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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.218.91199" ID-Pensoft-Pub="1314-2003-218-117" ID-Pensoft-UUID="ED39B229D5EF5C2A9DF976BB34698535" ModsDocID="1314-2003-218-117" checkinTime="1673667842713" checkinUser="pensoft" docAuthor="Wang, Long-Yuan, Zhao, Wan-Yi, Chen, Zai-Xiong, Huang, Wei-Cheng, Ding, Ming-Yan, Luo, Jin-Chu, Liao, Wen-Bo, Guo, Wei &amp; Fan, Qiang" docDate="2023" docId="1BDA1A62789257D7B50A8B9C95A17CE7" docLanguage="en" docName="PhytoKeys 218: 117-126" docOrigin="PhytoKeys 218" docPubDate="2023-01-13" docSource="http://dx.doi.org/10.3897/phytokeys.218.91199" docTitle="Commelina danxiaensis Q. Fan, Long Y. Wang &amp; W. Guo 2023, sp. nov." docType="treatment" docVersion="2" id="ED39B229D5EF5C2A9DF976BB34698535" lastPageNumber="117" masterDocId="ED39B229D5EF5C2A9DF976BB34698535" masterDocTitle="Commelina danxiaensis (Commelinaceae), a new species from Guangdong, China" masterLastPageNumber="126" masterPageNumber="117" pageNumber="117" updateTime="1673668144256" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Commelina danxiaensis (Commelinaceae), a new species from Guangdong, China</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wang, Long-Yuan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-2845-8171</mods:nameIdentifier>
<mods:affiliation>Department of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering, Guangzhou, Guangdong, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zhao, Wan-Yi</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocontrol and Guangdong Provincial Key Laboratory of Plant Resources, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Chen, Zai-Xiong</mods:namePart>
<mods:affiliation>Administrative Commission of Danxiashan National Park, Shaoguan, Guangdong, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Huang, Wei-Cheng</mods:namePart>
<mods:affiliation>Department of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering, Guangzhou, Guangdong, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ding, Ming-Yan</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1852-8167</mods:nameIdentifier>
<mods:affiliation>Shunde Polytechnic, Foshan, Guangdong, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Luo, Jin-Chu</mods:namePart>
<mods:affiliation>Department of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering, Guangzhou, Guangdong, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liao, Wen-Bo</mods:namePart>
<mods:affiliation>State Key Laboratory of Biocontrol and Guangdong Provincial Key Laboratory of Plant Resources, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong, China</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Guo, Wei</mods:namePart>
<mods:affiliation>Department of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering, Guangzhou, Guangdong, China</mods:affiliation>
<mods:nameIdentifier type="email">gwei717@163.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fan, Qiang</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4254-6936</mods:nameIdentifier>
<mods:affiliation>State Key Laboratory of Biocontrol and Guangdong Provincial Key Laboratory of Plant Resources, School of Life Sciences, Sun Yat-sen University, Guangzhou, Guangdong, China</mods:affiliation>
<mods:nameIdentifier type="email">fanqiang@mail.sysu.edu.cn</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>218</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>117</mods:start>
<mods:end>126</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/phytokeys.218.91199</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.218.91199</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-218-117</mods:identifier>
<mods:identifier type="Pensoft-UUID">ED39B229D5EF5C2A9DF976BB34698535</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:1BDA1A62789257D7B50A8B9C95A17CE7" httpUri="http://treatment.plazi.org/id/1BDA1A62789257D7B50A8B9C95A17CE7" lastPageNumber="117" pageId="0" pageNumber="117">
<subSubSection pageId="0" pageNumber="117" type="nomenclature">
<paragraph pageId="0" pageNumber="117">
<taxonomicName LSID="1BDA1A62-7892-57D7-B50A-8B9C95A17CE7" authority="Q. Fan, Long Y. Wang &amp; W. Guo" authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis" status="sp. nov.">Commelina danxiaensis Q.Fan, Long Y.Wang &amp; W.Guo</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="117">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="description">
<paragraph pageId="0" pageNumber="117">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Commelina danxiaensis sp. nov. A habitat B involucral bract C inflorescence D flower E fertile, sterile stamens and pistil F fruit in transverse section G seed." figureDoi="10.3897/phytokeys.218.91199.figure3" httpUri="https://binary.pensoft.net/fig/794736" pageId="0" pageNumber="117">Figs 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Commelina danxiaensis sp. nov. A habitat, growing on top of the Danxia landform B habit, plants ascending C-E leaf-blade, lanceolate to ovate-lanceolate F leaf-sheaths, with pubescence G-H flower I inflorescence J spathe K fruit in transverse section, 2 seeds per valve L seed. Scale bars: 1 cm (D, E, G, I, J); 3 mm (K, L)." figureDoi="10.3897/phytokeys.218.91199.figure4" httpUri="https://binary.pensoft.net/fig/794737" pageId="0" pageNumber="117">
, 4 Chinese name.
<normalizedToken originalValue="丹霞鸭跖草">丹霞鸭跖草</normalizedToken>
[
<normalizedToken originalValue="dān">dan</normalizedToken>
<normalizedToken originalValue="xiá">xia</normalizedToken>
<normalizedToken originalValue="yā">ya</normalizedToken>
<normalizedToken originalValue="zhǐ">zhǐ</normalizedToken>
<normalizedToken originalValue="cǎo">cǎo</normalizedToken>
]
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="materials_examined">
<paragraph pageId="0" pageNumber="117">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="117">
<materialsCitation collectingDate="2020-05-23" collectorName="Q. Fan" country="China" county="Renhua County" elevation="350" latitude="24.941591" location="Mount Danxia" longLatPrecision="1" longitude="113.7561" municipality="Shaoguan City" specimenCount="1" stateProvince="Guangdong" typeStatus="holotype">
<collectingCountry name="China">China</collectingCountry>
.
<collectingRegion country="China" name="Guangdong">Guangdong</collectingRegion>
:
<collectingMunicipality>Shaoguan City</collectingMunicipality>
,
<collectingCounty>Renhua County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:1BDA1A62789257D7B50A8B9C95A17CE7:BF36B21D897E7A91BDAB982417707E08" country="China" county="Renhua County" latitude="24.941591" longLatPrecision="1" longitude="113.7561" municipality="Shaoguan City" name="Mount Danxia" stateProvince="Guangdong">Mount Danxia</location>
,
<geoCoordinate degrees="24" direction="north" minutes="56" orientation="latitude" precision="1" seconds="29.73" value="24.941591">24°56'29.73&quot;N</geoCoordinate>
,
<geoCoordinate degrees="113" direction="east" minutes="45" orientation="longitude" precision="1" seconds="21.96" value="113.7561">113°45'21.96&quot;E</geoCoordinate>
, dry mountaintop, ca.
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</elevation>
</quantity>
elev.,
<collectingDate value="2020-05-23">23 May 2020</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="117">
<collectorName>Q. Fan</collectorName>
17910
</emphasis>
(
<typeStatus>holotype</typeStatus>
: SYS!; isotypes: SYS!, IBSC!)
</materialsCitation>
.
</paragraph>
<caption doi="10.3897/phytokeys.218.91199.figure3" httpUri="https://binary.pensoft.net/fig/794736" pageId="0" pageNumber="117" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Figure 3.</emphasis>
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="117">A</emphasis>
habitat
<emphasis bold="true" pageId="0" pageNumber="117">B</emphasis>
involucral bract
<emphasis bold="true" pageId="0" pageNumber="117">C</emphasis>
inflorescence
<emphasis bold="true" pageId="0" pageNumber="117">D</emphasis>
flower
<emphasis bold="true" pageId="0" pageNumber="117">E</emphasis>
fertile, sterile stamens and pistil
<emphasis bold="true" pageId="0" pageNumber="117">F</emphasis>
fruit in transverse section
<emphasis bold="true" pageId="0" pageNumber="117">G</emphasis>
seed.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="diagnosis">
<paragraph pageId="0" pageNumber="117">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
is morphologically similar to
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. communis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="communis">
<emphasis italics="true" pageId="0" pageNumber="117">C. communis</emphasis>
</taxonomicName>
due to their diffuse stems, lanceolate leaf-blades and approximate floral forms. However, it differs in its ascending stems (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Commelina danxiaensis sp. nov. A habitat, growing on top of the Danxia landform B habit, plants ascending C-E leaf-blade, lanceolate to ovate-lanceolate F leaf-sheaths, with pubescence G-H flower I inflorescence J spathe K fruit in transverse section, 2 seeds per valve L seed. Scale bars: 1 cm (D, E, G, I, J); 3 mm (K, L)." figureDoi="10.3897/phytokeys.218.91199.figure4" httpUri="https://binary.pensoft.net/fig/794737" pageId="0" pageNumber="117">4B</figureCitation>
), tomentose leaf-blades and sheaths (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Commelina danxiaensis sp. nov. A habitat, growing on top of the Danxia landform B habit, plants ascending C-E leaf-blade, lanceolate to ovate-lanceolate F leaf-sheaths, with pubescence G-H flower I inflorescence J spathe K fruit in transverse section, 2 seeds per valve L seed. Scale bars: 1 cm (D, E, G, I, J); 3 mm (K, L)." figureDoi="10.3897/phytokeys.218.91199.figure4" httpUri="https://binary.pensoft.net/fig/794737" pageId="0" pageNumber="117">4C-F</figureCitation>
), and sky-blue lateral petals with a white basal third (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Commelina danxiaensis sp. nov. A habitat, growing on top of the Danxia landform B habit, plants ascending C-E leaf-blade, lanceolate to ovate-lanceolate F leaf-sheaths, with pubescence G-H flower I inflorescence J spathe K fruit in transverse section, 2 seeds per valve L seed. Scale bars: 1 cm (D, E, G, I, J); 3 mm (K, L)." figureDoi="10.3897/phytokeys.218.91199.figure4" httpUri="https://binary.pensoft.net/fig/794737" pageId="0" pageNumber="117">4G</figureCitation>
).
</paragraph>
<caption doi="10.3897/phytokeys.218.91199.figure4" httpUri="https://binary.pensoft.net/fig/794737" pageId="0" pageNumber="117" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Figure 4.</emphasis>
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="117">A</emphasis>
habitat, growing on top of the Danxia landform
<emphasis bold="true" pageId="0" pageNumber="117">B</emphasis>
habit, plants ascending
<emphasis bold="true" pageId="0" pageNumber="117">C-E</emphasis>
leaf-blade, lanceolate to ovate-lanceolate
<emphasis bold="true" pageId="0" pageNumber="117">F</emphasis>
leaf-sheaths, with pubescence
<emphasis bold="true" pageId="0" pageNumber="117">G-H</emphasis>
flower
<emphasis bold="true" pageId="0" pageNumber="117">I</emphasis>
inflorescence
<emphasis bold="true" pageId="0" pageNumber="117">J</emphasis>
spathe
<emphasis bold="true" pageId="0" pageNumber="117">K</emphasis>
fruit in transverse section, 2 seeds per valve
<emphasis bold="true" pageId="0" pageNumber="117">L</emphasis>
seed. Scale bars: 1 cm (
<emphasis bold="true" pageId="0" pageNumber="117">D, E, G, I, J</emphasis>
); 3 mm (
<emphasis bold="true" pageId="0" pageNumber="117">K, L</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="description">
<paragraph pageId="0" pageNumber="117">Description.</paragraph>
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Herbs</emphasis>
over 30 cm tall, annual.
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Roots</emphasis>
thin, fibrous.
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Stems</emphasis>
ascending, many-branched; young shoots tomentose, becoming glabrous when old, hairs hyaline.
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Leaves</emphasis>
spirally-alternate, evenly distributed along the stems; sheaths tomentose, hairs hyaline, margin ciliate, hairs setose, white; blades 3-9
<normalizedToken originalValue="×">x</normalizedToken>
1.5-2.0 cm, lanceolate to ovate-lanceolate, tomentose on both sides, evently, hairs hyaline, base obtuse, margin glabrous, green to light-purple,, apex lanceolate.
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Inflorescences</emphasis>
leaf-opposite, restricted to the apex of the stems; peduncle 1.5-4.0 cm long, tomentose, hairs hyaline; spathe 1.2-2.5
<normalizedToken originalValue="×">x</normalizedToken>
1.0-2.2 cm, cordate, 6-8 veined, adaxially tomentose, long pilose hairs sometimes along the veins, hairs hyaline, abaxially glabrous, base cordate, free, margin green, apex acute; upper cincinnus developed, 1-flowered, flower staminate, peduncle ca. 8 mm long, exerted, sparsely tomentose to nearly glabrous, hairs hyaline; lateral cincinnus 2-3-flowered, flowers bisexual, peduncle 5-8 mm long, nearly included, tomentose to nearly glabrous, hairs hyaline.
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Flowers</emphasis>
chasmogamous, bisexual or staminate, strongly zygomorphic; pedicels c. 3 mm long at anthesis, less than 6 mm long in fruit, reflected, nearly glabrous, hairs hyaline; sepals green, membranous, glabrous, dorsal sepal c. 5
<normalizedToken originalValue="×">x</normalizedToken>
2 mm, triangular-lanceolate, apex lanceolate, lateral sepals c. 5 mm
<normalizedToken originalValue="×">x</normalizedToken>
3 mm, widely oblique-elliptic, connate for 3.6-4.2 mm, apex obtuse to round; paired petals short-clawed, claw c. 2-3 mm long, limb c. 1 cm
<normalizedToken originalValue="×">x</normalizedToken>
1 cm, orbicular to sub-orbicular, sky-blue with a white basal third, medial petal c. 5
<normalizedToken originalValue="×">x</normalizedToken>
2 mm, narrowly rhombic, white, nearlly hyaline; staminodes 3, subequal, the medial slightly shorter, filaments c. 3 mm long, antherodes X-shaped, distinctly four-lobed, upper lobes c. 1/2 to the lower ones, yellow, with a dark maroon at centre; lateral stamen filaments ca. 1 cm long, curved, anthers 1.8-2.2
<normalizedToken originalValue="×">x</normalizedToken>
1.2-1.4 mm, elliptic; medial stamen filament ca. 5 mm long, anther 2.2-2.4
<normalizedToken originalValue="×">x</normalizedToken>
1.8-2.2 mm, saddle-shaped; ovary c. 1
<normalizedToken originalValue="×">x</normalizedToken>
1 mm, ovoid, glabrous, 2-locular, ovules 2 per locule, style c. 1.2 cm long, strongly curved at apex, stigma trilobate, white.
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Capsule</emphasis>
c. 5
<normalizedToken originalValue="×">x</normalizedToken>
3 mm, ellipsoid, glabrous, equally 2-valved.
<emphasis bold="true" italics="true" pageId="0" pageNumber="117">Seeds</emphasis>
2 per valve, 3-4
<normalizedToken originalValue="×">x</normalizedToken>
2-2.5 mm, elliptic, ventrally flattened, testa brown, irregularly pitted.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="phenology">
<paragraph pageId="0" pageNumber="117">Phenology.</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
flowers from April to July and fruits from June to September.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="distribution">
<paragraph pageId="0" pageNumber="117">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
is only known from the type locality, Mount Danxia, Renhua County, Guangdong Province, China. Only two populations have been found, with several hundred individuals. It grows on dry mountaintops of the Danxia formation at elevations of 100-350 m.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="conservation assessment">
<paragraph pageId="0" pageNumber="117">Conservation assessment.</paragraph>
<paragraph pageId="0" pageNumber="117">
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
has only been found in Mount Danxia within an area of less than 20 km2, making it putatively
<normalizedToken originalValue="Vulnerable">'Vulnerable'</normalizedToken>
(VU D2) according to the guidelines of the IUCN Red List Categories and Criteria (
<bibRefCitation author="Hong, DY" journalOrPublisher="24. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="117" publicationUrl="https://nc.iucnredlist.org/redlist/content/attachment_files/RedListGuidelines.pdf" refId="B8" refString="2022. . https://nc.iucnredlist.org/redlist/content/attachment_files/RedListGuidelines.pdf" url="https://nc.iucnredlist.org/redlist/content/attachment_files/RedListGuidelines.pdf" year="2022">IUCN 2022</bibRefCitation>
). However, the threat risk seems low because it is not economically valuable, and the
<normalizedToken originalValue="areas">area's</normalizedToken>
conservation is good.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="materials_examined">
<paragraph pageId="0" pageNumber="117">Additional specimens examined</paragraph>
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">
(
<emphasis italics="true" pageId="0" pageNumber="117">
<typeStatus>paratypes</typeStatus>
</emphasis>
).
</emphasis>
<materialsCitation collectingDate="2020-07-03" collectingDateMax="2020-08-29" collectingDateMin="2020-07-03" collectorName="Q. Fan" country="China" county="Renhua County" location="Mount Danxia" municipality="Shaoguan City" specimenCount="1" stateProvince="Guangdong">
<collectingCountry name="China">China</collectingCountry>
.
<collectingRegion country="China" name="Guangdong">Guangdong</collectingRegion>
:
<collectingMunicipality>Shaoguan City</collectingMunicipality>
,
<collectingCounty>Renhua County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:1BDA1A62789257D7B50A8B9C95A17CE7:33C734A9CEB763D0A95AF3BCD41313A4" country="China" county="Renhua County" municipality="Shaoguan City" name="Mount Danxia" stateProvince="Guangdong">Mount Danxia</location>
, fl.,
<collectingDate value="2020-07-03">3 July 2020</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="117">
<collectorName>Q. Fan</collectorName>
18026
</emphasis>
(SYS); loc. cit., fr.,
<collectingDate value="2020-08-29">29 August 2020</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="117">
<collectorName>Q. Fan</collectorName>
18231
</emphasis>
(SYS); loc. cit., fr.,
<collectingDate value="2020-08-29">29 August 2020</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="117">
<collectorName>Q. Fan</collectorName>
18232
</emphasis>
(SYS)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="117" type="discussion">
<paragraph pageId="0" pageNumber="117">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="117">
The new species
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. communis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="communis">
<emphasis italics="true" pageId="0" pageNumber="117">C. communis</emphasis>
</taxonomicName>
and
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. diffusa" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="117">C. diffusa</emphasis>
</taxonomicName>
in some of its morphological characters, such as lanceolate leaf-blades, cordate spathes free at base, and blue paired petals. Morphological comparisons among them have been listed in Table
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison amongst Commelina danxiaensis, C. communis, C. diffusa and C. bicolor D. Q. Wang &amp; M. E. Cheng." httpUri="http://table.plazi.org/id/C0C09F9FECD2B731F0D50DFBC57E6BA3" pageId="0" pageNumber="117" tableUuid="C0C09F9FECD2B731F0D50DFBC57E6BA3">1</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="C0C09F9FECD2B731F0D50DFBC57E6BA3" httpUri="http://table.plazi.org/id/C0C09F9FECD2B731F0D50DFBC57E6BA3" pageId="0" pageNumber="117" start="Table 1" startId="T1">
<paragraph pageId="0" pageNumber="117">
<emphasis bold="true" pageId="0" pageNumber="117">Table 1.</emphasis>
Morphological comparison amongst
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. communis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="communis">
<emphasis italics="true" pageId="0" pageNumber="117">C. communis</emphasis>
</taxonomicName>
,
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. diffusa" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="117">C. diffusa</emphasis>
</taxonomicName>
and
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. bicolor" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="117">C. bicolor</emphasis>
</taxonomicName>
D.Q. Wang &amp; M.E. Cheng.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="117">
<table pageId="0" pageNumber="117">
<tr pageId="0" pageNumber="117">
<th colspan="1" pageId="0" pageNumber="117" rowspan="1">Character</th>
<th colspan="1" pageId="0" pageNumber="117" rowspan="1">
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">C. danxiaensis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="117" rowspan="1">
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. communis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="communis">
<emphasis italics="true" pageId="0" pageNumber="117">C. communis</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="117" rowspan="1">
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. diffusa" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="diffusa">
<emphasis italics="true" pageId="0" pageNumber="117">C. diffusa</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="117" rowspan="1">
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. bicolor" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="117">C. bicolor</emphasis>
</taxonomicName>
D.Q. Wang &amp; M.E. Cheng
</th>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Stem</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">ascending</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">creeping</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">creeping</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Phyllotaxy</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">spirally-alternate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">distichously-alternate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">distichously-alternate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Leaf-blades</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">tomentose</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">glabrous</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">glabrous or hispid</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Leaf-sheaths</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">tomentose</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">glabrous</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">hispid or hispid-ciliate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">glabrous</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Flower size</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">
2.5
<normalizedToken originalValue="×">x</normalizedToken>
2 cm
</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">
2
<normalizedToken originalValue="×">x</normalizedToken>
1.5 cm
</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">
2
<normalizedToken originalValue="×">x</normalizedToken>
1.5 cm
</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Lateral sepals connation</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">connate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">connate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">free</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Lateral petals colour</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">sky blue, basal third white</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">dark blue, sometimes lilac or white</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">light blue, sometimes lilac</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">upper most part deep-blue, basal part with claws (3 mm long) white</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Medial petal colour</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">white</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">white</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">light blue</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">white</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Medial petal claw</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">absent</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">present</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">absent</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Medial petal shape</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">narrowly rhombic</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">obtrullate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">rhombic</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">oribicular-ovata or broadly ovata</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Medial staminode development</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">developed</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">developed</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">not developed</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">developed</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Stigma colour</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">white</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">white</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">white</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">unknown</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Capsule valve number</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">equally 2-valved</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">equally 2-valved</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">unequally 2-valved</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">2-valved</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Seed number per valve</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">2-seeded</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">2-seeded</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">dorsal valved 1-seeded, ventral valve 2-seeded</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">2-seeded</td>
</tr>
<tr pageId="0" pageNumber="117">
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Seed testa ornamentation</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">irregularly pitted</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">irregularly pitted</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">Double-reticulate</td>
<td colspan="1" pageId="0" pageNumber="117" rowspan="1">irregularly foveolate</td>
</tr>
</table>
</paragraph>
<paragraph pageId="0" pageNumber="117">
Additionally, we found the name
<taxonomicName class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina bicolor" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina bicolor</emphasis>
</taxonomicName>
D.Q.Wang &amp; M.E.Cheng described by
<bibRefCitation author="Wang, DQ" journalOrPublisher="Journal of West Anhui University" pageId="0" pageNumber="117" pagination="67 - 73" refId="B17" refString="Wang, DQ, Qing, Z, Xie, J, Cheng, ME, Zhang, L, 2019. Four new species of flowering plants from Anhui. Journal of West Anhui University 35 (5): 67 - 73" title="Four new species of flowering plants from Anhui." volume="35" year="2019">Wang et al. (2019)</bibRefCitation>
, which is a later homonym of
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. bicolor" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="117">C. bicolor</emphasis>
</taxonomicName>
Poepp. ex
<bibRefCitation author="Kunth, K" journalOrPublisher="24. Science Press, Beijing and Missouri Botanical Garden Press, St. Louis" pageId="0" pageNumber="117" refId="B9" refString="Kunth, K, 1843. Enumeratio Plantarum Omnium Hucusque Cognitarum 4: 89." title="Enumeratio Plantarum Omnium Hucusque Cognitarum 4: 89." year="1843">Kunth (1843)</bibRefCitation>
. Therefore, it is illegitimate under Art. 53.1 of ICN (
<bibRefCitation DOI="https://doi.org/10.12705/Code.2018" author="Turland, NJ" journalOrPublisher="Nucleic Acids Research" pageId="0" pageNumber="117" refId="B16" refString="Turland, NJ, Wiersema, JH, Barrie, FR, Greuter, W, Hawksworth, DL, Herendeen, PS, Knapp, S, Kusber, WH, Li, D-Z, Marhold, K, May, TW, McNeill, J, Monro, AM, Prado, J, Price, MJ, Smith, GF [Eds], 2018. International Code of Nomenclature for algae, fungi and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten. https://doi.org/10.12705/Code.2018" title="International Code of Nomenclature for algae, fungi and plants (Shenzhen Code) adopted by the Nineteenth International Botanical Congress Shenzhen, China, July 2017. Regnum Vegetabile 159. Koeltz Botanical Books, Glashuetten." url="https://doi.org/10.12705/Code.2018" year="2018">Turland et al. 2018</bibRefCitation>
). This species is documented in Anhui and Hubei Provinces, and the type specimen (
<emphasis italics="true" pageId="0" pageNumber="117">D.Q. Wang, Y.Y. Lu &amp; L. Zhang W1406291</emphasis>
, PE) was collected at Daqian Shan of Feixi County in Anhui Province (
<bibRefCitation author="Wang, DQ" journalOrPublisher="Journal of West Anhui University" pageId="0" pageNumber="117" pagination="67 - 73" refId="B17" refString="Wang, DQ, Qing, Z, Xie, J, Cheng, ME, Zhang, L, 2019. Four new species of flowering plants from Anhui. Journal of West Anhui University 35 (5): 67 - 73" title="Four new species of flowering plants from Anhui." volume="35" year="2019">Wang et al. 2019</bibRefCitation>
). We previously believed the unknown species of
<taxonomicName class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina</emphasis>
</taxonomicName>
in Mount Danxia to be conspecific with
<taxonomicName class="Liliopsida" family="Commelinaceae" kingdom="Plantae" lsidName="C. bicolor" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="bicolor">
<emphasis italics="true" pageId="0" pageNumber="117">C. bicolor</emphasis>
</taxonomicName>
due to its similar morphological and ecological characteristics and treated it as a new distribution record for the Guangdong flora (
<bibRefCitation author="Dai, JM" journalOrPublisher="Subtropical Plant Science" pageId="0" pageNumber="117" pagination="495 - 499" refId="B1" refString="Dai, JM, Huang, YS, Sun, YY, Guo, JQ, Chen, ZX, Liao, WB, Fan, Q, 2021. New records of plants from Guangdong Province II. Subtropical Plant Science 50 (6): 495 - 499" title="New records of plants from Guangdong Province II." volume="50" year="2021">Dai et al. 2021</bibRefCitation>
). However, we have been unable to check the type material at PE, where the authors stated the holotype specimen was deposited, to confirm the
<normalizedToken originalValue="plants">plant's</normalizedToken>
identity and the
<normalizedToken originalValue="names">name's</normalizedToken>
application. We carried out fieldwork at Daqian Shan but were unable to find any individuals of this species. Hence, treatment has not been made for this species at the moment. It is planned that a further study will be carried out in the future.
</paragraph>
<paragraph pageId="0" pageNumber="117">
Up to the present time, four species of
<taxonomicName class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina</emphasis>
</taxonomicName>
have been documented in Danxiashan National Nature Reserve, including the new species
<taxonomicName authorityName="Q. Fan, Long Y. Wang &amp; W. Guo" authorityYear="2023" class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina danxiaensis" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="species" species="danxiaensis">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina danxiaensis</emphasis>
</taxonomicName>
. In order to facilitate identification, we here provide a key and checklist for the
<taxonomicName class="Liliopsida" family="Commelinaceae" genus="Commelina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Commelina" order="Commelinales" pageId="0" pageNumber="117" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="117">Commelina</emphasis>
</taxonomicName>
species in Mount Danxia.
</paragraph>
</subSubSection>
</treatment>
</document>