311 lines
38 KiB
XML
311 lines
38 KiB
XML
<document id="BD3942CE64A729BC1C02DC6A1F78B280" ID-DOI="10.11646/zootaxa.5338.1.1" ID-GBIF-Dataset="e7148a02-ac1a-4172-9071-606bd1e703fc" ID-ISSN="11755334" ID-Zenodo-Dep="8283738" ID-ZooBank="CAA37212-3008-43F4-98DE-94D7A3FFD3B6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="karina" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1692955869775" checkinUser="plazi" docAuthor="Mu, Yannan & Zhang, Feng" docDate="2023" docId="364E87DE867DFF9D5AA6F8E2DB3A215B" docLanguage="en" docName="zootaxa.5338.1.1.pdf" docOrigin="Zootaxa 5338 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5338.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Beatitas octomaculatus, sp. nov." docType="treatment" docVersion="7" lastPageNumber="21" masterDocId="CA77FFA68673FF895A31FFEBD84D203E" masterDocTitle="Further additions to the guardstone spider fauna from China (Araneae: Phrurolithidae)" masterLastPageNumber="104" masterPageNumber="1" pageNumber="15" updateTime="1720090975140" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="A9425CF3B4EC016B648748F1F6985CDA">
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<mods:title id="7F53496B80DE75224437A7B73CD88A0E">Further additions to the guardstone spider fauna from China (Araneae: Phrurolithidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3F5302C746FFE94934C6699ECE80C924" type="personal">
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<mods:roleTerm id="1EBE460F1F46E4D3E94059D29D801931">Author</mods:roleTerm>
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<mods:namePart id="A26A0395844F4EDD1F35D171E9CA02D9">Mu, Yannan</mods:namePart>
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<mods:affiliation id="1519EB64536525567537769AAF1A170A">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Developed, Hebei University, Baoding, Hebei 071002, P. R. China & 15188605531 @ 163. com; https: // orcid. org / 0000 - 0002 - 2504 - 673 X</mods:affiliation>
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<mods:roleTerm id="14718BD77393F155082F6727CAD6FABB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="71CC8234392156C56E57CD5593088A9F">Zhang, Feng</mods:namePart>
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<mods:affiliation id="B1508DD3FFADE1869EFE8F5A3D9971A1">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Developed, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<mods:title id="9D681085557E146E6CDAB4FA6D8F13DF">Zootaxa</mods:title>
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<mods:part id="6BAE706C166B8E29292DC61E63F5ED48">
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<mods:date id="3E85F55735AAAF6B450571D91F78A0D5">2023</mods:date>
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<mods:number id="CCB1A8A32571E798E6A68C6962199345">2023-08-25</mods:number>
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<mods:number id="7A30BEBB0C7F7F3CA5A7751FC7344EB7">5338</mods:number>
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<mods:url id="157C2C8B1DFD853F85335BD1003F531A">http://dx.doi.org/10.11646/zootaxa.5338.1.1</mods:url>
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<mods:classification id="BAA67351A9FF2CC16653C8D2C63704DA">journal article</mods:classification>
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<mods:identifier id="DAD1E1AEE4F636EECE771C084B02909D" type="DOI">10.11646/zootaxa.5338.1.1</mods:identifier>
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<mods:identifier id="63A8F372DB6B6729640AC64ECF2D7210" type="Zenodo-Dep">8283738</mods:identifier>
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<treatment id="364E87DE867DFF9D5AA6F8E2DB3A215B" ID-DOI="http://doi.org/10.5281/zenodo.8283793" ID-GBIF-Taxon="212459902" ID-Zenodo-Dep="8283793" LSID="urn:lsid:plazi:treatment:364E87DE867DFF9D5AA6F8E2DB3A215B" httpUri="http://treatment.plazi.org/id/364E87DE867DFF9D5AA6F8E2DB3A215B" lastPageId="20" lastPageNumber="21" pageId="14" pageNumber="15">
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<subSubSection id="F6FD6543867DFF875AA6F8E2DAEB271D" box="[151,678,1801,1828]" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="BE5836C8867DFF875AA6F8E2DAEB271D" blockId="14.[151,678,1801,1863]" box="[151,678,1801,1828]" pageId="14" pageNumber="15">
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<heading id="E51081A4867DFF875AA6F8E2DAEB271D" bold="true" box="[151,678,1801,1828]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="8C93EADA867DFF875AA6F8E2DAEB271D" bold="true" box="[151,678,1801,1828]" pageId="14" pageNumber="15">
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<taxonomicName id="79E74D4B867DFF875AA6F8E2D9FD271A" ID-CoL="C6GRM" authority="Mu & Zhang, 2023" box="[151,432,1801,1828]" class="Arachnida" family="Phrurolithidae" genus="Beatitas" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. nov.">
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<emphasis id="8C93EADA867DFF875AA6F8E2D9FD271A" bold="true" box="[151,432,1801,1828]" italics="true" pageId="14" pageNumber="15">Beatitas octomaculatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="97A057A1867DFF875B89F8E2DA58271D" box="[440,533,1801,1827]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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(ĀDzffiff)
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="F6FD6543867DFF875AA6F8C6D94B2779" box="[151,262,1837,1863]" pageId="14" pageNumber="15" type="description">
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||
<paragraph id="BE5836C8867DFF875AA6F8C6D94B2779" blockId="14.[151,678,1801,1863]" box="[151,262,1837,1863]" pageId="14" pageNumber="15">
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||
<figureCitation id="26DC2A4D867DFF875AA6F8C6D94B2779" box="[151,262,1837,1863]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStart-2="FIGURE 11" captionStart-3="FIGURE 12" captionStart-4="FIGURE 13" captionStartId-0="15.[151,250,1609,1633]" captionStartId-1="16.[151,250,1768,1792]" captionStartId-2="17.[151,250,1908,1932]" captionStartId-3="18.[151,250,1223,1247]" captionStartId-4="19.[151,250,1565,1589]" captionTargetBox-0="[159,1427,188,1578]" captionTargetBox-1="[158,1431,187,1735]" captionTargetBox-2="[203,1396,196,1884]" captionTargetBox-3="[152,1436,181,1199]" captionTargetBox-4="[151,1415,182,1520]" captionTargetId-0="figure-17@15.[151,1436,181,1586]" captionTargetId-1="figure-21@16.[151,1436,181,1745]" captionTargetId-2="figure-17@17.[191,1396,181,1884]" captionTargetId-3="figure-21@18.[151,1436,181,1199]" captionTargetId-4="figure-17@19.[151,1435,181,1542]" captionTargetPageId-0="15" captionTargetPageId-1="16" captionTargetPageId-2="17" captionTargetPageId-3="18" captionTargetPageId-4="19" captionText-0="FIGURE 9. Habitus of Beatitas octomaculatus sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." captionText-1="FIGURE 10. Left legs of Beatitas octomaculatus sp. nov. (male holotype): A. Leg I, prolateral view; B. Same, retrolateral view; C. Leg II, prolateral view; D. Same, retrolateral view; E. Leg III, prolateral view; F. Same retrolateral view; G. Leg IV, prolateral view; H. Same, retrolateral view." captionText-2="FIGURE 11. Beatitas octomaculatus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Ocular area, dorsal view; F. Cephalothorax, frontal view. Abbreviations: E—embolus; C— conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; RvTA—retroventral tibial apophysis; SD—sperm duct; TA—tegular apophysis." captionText-3="FIGURE 12. Beatitas octomaculatus sp. nov.: A. Epigyne, ventral view; B. Same, dorsal view; C. Ocular area, dorsal view; D. Male spinnerets; E. Female spinnerets. Abbreviations: B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; S—spermathecae." captionText-4="FIGURE 13. Copulatory organs of Beatitas octomaculatus sp. nov.: A. Male left palp, ventral view; B. Same, retrolateral view; C. Same, dorsal view; D. Epigyne, ventral view; E. Same, dorsal view. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8283765" figureDoi-1="http://doi.org/10.5281/zenodo.8283771" figureDoi-2="http://doi.org/10.5281/zenodo.8283773" figureDoi-3="http://doi.org/10.5281/zenodo.8283777" figureDoi-4="http://doi.org/10.5281/zenodo.8283781" httpUri-0="https://zenodo.org/record/8283765/files/figure.png" httpUri-1="https://zenodo.org/record/8283771/files/figure.png" httpUri-2="https://zenodo.org/record/8283773/files/figure.png" httpUri-3="https://zenodo.org/record/8283777/files/figure.png" httpUri-4="https://zenodo.org/record/8283781/files/figure.png" pageId="14" pageNumber="15">Figs 9–13</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F6FD6543867DFF875AA6F89EDAA327E9" pageId="14" pageNumber="15" type="materials_examined">
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<paragraph id="BE5836C8867DFF875AA6F89EDD48278A" blockId="14.[151,1436,1909,2010]" pageId="14" pageNumber="15">
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<emphasis id="8C93EADA867DFF875AA6F89ED90627B1" bold="true" box="[151,331,1909,1935]" pageId="14" pageNumber="15">Type material.</emphasis>
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<materialsCitation id="0E8F3C95867DFF875B69F89EDD48278A" ID-GBIF-Occurrence="4400675312" collectingDate="2021-05-11" collectorName="Yannan Mu." country="China" county="Autonomous Region" elevation="830" latitude="24.629858" location="Pinggui district" longLatPrecision="1" longitude="111.52969" municipality="Hezhou City" pageId="14" pageNumber="15" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
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<emphasis id="8C93EADA867DFF875B69F89ED98827B1" bold="true" box="[344,453,1909,1935]" pageId="14" pageNumber="15">
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||
<typeStatus id="615C886A867DFF875B69F89ED98827B1" box="[344,453,1909,1935]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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♁,
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<emphasis id="8C93EADA867DFF875BCAF89EDA2F27B1" bold="true" box="[507,610,1909,1935]" pageId="14" pageNumber="15">
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||
<collectingCountry id="C6F07658867DFF875BCAF89EDA1027B1" box="[507,605,1909,1935]" name="China" pageId="14" pageNumber="15">CHINA</collectingCountry>
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:
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</emphasis>
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<emphasis id="8C93EADA867DFF87585EF89DDC6A27B1" box="[623,1063,1909,1936]" italics="true" pageId="14" pageNumber="15">
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||
<collectingRegion id="7C23F82A867DFF87585EF89DDB7E27B1" box="[623,819,1909,1936]" country="China" name="Guangxi" pageId="14" pageNumber="15">Guangxi Zhuang</collectingRegion>
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<collectingCounty id="57394E44867DFF87590EF89DDC6A27B1" box="[831,1063,1910,1936]" pageId="14" pageNumber="15">Autonomous Region</collectingCounty>
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</emphasis>
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||
:
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<collectingMunicipality id="5E3CACB2867DFF875E0DF89EDC9F27B1" box="[1084,1234,1909,1935]" pageId="14" pageNumber="15">Hezhou City</collectingMunicipality>
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,
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<location id="BB386013867DFF875ED2F89EDDD527B1" LSID="urn:lsid:plazi:treatment:364E87DE867DFF9D5AA6F8E2DB3A215B:BB386013867DFF875ED2F89EDDD527B1" box="[1251,1432,1909,1935]" country="China" county="Autonomous Region" latitude="24.629858" longLatPrecision="1" longitude="111.52969" municipality="Hezhou City" name="Pinggui district" pageId="14" pageNumber="15" stateProvince="Guangxi">Pinggui district</location>
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,
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<location id="BB386013867DFF875AA6F872D944278D" LSID="urn:lsid:plazi:treatment:364E87DE867DFF9D5AA6F8E2DB3A215B:BB386013867DFF875AA6F872D944278D" box="[151,265,1945,1971]" country="China" county="Autonomous Region" latitude="24.629858" longLatPrecision="1" longitude="111.52969" municipality="Hezhou City" name="Guposhan" pageId="14" pageNumber="15" stateProvince="Guangxi">Guposhan</location>
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scenic spot (
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<geoCoordinate id="DBD3500F867DFF875BABF870DA0B2788" box="[410,582,1947,1974]" degrees="24" direction="north" minutes="37" orientation="latitude" pageId="14" pageNumber="15" precision="1" seconds="47.49" value="24.629858">24°37′47.49″N</geoCoordinate>
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,
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<geoCoordinate id="DBD3500F867DFF875863F870DB4B2788" box="[594,774,1947,1974]" degrees="111" direction="east" minutes="31" orientation="longitude" pageId="14" pageNumber="15" precision="1" seconds="46.89" value="111.52969">111°31′46.89″E</geoCoordinate>
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,
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<elevation id="35CAD1FB867DFF875920F872DBDE278D" box="[785,915,1945,1972]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="14" pageNumber="15" unit="m" value="830.0">
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<quantity id="791F9B2D867DFF875920F872DB15278A" box="[785,856,1945,1972]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="14" pageNumber="15" unit="m" value="830.0">830 m</quantity>
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a.s.l.
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</elevation>
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),
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<date id="CA591008867DFF87599BF872DC0C278D" box="[938,1089,1945,1972]" pageId="14" pageNumber="15" value="2021-05-11">
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<collectingDate id="DA1DE9E0867DFF87599BF872DC0C278D" box="[938,1089,1945,1972]" pageId="14" pageNumber="15" value="2021-05-11">11 May 2021</collectingDate>
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</date>
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, leg.
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<collectorName id="1312531E867DFF875E4DF871DD48278A" box="[1148,1285,1946,1972]" pageId="14" pageNumber="15">Yannan Mu.</collectorName>
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="BE5836C8867DFF875AF6F856DAA327E9" blockId="14.[151,1436,1909,2010]" box="[199,750,1981,2010]" pageId="14" pageNumber="15">
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||
<materialsCitation id="0E8F3C95867DFF875AF6F856DAA727E9" ID-GBIF-Occurrence="4400675330" box="[199,746,1981,2010]" collectingDate="2021-05-11" collectorName="Yannan Mu." country="China" county="Autonomous Region" elevation="830" latitude="24.629858" location="Pinggui district" longLatPrecision="1" longitude="111.52969" municipality="Hezhou City" pageId="14" pageNumber="15" specimenCount="9" specimenCount-female="9" stateProvince="Guangxi" typeStatus="paratype">
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||
<emphasis id="8C93EADA867DFF875AF6F856D97227E9" bold="true" box="[199,319,1981,2007]" pageId="14" pageNumber="15">
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||
<typeStatus id="615C886A867DFF875AF6F856D97227E9" box="[199,319,1981,2007]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
: 2 ♁
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||
<specimenCount id="A8E1FD41867DFF875B4EF856D9E427E6" box="[383,425,1981,2008]" pageId="14" pageNumber="15" type="female">9 ♀</specimenCount>
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, with same data as holotype
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<caption id="EA986640867CFF865AA6F9A2DA2826B8" ID-DOI="http://doi.org/10.5281/zenodo.8283765" ID-Zenodo-Dep="8283765" httpUri="https://zenodo.org/record/8283765/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,250,1609,1633]" targetBox="[159,1427,188,1578]" targetPageId="15" targetType="figure">
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||
<paragraph id="BE5836C8867CFF865AA6F9A2DA2826B8" blockId="15.[151,1437,1609,1670]" pageId="15" pageNumber="16">
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||
<emphasis id="8C93EADA867CFF865AA6F9A2D958265C" bold="true" box="[151,277,1609,1634]" pageId="15" pageNumber="16">FIGURE 9.</emphasis>
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Habitus of
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<taxonomicName id="79E74D4B867CFF865BA2F9A1DACF265C" box="[403,642,1610,1634]" class="Arachnida" family="Phrurolithidae" genus="Beatitas" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. nov.">
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<emphasis id="8C93EADA867CFF865BA2F9A1DACF265C" box="[403,642,1610,1634]" italics="true" pageId="15" pageNumber="16">Beatitas octomaculatus</emphasis>
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</taxonomicName>
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||
<emphasis id="8C93EADA867CFF8658BAF9A1DA90265C" bold="true" box="[651,733,1610,1634]" pageId="15" pageNumber="16">
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||
<taxonomicNameLabel id="97A057A1867CFF8658BAF9A1DA90265C" box="[651,733,1610,1634]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
:
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<emphasis id="8C93EADA867CFF8658DCF9A2DB4B265F" bold="true" box="[749,774,1609,1633]" pageId="15" pageNumber="16">A.</emphasis>
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Male holotype, dorsal view;
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<emphasis id="8C93EADA867CFF865E0BF9A1DC1C265C" bold="true" box="[1082,1105,1610,1634]" pageId="15" pageNumber="16">B.</emphasis>
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Same, ventral view;
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<emphasis id="8C93EADA867CFF865F01F9A2DD04265F" bold="true" box="[1328,1353,1609,1633]" pageId="15" pageNumber="16">C.</emphasis>
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||
Female paratype, dorsal view;
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<emphasis id="8C93EADA867CFF865B4FF985D9DA26B8" bold="true" box="[382,407,1646,1670]" pageId="15" pageNumber="16">D.</emphasis>
|
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Same, ventral view.
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||
</paragraph>
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</caption>
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<subSubSection id="F6FD6543867CFF865AF6F933DAD22728" pageId="15" pageNumber="16" type="etymology">
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||
<paragraph id="BE5836C8867CFF865AF6F933DAD22728" blockId="15.[151,1437,1752,1994]" pageId="15" pageNumber="16">
|
||
<emphasis id="8C93EADA867CFF865AF6F933D90326CC" bold="true" box="[199,334,1752,1778]" pageId="15" pageNumber="16">Etymology.</emphasis>
|
||
This species name is a combination of “
|
||
<emphasis id="8C93EADA867CFF86591EF932DBEA26CC" box="[815,935,1753,1778]" italics="true" pageId="15" pageNumber="16">octonarius</emphasis>
|
||
” and “
|
||
<emphasis id="8C93EADA867CFF8659CCF933DC0226CC" box="[1021,1103,1752,1778]" italics="true" pageId="15" pageNumber="16">macula</emphasis>
|
||
”, referring to the eight light markings on the abdominal dorsum; adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F6FD6543867CFF865AF6F8CBDCD627F4" pageId="15" pageNumber="16" type="diagnosis">
|
||
<paragraph id="BE5836C8867CFF865AF6F8CBDCD627F4" blockId="15.[151,1437,1752,1994]" pageId="15" pageNumber="16">
|
||
<emphasis id="8C93EADA867CFF865AF6F8CBD90F2704" bold="true" box="[199,322,1824,1850]" pageId="15" pageNumber="16">Diagnosis.</emphasis>
|
||
The male of this new species can be recognized by: 1) the strong and upward-extending embolus, 2) the membranous conductor, 3) the retrolateral tibial apophysis with a bifurcated tip, and curved dorsal tibial apophysis extending towards prolateral side of cymbium. The female can be recognized by: 1) the diamond-shaped atrium on the epigynal plate, 2) the widely separated copulatory openings located at the sides of the atrium, and 3) the globular spermathecae that are almost entirely surrounded by the long connecting tubes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="EA9866408663FF995AA6F903D9AA2777" ID-DOI="http://doi.org/10.5281/zenodo.8283771" ID-Zenodo-Dep="8283771" httpUri="https://zenodo.org/record/8283771/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,250,1768,1792]" targetBox="[158,1431,187,1735]" targetPageId="16" targetType="figure">
|
||
<paragraph id="BE5836C88663FF995AA6F903D9AA2777" blockId="16.[151,1436,1768,1865]" pageId="16" pageNumber="17">
|
||
<emphasis id="8C93EADA8663FF995AA6F903D952273F" bold="true" box="[151,287,1768,1793]" pageId="16" pageNumber="17">FIGURE 10.</emphasis>
|
||
Left legs of
|
||
<taxonomicName id="79E74D4B8663FF995BAAF902DACA273F" box="[411,647,1769,1793]" class="Arachnida" family="Phrurolithidae" genus="Beatitas" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. nov.">
|
||
<emphasis id="8C93EADA8663FF995BAAF902DACA273F" box="[411,647,1769,1793]" italics="true" pageId="16" pageNumber="17">Beatitas octomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8C93EADA8663FF9958BDF902DA91273F" bold="true" box="[652,732,1769,1793]" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="97A057A18663FF9958BDF902DA91273F" box="[652,732,1769,1793]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(male holotype):
|
||
<emphasis id="8C93EADA8663FF9959BFF903DBEB273E" bold="true" box="[910,934,1768,1792]" pageId="16" pageNumber="17">A.</emphasis>
|
||
Leg I, prolateral view;
|
||
<emphasis id="8C93EADA8663FF995EBEF902DCEA273F" bold="true" box="[1167,1191,1769,1793]" pageId="16" pageNumber="17">B.</emphasis>
|
||
Same, retrolateral view;
|
||
<emphasis id="8C93EADA8663FF995AA6F8E7D8FD271A" bold="true" box="[151,176,1804,1828]" pageId="16" pageNumber="17">C.</emphasis>
|
||
Leg II, prolateral view;
|
||
<emphasis id="8C93EADA8663FF995B90F8E6D9F7271B" bold="true" box="[417,442,1805,1829]" pageId="16" pageNumber="17">D.</emphasis>
|
||
Same, retrolateral view;
|
||
<emphasis id="8C93EADA8663FF995885F8E6DA86271B" bold="true" box="[692,715,1805,1829]" pageId="16" pageNumber="17">E.</emphasis>
|
||
Leg III, prolateral view;
|
||
<emphasis id="8C93EADA8663FF9959F4F8E6DB95271B" bold="true" box="[965,984,1805,1829]" pageId="16" pageNumber="17">F.</emphasis>
|
||
Same retrolateral view;
|
||
<emphasis id="8C93EADA8663FF995EFCF8E7DCAA271A" bold="true" box="[1229,1255,1804,1828]" pageId="16" pageNumber="17">G.</emphasis>
|
||
Leg IV, prolateral view;
|
||
<emphasis id="8C93EADA8663FF995AE7F8DAD8BD2777" bold="true" box="[214,240,1841,1865]" pageId="16" pageNumber="17">H.</emphasis>
|
||
Same, retrolateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EA9866408662FF985AA6F89FD9C727C6" ID-DOI="http://doi.org/10.5281/zenodo.8283773" ID-Zenodo-Dep="8283773" httpUri="https://zenodo.org/record/8283773/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,250,1908,1932]" targetBox="[203,1396,196,1884]" targetPageId="17" targetType="figure">
|
||
<paragraph id="BE5836C88662FF985AA6F89FD9C727C6" blockId="17.[151,1436,1908,2040]" pageId="17" pageNumber="18">
|
||
<emphasis id="8C93EADA8662FF985AA6F89FD95227B2" bold="true" box="[151,287,1908,1932]" pageId="17" pageNumber="18">FIGURE 11.</emphasis>
|
||
<taxonomicName id="79E74D4B8662FF985B14F89EDA5E27B2" box="[293,531,1908,1932]" class="Arachnida" family="Phrurolithidae" genus="Beatitas" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. nov.">
|
||
<emphasis id="8C93EADA8662FF985B14F89EDA5E27B2" box="[293,531,1908,1932]" italics="true" pageId="17" pageNumber="18">Beatitas octomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8C93EADA8662FF985828F89EDA2727B3" bold="true" box="[537,618,1909,1933]" pageId="17" pageNumber="18">
|
||
<taxonomicNameLabel id="97A057A18662FF985828F89EDA2727B3" box="[537,618,1909,1933]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="8C93EADA8662FF985849F89FDADC27B2" bold="true" box="[632,657,1908,1932]" pageId="17" pageNumber="18">A.</emphasis>
|
||
Male left palp, prolateral view;
|
||
<emphasis id="8C93EADA8662FF9859EBF89EDBBC27B3" bold="true" box="[986,1009,1909,1933]" pageId="17" pageNumber="18">B.</emphasis>
|
||
Same, retrolateral view;
|
||
<emphasis id="8C93EADA8662FF985EC0F89FDD4727B2" bold="true" box="[1265,1290,1908,1932]" pageId="17" pageNumber="18">C.</emphasis>
|
||
Same, ventral view;
|
||
<emphasis id="8C93EADA8662FF985AE9F872D8BC278F" bold="true" box="[216,241,1945,1969]" pageId="17" pageNumber="18">D.</emphasis>
|
||
Same, dorsal view;
|
||
<emphasis id="8C93EADA8662FF985BFBF872D9AC278F" bold="true" box="[458,481,1945,1969]" pageId="17" pageNumber="18">E.</emphasis>
|
||
Ocular area, dorsal view;
|
||
<emphasis id="8C93EADA8662FF9858C8F872DB41278F" bold="true" box="[761,780,1945,1969]" pageId="17" pageNumber="18">F.</emphasis>
|
||
Cephalothorax, frontal view. Abbreviations: E—embolus; C— conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; RvTA—retroventral tibial apophysis; SD—sperm duct; TA—tegular apophysis.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="EA9866408661FF9B5AA6FB2CDAA32519" ID-DOI="http://doi.org/10.5281/zenodo.8283777" ID-Zenodo-Dep="8283777" httpUri="https://zenodo.org/record/8283777/files/figure.png" pageId="18" pageNumber="19" startId="18.[151,250,1223,1247]" targetBox="[152,1436,181,1199]" targetPageId="18" targetType="figure">
|
||
<paragraph id="BE5836C88661FF9B5AA6FB2CDAA32519" blockId="18.[151,1436,1223,1319]" pageId="18" pageNumber="19">
|
||
<emphasis id="8C93EADA8661FF9B5AA6FB2CD96C24E1" bold="true" box="[151,289,1223,1247]" pageId="18" pageNumber="19">FIGURE 12.</emphasis>
|
||
<taxonomicName id="79E74D4B8661FF9B5B19FB23DA5A24E1" box="[296,535,1223,1247]" class="Arachnida" family="Phrurolithidae" genus="Beatitas" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. nov.">
|
||
<emphasis id="8C93EADA8661FF9B5B19FB23DA5A24E1" box="[296,535,1223,1247]" italics="true" pageId="18" pageNumber="19">Beatitas octomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8C93EADA8661FF9B582FFB2CDA3D24E1" bold="true" box="[542,624,1223,1247]" pageId="18" pageNumber="19">
|
||
<taxonomicNameLabel id="97A057A18661FF9B582FFB2CDA3D24E1" box="[542,624,1223,1247]" pageId="18" pageNumber="19" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="8C93EADA8661FF9B584EFB2CDAD524E1" bold="true" box="[639,664,1223,1247]" pageId="18" pageNumber="19">A.</emphasis>
|
||
Epigyne, ventral view;
|
||
<emphasis id="8C93EADA8661FF9B59BEFB2CDBEA24E1" bold="true" box="[911,935,1223,1247]" pageId="18" pageNumber="19">B.</emphasis>
|
||
Same, dorsal view;
|
||
<emphasis id="8C93EADA8661FF9B5E4BFB2CDCDF24E1" bold="true" box="[1146,1170,1223,1247]" pageId="18" pageNumber="19">C.</emphasis>
|
||
Ocular area, dorsal view;
|
||
<emphasis id="8C93EADA8661FF9B5AA6FB00D8FD253D" bold="true" box="[151,176,1259,1283]" pageId="18" pageNumber="19">D.</emphasis>
|
||
Male spinnerets;
|
||
<emphasis id="8C93EADA8661FF9B5B5DFB00D9CE253D" bold="true" box="[364,387,1259,1283]" pageId="18" pageNumber="19">E.</emphasis>
|
||
Female spinnerets. Abbreviations: B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; S—spermathecae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="F6FD65438661FF9D5AF6FA92D97A217F" lastPageId="20" lastPageNumber="21" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="BE5836C88661FF9B5AF6FA92DBA92655" blockId="18.[151,1437,1401,1968]" pageId="18" pageNumber="19">
|
||
<emphasis id="8C93EADA8661FF9B5AF6FA92D91425AD" bold="true" box="[199,345,1401,1427]" pageId="18" pageNumber="19">Description.</emphasis>
|
||
Male (
|
||
<typeStatus id="615C886A8661FF9B5B96FA92DA5B25AD" box="[423,534,1401,1427]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
|
||
): total length 2.27, carapace 1.06 long, 0.89 wide; abdomen 1.21 long, 0.84 wide. Eye sizes and interdistances:AME 0.06, ALE 0.08, PME 0.05, PLE 0.06; AME–AME 0.03, AME–ALE 0.01, ALE– ALE 0.15, PME–PME 0.05, PME–PLE 0.04, PLE–PLE 0.23, ALE–PLE 0.03. EAW 0.32, CRW 0.45, EAW/CRW 0.71, CRW/CW 0.51. MOA 0.15 long, anterior width 0.15, posterior width 0.15.
|
||
<collectingCountry id="C6F076588661FF9B5E15FA0EDC0625C1" box="[1060,1099,1509,1535]" name="Switzerland" pageId="18" pageNumber="19">CH</collectingCountry>
|
||
0.13,
|
||
<collectingCountry id="C6F076588661FF9B5EA3FA0EDCF525C1" box="[1170,1208,1509,1535]" name="Switzerland" pageId="18" pageNumber="19">CH</collectingCountry>
|
||
/AME 2.17. Labium 0.12 long, 0.17 wide. Sternum 0.61 long, 0.56 wide. Leg measurements: Ⅰ 3.22 (0.88, 0.37, 0.84, 0.78, 0.35), II 2.87 (0.80, 0.36, 0.65, 0.65, 0.41), III 2.56 (0.71, 0.32, 0.49, 0.62, 0.42), IV 3.80 (0.98, 0.36, 0.88, 0.97, 0.61). Spination: femora I–IV d 1, tibia Ⅰ pv 6 rv 5, tibia II pv 5 rv 5, metatarsi Ⅰ–II pv 4 rv 3.
|
||
</paragraph>
|
||
<paragraph id="BE5836C88661FF9B5AF6F99EDD2B26E9" blockId="18.[151,1437,1401,1968]" pageId="18" pageNumber="19">
|
||
Coloration. Carapace black, fovea round, with several markings resembling flowing water droplets beside fovea. Abdomen black, covered entirely by dorsal scutum, with four pairs of light markings on abdominal dorsum (
|
||
<figureCitation id="26DC2A4D8661FF9B5AAEF956D95426E9" box="[159,281,1725,1751]" captionStart="FIGURE 9" captionStartId="15.[151,250,1609,1633]" captionTargetBox="[159,1427,188,1578]" captionTargetId="figure-17@15.[151,1436,181,1586]" captionTargetPageId="15" captionText="FIGURE 9. Habitus of Beatitas octomaculatus sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8283765" httpUri="https://zenodo.org/record/8283765/files/figure.png" pageId="18" pageNumber="19">Fig. 9A, B</figureCitation>
|
||
). Legs yellow except black femora; patellae, tibiae and metatarsi I–IV with black annuli (
|
||
<figureCitation id="26DC2A4D8661FF9B5F34F956DD1426E9" box="[1285,1369,1725,1751]" captionStart="FIGURE 10" captionStartId="16.[151,250,1768,1792]" captionTargetBox="[158,1431,187,1735]" captionTargetId="figure-21@16.[151,1436,181,1745]" captionTargetPageId="16" captionText="FIGURE 10. Left legs of Beatitas octomaculatus sp. nov. (male holotype): A. Leg I, prolateral view; B. Same, retrolateral view; C. Leg II, prolateral view; D. Same, retrolateral view; E. Leg III, prolateral view; F. Same retrolateral view; G. Leg IV, prolateral view; H. Same, retrolateral view." figureDoi="http://doi.org/10.5281/zenodo.8283771" httpUri="https://zenodo.org/record/8283771/files/figure.png" pageId="18" pageNumber="19">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BE5836C88661FF9B5AF6F90ADC112791" blockId="18.[151,1437,1401,1968]" pageId="18" pageNumber="19">
|
||
Palp as in
|
||
<figureCitation id="26DC2A4D8661FF9B5B77F90AD99826C5" box="[326,469,1761,1787]" captionStart="FIGURE 11" captionStartId="17.[151,250,1908,1932]" captionTargetBox="[203,1396,196,1884]" captionTargetId="figure-17@17.[191,1396,181,1884]" captionTargetPageId="17" captionText="FIGURE 11. Beatitas octomaculatus sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Ocular area, dorsal view; F. Cephalothorax, frontal view. Abbreviations: E—embolus; C— conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; RvTA—retroventral tibial apophysis; SD—sperm duct; TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.8283773" httpUri="https://zenodo.org/record/8283773/files/figure.png" pageId="18" pageNumber="19">Figs 11A–D</figureCitation>
|
||
,
|
||
<figureCitation id="26DC2A4D8661FF9B5BD5F90ADA7426C5" box="[484,569,1761,1787]" captionStart="FIGURE 13" captionStartId="19.[151,250,1565,1589]" captionTargetBox="[151,1415,182,1520]" captionTargetId="figure-17@19.[151,1435,181,1542]" captionTargetPageId="19" captionText="FIGURE 13. Copulatory organs of Beatitas octomaculatus sp. nov.: A. Male left palp, ventral view; B. Same, retrolateral view; C. Same, dorsal view; D. Epigyne, ventral view; E. Same, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8283781" httpUri="https://zenodo.org/record/8283781/files/figure.png" pageId="18" pageNumber="19">13A–C</figureCitation>
|
||
. Femoral apophysis round, protruded near tip of femur. Retroventral tibial apophysis strong, strongly curved, extending towards ventral side, middle parts wider than base and tip bifurcated as U-shaped; dorsal tibial apophysis digitiform, base wide and tip curved towards prolateral side of cymbium. Tegulum as wide as cymbium, diamond-shaped; tegular apophysis triangular, located at retrolateral tip of tegulum. Sperm duct distinct, U-shaped, tapering from retrolateral side of tegulum to embolus. Embolus strong, tip slightly curved, beak-shaped, directed retroventrally. Conductor large, membranous, close to embolus.
|
||
</paragraph>
|
||
<caption id="EA9866408660FF9A5AA6F9F6DC7B2667" ID-DOI="http://doi.org/10.5281/zenodo.8283781" ID-Zenodo-Dep="8283781" httpUri="https://zenodo.org/record/8283781/files/figure.png" pageId="19" pageNumber="20" startId="19.[151,250,1565,1589]" targetBox="[151,1415,182,1520]" targetPageId="19" targetType="figure">
|
||
<paragraph id="BE5836C88660FF9A5AA6F9F6DC7B2667" blockId="19.[151,1436,1565,1626]" pageId="19" pageNumber="20">
|
||
<emphasis id="8C93EADA8660FF9A5AA6F9F6D952260B" bold="true" box="[151,287,1565,1589]" pageId="19" pageNumber="20">FIGURE 13.</emphasis>
|
||
Copulatory organs of
|
||
<taxonomicName id="79E74D4B8660FF9A5BCEF9F5DAA12608" box="[511,748,1566,1590]" class="Arachnida" family="Phrurolithidae" genus="Beatitas" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="octomaculatus" status="sp. nov.">
|
||
<emphasis id="8C93EADA8660FF9A5BCEF9F5DAA12608" box="[511,748,1566,1590]" italics="true" pageId="19" pageNumber="20">Beatitas octomaculatus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="8C93EADA8660FF9A58C0F9F5DB0C2608" bold="true" box="[753,833,1566,1590]" pageId="19" pageNumber="20">
|
||
<taxonomicNameLabel id="97A057A18660FF9A58C0F9F5DB0C2608" box="[753,833,1566,1590]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
:
|
||
<emphasis id="8C93EADA8660FF9A597FF9F6DB2A260B" bold="true" box="[846,871,1565,1589]" pageId="19" pageNumber="20">A.</emphasis>
|
||
Male left palp, ventral view;
|
||
<emphasis id="8C93EADA8660FF9A5EBFF9F5DCE82608" bold="true" box="[1166,1189,1566,1590]" pageId="19" pageNumber="20">B.</emphasis>
|
||
Same, retrolateral view;
|
||
<emphasis id="8C93EADA8660FF9A5AA6F9AAD8FD2667" bold="true" box="[151,176,1601,1625]" pageId="19" pageNumber="20">C.</emphasis>
|
||
Same, dorsal view;
|
||
<emphasis id="8C93EADA8660FF9A5B4FF9A9D9DA2664" bold="true" box="[382,407,1602,1626]" pageId="19" pageNumber="20">D.</emphasis>
|
||
Epigyne, ventral view;
|
||
<emphasis id="8C93EADA8660FF9A58B8F9A9DAEC2664" bold="true" box="[649,673,1602,1626]" pageId="19" pageNumber="20">E.</emphasis>
|
||
Same, dorsal view. Scale bars: 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="BE5836C88660FF9A5AF6F947DA5427FC" blockId="19.[151,1437,1708,2022]" pageId="19" pageNumber="20">
|
||
Female (One
|
||
<typeStatus id="615C886A8660FF9A5B50F947D98A26F8" box="[353,455,1708,1734]" pageId="19" pageNumber="20" type="paratype">paratype</typeStatus>
|
||
): Total length 2.86, carapace 1.20 long, 1.02 wide; abdomen 1.66 long, 1.37 wide. Eye sizes and interdistances: AME 0.05, ALE 0.08, PME 0.06, PLE 0.07; AME–AME 0.05, AME–ALE 0.01, ALE– ALE 0.17, PME–PME 0.05, PME–PLE 0.04, PLE–PLE 0.24, ALE–PLE 0.04. EAW 0.35, CRW 0.50, EAW/CRW 0.70, CRW/CW 0.49. MOA 0.16 long, anterior width 0.16, posterior width 0.16.
|
||
<collectingCountry id="C6F076588660FF9A5E15F8F3DC06270C" box="[1060,1099,1816,1842]" name="Switzerland" pageId="19" pageNumber="20">CH</collectingCountry>
|
||
0.14,
|
||
<collectingCountry id="C6F076588660FF9A5EA3F8F3DCF5270C" box="[1170,1208,1816,1842]" name="Switzerland" pageId="19" pageNumber="20">CH</collectingCountry>
|
||
/AME 2.80. Labium 0.15 long, 0.19 wide. Sternum 0.71 long, 0.64 wide. Leg measurements: Ⅰ 3.65 (0.97, 0.41, 0.99, 0.88, 0.40), II 3.10 (0.82, 0.38, 0.75, 0.74, 0.41), III 2.85 (0.79, 0.38, 0.54, 0.68, 0.46), IV4.05 (1.03, 0.37, 0.98, 1.02, 0.65). Spination: femora I–IV d 1, tibia Ⅰ pv 6 rv 6, tibia II pv 6 rv 5, metatarsi Ⅰ–II pv 4 rv 3. Other characters as for male, except dorsal scutum absent (
|
||
<figureCitation id="26DC2A4D8660FF9A5BA0F843DA4127FC" box="[401,524,1960,1986]" captionStart="FIGURE 9" captionStartId="15.[151,250,1609,1633]" captionTargetBox="[159,1427,188,1578]" captionTargetId="figure-17@15.[151,1436,181,1586]" captionTargetPageId="15" captionText="FIGURE 9. Habitus of Beatitas octomaculatus sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8283765" httpUri="https://zenodo.org/record/8283765/files/figure.png" pageId="19" pageNumber="20">Fig. 9C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BE5836C88660FF9D5AF6F827D97A217F" blockId="19.[151,1437,1708,2022]" lastBlockId="20.[151,1437,151,357]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
|
||
Epigyne as in
|
||
<figureCitation id="26DC2A4D8660FF9A5B5AF827D9BA27D8" box="[363,503,1996,2022]" captionStart="FIGURE 12" captionStartId="18.[151,250,1223,1247]" captionTargetBox="[152,1436,181,1199]" captionTargetId="figure-21@18.[151,1436,181,1199]" captionTargetPageId="18" captionText="FIGURE 12. Beatitas octomaculatus sp. nov.: A. Epigyne, ventral view; B. Same, dorsal view; C. Ocular area, dorsal view; D. Male spinnerets; E. Female spinnerets. Abbreviations: B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8283777" httpUri="https://zenodo.org/record/8283777/files/figure.png" pageId="19" pageNumber="20">Figs 12A–B</figureCitation>
|
||
,
|
||
<figureCitation id="26DC2A4D8660FF9A5835F827DA1A27D8" box="[516,599,1996,2022]" captionStart="FIGURE 13" captionStartId="19.[151,250,1565,1589]" captionTargetBox="[151,1415,182,1520]" captionTargetId="figure-17@19.[151,1435,181,1542]" captionTargetPageId="19" captionText="FIGURE 13. Copulatory organs of Beatitas octomaculatus sp. nov.: A. Male left palp, ventral view; B. Same, retrolateral view; C. Same, dorsal view; D. Epigyne, ventral view; E. Same, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8283781" httpUri="https://zenodo.org/record/8283781/files/figure.png" pageId="19" pageNumber="20">13D–E</figureCitation>
|
||
. Epigynal plate weakly sclerotized, posterior edge nearly straight. Atrium diamond-shaped. Median septum thin, Y-shaped. Copulatory openings large, horn-shaped, located at lateral margin of atrium. Copulatory ducts short and thick, straight and connected with oval bursa. Glandular appendages absent. Connecting tube long, thinner than copulatory ducts, nearly surrounding entire spermathecae. Bursa oval, at anterior of the plate. Spermathecae globular, slightly separated. Fertilization ducts located anterior margin of spermathecae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F6FD65438667FF9D5AF6FEA1DB3A215B" box="[199,887,330,357]" pageId="20" pageNumber="21" type="distribution">
|
||
<paragraph id="BE5836C88667FF9D5AF6FEA1DB3A215B" blockId="20.[151,1437,151,357]" box="[199,887,330,357]" pageId="20" pageNumber="21">
|
||
<emphasis id="8C93EADA8667FF9D5AF6FEA1D92C215A" bold="true" box="[199,353,330,356]" pageId="20" pageNumber="21">Distribution.</emphasis>
|
||
<collectingCountry id="C6F076588667FF9D5B56FEA0D9E7215B" box="[359,426,331,357]" name="China" pageId="20" pageNumber="21">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="7C23F82A8667FF9D5B88FEA0DA35215B" box="[441,632,331,357]" country="China" name="Guangxi" pageId="20" pageNumber="21">Guangxi Zhuang</collectingRegion>
|
||
Autonomous Region).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |