3F80107-BABB-495D-BFB7-721DCA10C4F8" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius McCulloch 1923:121" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Ecsenius</taxonomicName>
|
|
now comprises 53 species, more than twice as many as the next speciose blenniid genus,
|
|
<taxonomicName id="6A250E48355062F609F41B51A95BF477" LSID-ZBK="urn:lsid:zoobank.org:act:F0C4AFAF-0033-43C6-9965-4D47EA2DBF13" family="Blenniidae" genus="Meiacanthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Meiacanthus Norman 1944:805" order="Perciformes" pageNumber="1" phylum="Chordata" rank="genus">Meiacanthus Norman</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="1F54C06CA0ECA6B5B731A695CB31B84B" lastPageNumber="2" pageNumber="1">
|
|
The
|
|
<taxonomicName id="192E042FB6CDB4E516D721B8A3C3B96D" LSID-ZBK="urn:lsid:zoobank.org:act:669CE437-D461-4DF0-AF57-F94B3740CCAE" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius prooculis Chapman & Schultz 1952:519" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
|
|
species group is defined by a combination of characters, only one of which appears to be synapomorphic: presence on ventral surface of head of two bilateral pairs of small dark spots (Figure 1a, b; also Springer, 1988:figure 57) or single bilateral pair of elongate dark crescentic marks (Springer, 1988:figure 58) in at least some males, or two pairs of pale, round spots (Figure 2c; also Springer, 1988:figure 56; 1991:figure 8) or single pair of pale crescentic marks in some females and dark crescentic marks in some males. The central portions of the crescentic dark marks are absent in some specimens of one species (
|
|
<taxonomicName id="340AD1353FBF786990B3B59B4D8A3564" LSID-ZBK="urn:lsid:zoobank.org:act:669CE437-D461-4DF0-AF57-F94B3740CCAE" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius prooculis Chapman & Schultz 1952:519" order="Perciformes" pageNumber="1" phylum="Chordata" rank="species" species="prooculis">E. prooculis</taxonomicName>
|
|
) that exhibits crescentic markings, thus duplicating the dark spots of other species, and it is inferred from this that the character states are homologs. The positions of the dark or pale spots on the ventral surface of the head are the same in both sexes of the same species and occupy the same relative positions in all species that have them. The anterior and posterior ends of the crescentic marks occupy the same positions as the spots. The anterior spot on each side is associated with the anteriormost sensory pore of the mandibular series. The posterior spot is in the area external to the attachments of the branchiostegals to the anterior ceratohyal, which attachments can be seen through the skin.
|
|
</paragraph>
|
|
<paragraph id="9750E49EC7A22F7EB6E582B7455D1C7A" pageNumber="2">
|
|
The presence of these markings appears to be related to size or sexual maturity of individuals, but apparently sexually mature individuals may also fail to exhibit them. Even though these markings are not exhibited by every specimen, or even most of the specimens of any of the species, their presence in some specimens of each species and not, with one exception, in any other
|
|
<taxonomicName id="DA407641490966E4E1C63400A4D4DB2E" LSID-ZBK="urn:lsid:zoobank.org:act:B3F80107-BABB-495D-BFB7-721DCA10C4F8" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius McCulloch 1923:121" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Ecsenius</taxonomicName>
|
|
species, suffices our hypothesis of synapomorphy. The exception is
|
|
<taxonomicName id="2028F479539E50D9BDB156E0F9DBA5FF" LSID-ZBK="urn:lsid:zoobank.org:act:5B983FD8-DF45-4378-B931-F1BA6B80A391" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius isos McKinney & Springer 1976:5" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="isos">E. isos McKinney & Springer</taxonomicName>
|
|
, one of the three species of the
|
|
<taxonomicName id="F10F938EC23014EC2FCCE545707E8483" LSID-ZBK="urn:lsid:zoobank.org:act:237E3222-7E4C-4ABA-9D06-C304025DABAC" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius Ecsenius trilineatus Springer 1972:2" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="trilineatus">Trilineatus</taxonomicName>
|
|
species group. In
|
|
<taxonomicName id="D3D5BDA95574F5A336C2AEC143E798D7" LSID-ZBK="urn:lsid:zoobank.org:act:5B983FD8-DF45-4378-B931-F1BA6B80A391" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius isos McKinney & Springer 1976:5" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="isos">E. isos</taxonomicName>
|
|
, there is a dark spot (Springer, 1988, figure 53) in the same area as the posterior spot of the
|
|
<taxonomicName id="EEC5B061C88268EFAC6D8DC5AB936061" LSID-ZBK="urn:lsid:zoobank.org:act:669CE437-D461-4DF0-AF57-F94B3740CCAE" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius prooculis Chapman & Schultz 1952:519" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
|
|
group species. This spot appears to be an isolation of the ventral end of a dark crescentic marking that begins dorsally on the opercle and curves ventroanteriorly, ending on the ventral surface of the head in the other two species of the group. There may be other isolated spots continuing dorsal to the ventral one in
|
|
<taxonomicName id="339FD4D933FF807E0D7C95E1B9B1EAE9" LSID-ZBK="urn:lsid:zoobank.org:act:5B983FD8-DF45-4378-B931-F1BA6B80A391" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius isos McKinney & Springer 1976:5" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="isos">E. isos</taxonomicName>
|
|
; these represent positions along which the dark crescent mark lies in the other two species (Springer, 1991: figure 2).
|
|
</paragraph>
|
|
<paragraph id="DC119F78D4FC3ED7AC04A82EDFC84501" pageNumber="2">
|
|
The validity of the
|
|
<taxonomicName id="3BA6042D8C8FB654F20D1AB50E7892C7" LSID-ZBK="urn:lsid:zoobank.org:act:669CE437-D461-4DF0-AF57-F94B3740CCAE" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius prooculis Chapman & Schultz 1952:519" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
|
|
group is supported by the distributions of its component species. Seven of the eight species are allopatrically distributed; the eighth species,
|
|
<taxonomicName id="2C2CCC4CAD140EE6990A5FFDA58797F5" LSID-ZBK="urn:lsid:zoobank.org:act:C6554AD6-8A16-41DD-B379-168B0416A3FB" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius Ecsenius bimaculatus Springer 1971:38" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="bimaculatus">E. bimaculatus Springer</taxonomicName>
|
|
, has a partially allopatric distribution, but overlaps the distributions of two of the other species (Figure 3).
|
|
</paragraph>
|
|
<paragraph id="99875AFFEA31CFB7FB52DB9EA913AA49" pageNumber="2">
|
|
Other characters defining the group within
|
|
<taxonomicName id="F202AD4680830994E47D50F5F0FA4B3F" LSID-ZBK="urn:lsid:zoobank.org:act:B3F80107-BABB-495D-BFB7-721DCA10C4F8" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius McCulloch 1923:121" order="Perciformes" pageNumber="2" phylum="Chordata" rank="genus">Ecsenius</taxonomicName>
|
|
: Small species (largest known specimen 40 mm SL, attained by only one species,
|
|
<taxonomicName id="800AA294E9FE3F3785C598E2477CE622" LSID-ZBK="urn:lsid:zoobank.org:act:669CE437-D461-4DF0-AF57-F94B3740CCAE" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius prooculis Chapman & Schultz 1952:519" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="prooculis">E. prooculis Chapman & Schultz</taxonomicName>
|
|
other species not exceeding 34 mm SL); dorsal-fin spines XI-XIII (strongly modally XII), segmented rays 13-15 (strongly modally 14), deeply notched between spines and segmented rays; anal fin II,15-17 (modally 16); pectoral-fin rays 12-14 (strongly modally 14); segmented caudal-fin rays 13, third from dorsalmost and third from ventralmost filamentous in adult males; vertebrae 10 + 21-23 (strongly modally 22); total dentary teeth (including anterior canines, but excluding posterior canines) 39-50, posterior canine teeth 1 on each side; lateral-line a continuous tube extending posteriorly to vertical from 8th to 11th dorsalfin spine (to 11th in only one specimen of
|
|
<taxonomicName id="F261DAF9B40F2861734E8B9EAC0E6F6D" LSID-ZBK="urn:lsid:zoobank.org:act:C6554AD6-8A16-41DD-B379-168B0416A3FB" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius Ecsenius bimaculatus Springer 1971:38" order="Perciformes" pageNumber="2" phylum="Chordata" rank="species" species="bimaculatus">E. bimaculatus</taxonomicName>
|
|
); nasal cirrus minute; no distinct dark stripes or spots on fleshy pectoral-fin base.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |