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<document id="4920BA6CEDD79D6A79A0B7BD0FD871E9" ID-DOI="10.11646/zootaxa.1170.1.2" ID-ISSN="1175­5334" ID-Zenodo-Dep="5058686" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625191004336" checkinUser="felipe" docAuthor="Shaverdo, Helena V." docDate="2006" docId="1C71EF581F4C847AFEE3F9E93F9CFCA8" docLanguage="en" docName="zt01170p056.pdf" docOrigin="Zootaxa 1170 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1170.1.2" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Hydroporus simplex Gordon. During 1981" docType="treatment" docVersion="3" lastPageNumber="55" masterDocId="E04897201F578466FFEBFFC63D63FFCE" masterDocTitle="Revision of the longiusculus-group of the genus Hydroporus Clairville, 1806 (Coleoptera: Dytiscidae)" masterLastPageNumber="56" masterPageNumber="27" pageNumber="54" updateTime="1698983326455" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="95348A6CE33C51D49DA123970CCA7B1F">Revision of the longiusculus-group of the genus Hydroporus Clairville, 1806 (Coleoptera: Dytiscidae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="DD233834FEC6F056BFFBA1F6F8C26D39">Shaverdo, Helena V.</mods:namePart>
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<mods:date id="938E09F2F246FAED83513371EE3DAE2B">2006</mods:date>
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<mods:number id="0B14FCB8A10A57D1F7F007AB35F790AF">2006-04-10</mods:number>
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<mods:number id="86564C42FE0D9E11269847D5493C97B9">1170</mods:number>
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</mods:location>
<mods:classification id="B881D6ED107F5661E517B6C056AE661A">journal article</mods:classification>
<mods:identifier id="81C9A93D86AA5D0975DBD9A0B7D6FDC1" type="DOI">10.11646/zootaxa.1170.1.2</mods:identifier>
<mods:identifier id="3B525D05D81ADA29BD0B769333AB2A3F" type="ISSN">1175­5334</mods:identifier>
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</mods:mods>
<treatment id="1C71EF581F4C847AFEE3F9E93F9CFCA8" LSID="urn:lsid:plazi:treatment:1C71EF581F4C847AFEE3F9E93F9CFCA8" httpUri="http://treatment.plazi.org/id/1C71EF581F4C847AFEE3F9E93F9CFCA8" lastPageId="28" lastPageNumber="55" pageId="27" pageNumber="54">
<subSubSection id="DCC20DC51F4C847DFEE3F9E93F06F987" box="[264,613,1583,1609]" pageId="27" pageNumber="54" type="nomenclature">
<paragraph id="94675E4E1F4C847DFEE3F9E93F06F987" blockId="27.[264,613,1583,1609]" box="[264,613,1583,1609]" pageId="27" pageNumber="54">
<heading id="CF2FE9221F4C847DFEE3F9E93F06F987" bold="true" box="[264,613,1583,1609]" fontSize="11" level="1" pageId="27" pageNumber="54" reason="1">
<taxonomicName id="53D825CD1F4C847DFEE3F9E93F06F987" ID-CoL="3N8XM" authority="Gordon" authorityName="Gordon. During" authorityYear="1981" box="[264,613,1583,1609]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="54" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A6AC825C1F4C847DFEE3F9E93F06F987" bold="true" box="[264,613,1583,1609]" pageId="27" pageNumber="54">
<emphasis id="A6AC825C1F4C847DFEE3F9E93C9EF987" bold="true" box="[264,509,1583,1609]" italics="true" pageId="27" pageNumber="54">Hydroporus simplex</emphasis>
Gordon
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DCC20DC51F4C847DFEE3F9A93C8BF968" pageId="27" pageNumber="54" type="reference_group">
<paragraph id="94675E4E1F4C847DFEE3F9A93C8BF968" blockId="27.[264,1322,1647,1702]" pageId="27" pageNumber="54">
<treatmentCitationGroup id="B4C879601F4C847DFEE3F9A93C8BF968" pageId="27" pageNumber="54">
<treatmentCitation id="1579785F1F4C847DFEE3F9A93FDCF948" author="Gordon, R. D." box="[264,703,1647,1670]" page="114" pageId="27" pageNumber="54" year="1981">
<taxonomicName id="53D825CD1F4C847DFEE3F9A93FDCF948" ID-CoL="3N8XM" authority="Gordon, 1981: 114" authorityName="Gordon. During" authorityPageNumber="114" authorityYear="1981" box="[264,703,1647,1670]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="54" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A6AC825C1F4C847DFEE3F9A93C80F948" box="[264,483,1647,1670]" italics="true" pageId="27" pageNumber="54">Hydroporus simplex</emphasis>
<bibRefCitation id="F04923BF1F4C847DFE05F9A93FDCF948" author="Gordon, R. D." box="[494,703,1647,1670]" pageId="27" pageNumber="54" pagination="105 - 123" refId="ref18345" refString="Gordon, R. D. (1981) New species of North American Hydroporus, niger-tenebrosus group (Coleoptera: Dytiscidae). Pan-Pacific Entomologist, 57 (1), 105 - 123." type="journal article" year="1981">Gordon, 1981: 114</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(orig. descr.);
<treatmentCitation id="1579785F1F4C847DFC8DF9A9390EF948" author="Larson, D. J. &amp; Alarie Y. &amp; Roughley, R. E." box="[870,1133,1647,1670]" page="379" pageId="27" pageNumber="54" year="2000">
<bibRefCitation id="F04923BF1F4C847DFC8DF9A9390EF948" author="Larson, D. J. &amp; Alarie Y. &amp; Roughley, R. E." box="[870,1133,1647,1670]" pageId="27" pageNumber="54" refId="ref18481" refString="Larson, D. J., Alarie Y., &amp; Roughley, R. E. (2000) Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, Ontario, Canada, 982 pp." type="book" year="2000">
Larson
<emphasis id="A6AC825C1F4C847DFC50F9A93E8DF948" box="[955,1006,1647,1670]" italics="true" pageId="27" pageNumber="54">et al</emphasis>
. 2000: 379
</bibRefCitation>
</treatmentCitation>
(descr.);
<treatmentCitation id="1579785F1F4C847DFB32F9A93CC0F968" author="Nilsson, A. N." page="159" pageId="27" pageNumber="54" year="2001">
<bibRefCitation id="F04923BF1F4C847DFB32F9A93CC0F968" author="Nilsson, A. N." pageId="27" pageNumber="54" refId="ref18675" refString="Nilsson, A. N. (2001) Dytiscidae. World Catalogue of Insects, Vol. 3. Apollo Books, Stenstrup, 395 pp." type="book" year="2001">Nilsson 2001: 159</bibRefCitation>
</treatmentCitation>
(cat.).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DCC20DC51F4C847AFEE3F90B3F9CFCA8" lastPageId="28" lastPageNumber="55" pageId="27" pageNumber="54" type="materials_examined">
<paragraph id="94675E4E1F4C847DFEE3F90B3CCAF928" blockId="27.[264,1323,1741,1886]" box="[264,425,1741,1766]" pageId="27" pageNumber="54">
<emphasis id="A6AC825C1F4C847DFEE3F90B3CCAF928" box="[264,425,1741,1766]" italics="true" pageId="27" pageNumber="54">
<typeStatus id="4B63E0EC1F4C847DFEE3F90B3C5CF928" box="[264,319,1741,1766]" pageId="27" pageNumber="54">Type</typeStatus>
material
</emphasis>
</paragraph>
<paragraph id="94675E4E1F4C847DFED3F9333E21F890" blockId="27.[264,1323,1741,1886]" pageId="27" pageNumber="54">
<taxonomicName id="53D825CD1F4C847DFED3F9333F4AF8C0" authorityName="Gordon. During" authorityYear="1981" box="[312,553,1781,1806]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="54" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A6AC825C1F4C847DFED3F9333F4AF8C0" box="[312,553,1781,1806]" italics="true" pageId="27" pageNumber="54">Hydroporus simplex</emphasis>
</taxonomicName>
:
<materialsCitation id="24B054131F4C847DFDD6F9323E5DF890" collectingDate="1948-07-13" collectionCode="CAS" collectorName="P. H. Arnaud" county="Tuolumne Co." location="Cal" municipality="Pinecrest" pageId="27" pageNumber="54" specimenCount="1" specimenCount-male="1" stateProvince="California" typeStatus="holotype">
<typeStatus id="4B63E0EC1F4C847DFDD6F9323FCCF8C0" box="[573,687,1780,1806]" pageId="27" pageNumber="54" type="holotype">Holotype</typeStatus>
:
<specimenCount id="82DE95C71F4C847DFD56F9313FACF8C9" box="[701,719,1783,1799]" pageId="27" pageNumber="54" type="male">♂</specimenCount>
&quot;
<collectingMunicipality id="7403C4341F4C847DFD08F9323E36F8C0" box="[739,853,1780,1806]" pageId="27" pageNumber="54">Pinecrest</collectingMunicipality>
,
<location id="910708951F4C847DFC8EF9323EEDF8C0" LSID="urn:lsid:plazi:treatment:1C71EF581F4C847AFEE3F9E93F9CFCA8:910708951F4C847DFC8EF9323EEDF8C0" box="[869,910,1780,1806]" county="Tuolumne Co." municipality="Pinecrest" name="Cal" pageId="27" pageNumber="54" stateProvince="California">Cal</location>
\
<collectingCounty id="7D0626C21F4C847DFC46F9323939F8C0" box="[941,1114,1780,1806]" pageId="27" pageNumber="54">Tuolumne Co.</collectingCounty>
\
<date id="E066788E1F4C847DFB92F9323874F8C0" box="[1145,1303,1780,1806]" pageId="27" pageNumber="54" value="1948-07-13">
<collectingDate id="F02281661F4C847DFB92F9323874F8C0" box="[1145,1303,1780,1806]" pageId="27" pageNumber="54" value="1948-07-13">13–VII–1948</collectingDate>
</date>
\
<collectorName id="392D3B981F4C847DFEE3F8DA3CC5F8F8" box="[264,422,1820,1846]" pageId="27" pageNumber="54">P.H. Arnaud</collectorName>
&quot;, &quot;
<typeStatus id="4B63E0EC1F4C847DFE27F8DA3F5EF8F8" box="[460,573,1820,1846]" pageId="27" pageNumber="54" type="holotype">Holotype</typeStatus>
\
<taxonomicName id="53D825CD1F4C847DFD8FF8DA3E3FF8F8" authorityName="Gordon. During" authorityYear="1981" box="[612,860,1820,1846]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="54" phylum="Arthropoda" rank="species" species="simplex">Hydroporus simplex</taxonomicName>
\ Robert Gordon&quot; [red], &quot;
<collectingRegion id="561C90AC1F4C847DFB5BF8DA3848F8F8" box="[1200,1323,1820,1846]" country="United States of America" name="California" pageId="27" pageNumber="54">California</collectingRegion>
Academy \ of Sciences \ Type No. 9839&quot; (
<collectionCode id="F2C9C68B1F4C847DFD1DF8823E56F890" box="[758,821,1860,1886]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="27" pageNumber="54" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="94675E4E1F4B847AFED3FEDA3966FEF8" blockId="28.[264,1324,284,870]" box="[312,1029,284,310]" pageId="28" pageNumber="55">
<materialsCitation id="24B054131F4B847AFED3FEDA3963FEF8" box="[312,1024,284,310]" country="United States of America" county="Tuolumne County" location="Pinecrest" pageId="28" pageNumber="55" specimenCount="1" stateProvince="California" typeStatus="holotype">
<typeStatus id="4B63E0EC1F4B847AFED3FEDA3C11FEF8" box="[312,370,284,310]" pageId="28" pageNumber="55">Type</typeStatus>
locality:
<collectingCountry id="ECCF1EDE1F4B847AFE0BFEDA3F7EFEF8" box="[480,541,284,310]" name="United States of America" pageId="28" pageNumber="55">USA</collectingCountry>
,
<collectingRegion id="561C90AC1F4B847AFDC1FEDA3FC7FEF8" box="[554,676,284,310]" country="United States of America" name="California" pageId="28" pageNumber="55">California</collectingRegion>
,
<collectingCounty id="7D0626C21F4B847AFD44FEDA3EE5FEF8" box="[687,902,284,310]" pageId="28" pageNumber="55">Tuolumne County</collectingCounty>
,
<location id="910708951F4B847AFC7AFEDA3963FEF8" LSID="urn:lsid:plazi:treatment:1C71EF581F4C847AFEE3F9E93F9CFCA8:910708951F4B847AFC7AFEDA3963FEF8" box="[913,1024,284,310]" country="United States of America" county="Tuolumne County" name="Pinecrest" pageId="28" pageNumber="55" stateProvince="California">Pinecrest</location>
</materialsCitation>
.
</paragraph>
<paragraph id="94675E4E1F4B847AFED3FE823F9CFCA8" blockId="28.[264,1324,284,870]" pageId="28" pageNumber="55">
In
<bibRefCitation id="F04923BF1F4B847AFEB2FE823F27FE90" author="Larson, D. J. &amp; Alarie Y. &amp; Roughley, R. E." box="[345,580,324,350]" pageId="28" pageNumber="55" refId="ref18481" refString="Larson, D. J., Alarie Y., &amp; Roughley, R. E. (2000) Predaceous Diving Beetles (Coleoptera: Dytiscidae) of the Nearctic Region, with emphasis on the fauna of Canada and Alaska. NRC Research Press, Ottawa, Ontario, Canada, 982 pp." type="book" year="2000">
Larson
<emphasis id="A6AC825C1F4B847AFE59FE823C84FE93" box="[434,487,324,349]" italics="true" pageId="28" pageNumber="55">et al</emphasis>
. (2000)
</bibRefCitation>
<taxonomicName id="53D825CD1F4B847AFDA6FE823FA8FE93" authorityName="Gordon. During" authorityYear="1981" box="[589,715,324,349]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A6AC825C1F4B847AFDA6FE823FA8FE93" box="[589,715,324,349]" italics="true" pageId="28" pageNumber="55">H. simplex</emphasis>
</taxonomicName>
was recognized as a member of the
<taxonomicName id="53D825CD1F4B847AFB6BFE823842FE93" authorityName="LeConte. Being" authorityYear="1852" box="[1152,1313,324,349]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="subpubescens">
<emphasis id="A6AC825C1F4B847AFB6BFE823842FE93" box="[1152,1313,324,349]" italics="true" pageId="28" pageNumber="55">subpubescens</emphasis>
</taxonomicName>
­ group. According to the results of the present work, this group now includes only one species,
<taxonomicName id="53D825CD1F4B847AFE85FE523F55FE63" authorityName="LeConte. Being" authorityYear="1852" box="[366,566,404,429]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="subpubescens">
<emphasis id="A6AC825C1F4B847AFE85FE523F55FE63" box="[366,566,404,429]" italics="true" pageId="28" pageNumber="55">H. subpubescens</emphasis>
</taxonomicName>
. This species is very similar to representatives of the
<emphasis id="A6AC825C1F4B847AFB2FFE523842FE63" box="[1220,1313,404,429]" italics="true" pageId="28" pageNumber="55">nigellus</emphasis>
­ group, especially to
<taxonomicName id="53D825CD1F4B847AFE13FE7A3FF8FE1B" authorityName="LeConte" authorityYear="1850" box="[504,667,444,469]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="tenebrosus">
<emphasis id="A6AC825C1F4B847AFE13FE7A3FF8FE1B" box="[504,667,444,469]" italics="true" pageId="28" pageNumber="55">H. tenebrosus</emphasis>
</taxonomicName>
; therefore, distinguishing of the
<taxonomicName id="53D825CD1F4B847AFBF4FE7A39A3FE1B" authorityName="LeConte. Being" authorityYear="1852" box="[1055,1216,444,469]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="subpubescens">
<emphasis id="A6AC825C1F4B847AFBF4FE7A39A3FE1B" box="[1055,1216,444,469]" italics="true" pageId="28" pageNumber="55">subpubescens</emphasis>
</taxonomicName>
­group is found unnecessary. The
<taxonomicName id="53D825CD1F4B847AFDA6FE223F8DFE33" authorityName="LeConte. Being" authorityYear="1852" box="[589,750,484,509]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="subpubescens">
<emphasis id="A6AC825C1F4B847AFDA6FE223F8DFE33" box="[589,750,484,509]" italics="true" pageId="28" pageNumber="55">subpubescens</emphasis>
</taxonomicName>
­group is omitted, and
<taxonomicName id="53D825CD1F4B847AFBC4FE223861FE33" authorityName="LeConte. Being" authorityYear="1852" box="[1071,1282,484,509]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="subpubescens">
<emphasis id="A6AC825C1F4B847AFBC4FE223861FE33" box="[1071,1282,484,509]" italics="true" pageId="28" pageNumber="55">H. subpubescens</emphasis>
</taxonomicName>
is transferred to the
<emphasis id="A6AC825C1F4B847AFE0AFDCA3F5DFDEB" box="[481,574,524,549]" italics="true" pageId="28" pageNumber="55">nigellus</emphasis>
­group.
<bibRefCitation id="F04923BF1F4B847AFD75FDCA3E33FDE8" author="Nilsson, A. N." box="[670,848,524,550]" pageId="28" pageNumber="55" refId="ref18675" refString="Nilsson, A. N. (2001) Dytiscidae. World Catalogue of Insects, Vol. 3. Apollo Books, Stenstrup, 395 pp." type="book" year="2001">Nilsson (2001)</bibRefCitation>
placed
<taxonomicName id="53D825CD1F4B847AFC59FDCA3951FDEB" authorityName="Gordon. During" authorityYear="1981" box="[946,1074,524,549]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A6AC825C1F4B847AFC59FDCA3951FDEB" box="[946,1074,524,549]" italics="true" pageId="28" pageNumber="55">H. simplex</emphasis>
</taxonomicName>
in the
<emphasis id="A6AC825C1F4B847AFB67FDCA3C2FFD80" italics="true" pageId="28" pageNumber="55">longiusculusgroup</emphasis>
probably because of its slightly angulate metacoxal processes. As shown above, this character is variable in the representatives of the group and alone cannot support placing
<taxonomicName id="53D825CD1F4B847AFEE3FD423CE6FD53" authorityName="Gordon. During" authorityYear="1981" box="[264,389,644,669]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A6AC825C1F4B847AFEE3FD423CE6FD53" box="[264,389,644,669]" italics="true" pageId="28" pageNumber="55">H. simplex</emphasis>
</taxonomicName>
in the
<emphasis id="A6AC825C1F4B847AFE3EFD423F10FD53" box="[469,627,644,669]" italics="true" pageId="28" pageNumber="55">longiusculus­</emphasis>
group. Based on my examination of this species I think that it is rather similar to species of the
<emphasis id="A6AC825C1F4B847AFD4BFD6A3F9EFD0B" box="[672,765,684,709]" italics="true" pageId="28" pageNumber="55">nigellus</emphasis>
­group, especially more similar to
<taxonomicName id="53D825CD1F4B847AFB65FD6A3840FD0B" authorityName="Sharp" authorityYear="1882" box="[1166,1315,684,709]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="despectus">
<emphasis id="A6AC825C1F4B847AFB65FD6A3840FD0B" box="[1166,1315,684,709]" italics="true" pageId="28" pageNumber="55">H. despectus</emphasis>
</taxonomicName>
, than to
<taxonomicName id="53D825CD1F4B847AFEB4FD123F74FD23" box="[351,535,724,749]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="longiusculus">
<emphasis id="A6AC825C1F4B847AFEB4FD123F74FD23" box="[351,535,724,749]" italics="true" pageId="28" pageNumber="55">H. longiusculus</emphasis>
</taxonomicName>
and
<taxonomicName id="53D825CD1F4B847AFDA4FD123F8FFD23" authorityName="Fall" authorityYear="1923" box="[591,748,724,749]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="pervicinus">
<emphasis id="A6AC825C1F4B847AFDA4FD123F8FFD23" box="[591,748,724,749]" italics="true" pageId="28" pageNumber="55">H. pervicinus</emphasis>
</taxonomicName>
. It shares with them a single puncture row on the anterior surface of the protibia, the non­produced posterolateral angles of the pronotum, and the median lobe of the aedeagus sinuate in lateral view. Therefore, it is suggested that
<taxonomicName id="53D825CD1F4B847AFEE3FC8A3CE6FCAB" authorityName="Gordon. During" authorityYear="1981" box="[264,389,844,869]" class="Insecta" family="Dytiscidae" genus="Hydroporus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="55" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="A6AC825C1F4B847AFEE3FC8A3CE6FCAB" box="[264,389,844,869]" italics="true" pageId="28" pageNumber="55">H. simplex</emphasis>
</taxonomicName>
be placed in the
<emphasis id="A6AC825C1F4B847AFDA6FC8A3FC9FCAB" box="[589,682,844,869]" italics="true" pageId="28" pageNumber="55">nigellus</emphasis>
­group.
</paragraph>
</subSubSection>
</treatment>
</document>
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