142 lines
19 KiB
XML
142 lines
19 KiB
XML
<document id="47B1344C635E4BFC4B6AA51384C0004C" ENCODING="UTF8" ID-GBIF-Dataset="9fcd647a-09f8-4ce9-ba68-778f173a6c1c" ModsDocID="22783" checkinTime="1249509757754" checkinUser="christiana" docAuthor="Hosoishi, S. & Ogata, K." docDate="2009" docId="1B2643154671162B18051F6C82FEE763" docLanguage="en" docName="22783.xml" docOrigin="Zootaxa 2062" docSource="http://hol.osu.edu/reference-full.html?id=22783" docTitle="Crematogaster (Physocrema) sewardi Forel" docType="treatment" docVersion="10" lastPageNumber="29" masterDocId="A92802EA8A918BF15D7133FD48DB5498" masterDocTitle="A taxonomic revision of the Asian endemic subgenus Physocrema of the genus Crematogaster (Hymenoptera: Formicidae)." masterLastPageNumber="36" masterPageNumber="15" pageNumber="28" updateTime="1694988214208" updateUser="admin">
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<mods:titleInfo id="FF99054C53F253CFB4FBB8C95C36ACD7">
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<mods:title id="F9169041997057DC8BEF5407CD1C0B1A">A taxonomic revision of the Asian endemic subgenus Physocrema of the genus Crematogaster (Hymenoptera: Formicidae).</mods:title>
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<mods:namePart id="BBEBB5A74E7DBF34A94A304C5E2DC92A">Hosoishi, S.</mods:namePart>
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<mods:namePart id="CC8C0D91D63A00813EF57948DA5FE83C">Ogata, K.</mods:namePart>
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<mods:typeOfResource id="CA63A5012C24BDDC4800C3D857E4032D">text</mods:typeOfResource>
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<mods:title id="70625623B3F334474346D59FB1C7599F">Zootaxa</mods:title>
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<mods:date id="EF3ECAA7BE0176E13038BE721E4F3DF8">2009</mods:date>
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<mods:detail id="C0E82498A91A2A42DC60800DF696E01C" type="volume">
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<mods:number id="4EC87DE372A693074B598114AE8BBB1E">2062</mods:number>
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<mods:start id="569D772829AE62CD0E9295C9174090C5">15</mods:start>
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<mods:url id="51FADCEB13A1A955C6CCC84A6CB61691">http://hol.osu.edu/reference-full.html?id=22783</mods:url>
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<mods:classification id="1DEE821876A2704FC3E78B5DC9A27F7C">journal article</mods:classification>
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<mods:identifier id="D52D0586306EE52F82582871F9A1F8B0" type="HNS-PUB">22783</mods:identifier>
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<treatment id="1B2643154671162B18051F6C82FEE763" ID-DOI="http://doi.org/10.5281/zenodo.6214302" ID-GBIF-Taxon="100122494" ID-Zenodo-Dep="6214302" LSID="urn:lsid:plazi:treatment:1B2643154671162B18051F6C82FEE763" httpUri="http://treatment.plazi.org/id/1B2643154671162B18051F6C82FEE763" lastPageNumber="29" pageNumber="28">
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<subSubSection id="25EE3156822BDB7A06BF8B61ECF75261" type="nomenclature">
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<paragraph id="B0D6F8C14963FB32F15DE3E62998226D" pageNumber="28">
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<taxonomicName id="70137A6936E3F8795E28A6C1D674822D" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:251853" authority="Forel" authorityName="Forel" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) sewardi" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Physocrema">Crematogaster (Physocrema) sewardi Forel</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="3BAA0CE87C61818EDEDEAD509136FBFE" type="description">
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<paragraph id="85D2D8B25947930AE870C03961240D19" pageNumber="28">(Figs 9, 17, 29)</paragraph>
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</subSubSection>
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<subSubSection id="732BBA8B641BEF912CA9537989DFD99D" type="materials_examined">
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<paragraph id="7DD19EE01B5C325E7C0E96075004DC72" pageNumber="28">
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<taxonomicName id="E5884A4F2B384C4A5F2F9DCC8B15A57B" authority="Forel, 1901: 64" authorityName="Forel" authorityPageNumber="64" authorityYear="1901" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subSpecies" species="deformis" subSpecies="sewardi">Crematogaster deformis r. sewardi Forel, 1901: 64</taxonomicName>
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. LECTOTYPE worker (top specimen of two on one pin) (by present designation) and one paralectotype worker and queen from MALAYSIA: Borneo (H. Seward) (
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<collectionCode id="5572524E335BA02A5F7741DFFE7D353F" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">
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<collectionCode id="712B8C07C235B0D82EA8075074D0E613">MHNG</collectionCode>
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</collectionCode>
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) [examined]. Combination in
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<taxonomicName id="FB59D65094BA6931FFD0F1676E33EED3" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:183849" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">
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<taxonomicName id="3608621B8ECF81A03424FCE3119566B0" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" subGenus="Physocrema">C. (Physocrema)</taxonomicName>
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</taxonomicName>
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by Emery, 1922: 140. Raised to species by Hosoishi & Ogata 2008: 7.
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</paragraph>
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</subSubSection>
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<subSubSection id="F104E8AF9A960F3B661E1DF23C8DD993" type="description">
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<paragraph id="A4B27DCAD8C5ACF8E2483DE625F413D2" pageNumber="28">Measurements and indices. HW 0.85-1.32; HL 0.77-1.22; CI 105-111; SL 0.70-0.96; SI 72-83; EL 0.16-0.24; PW 0.51-0.74; WL 0.96-1.40; PSL 0; PtL 0.28-0.42; PtW 0.21-0.36; PtH 0.15-0.22; PpL 0.19-0.28; PpW 0.23-0.34; PtHI 52-57; PtWI 74-86; PpWI 113-129; WI 94-116 (Three workers measured).</paragraph>
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<paragraph id="419695E3F2508439A06BA9F2EF404FE2" pageNumber="28">Description of worker. Workers with pronounced size polymorphism.</paragraph>
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<paragraph id="69F1DA7EC8E3DA0EDE882D9B94380AD8" pageNumber="28">Scape not reaching posterior corner of head in large workers.</paragraph>
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<paragraph id="6FAF9640601ABA9563B894A35E4DBB28" pageNumber="28">In lateral view, dorsal profile of promesonotum convex; propodeum distinctly raised relative to promesonotum. Anterior margin of pronotal collar slightly concave in dorsal view. Metanotal groove convex posteriorly in dorsal view in large workers, deep and situated posteriorly to the posterior ridge of pronotum in lateral view; thorax strongly constricted in middle in dorsal view. In dorsal view, some longitudinal rugulae connecting between mesonotum and propodeum; the boundary distinct. Propodeal spiracle situated away from the metapleural gland orifice, the distance between them much greater than the propodeal spiracle diameter. Propodeal spines absent.</paragraph>
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<paragraph id="F2410A36AB4E387C6FAC231034AF6482" pageNumber="28">Clypeus sculptured with longitudinal rugulae.</paragraph>
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<paragraph id="EB0DE5D69F29C670D74B525F56AF1017" pageNumber="28">Standing pilosity sparse on dorsal face of head, abundant on promesonotum. Dorsal face of head with decumbent setae. Clypeus with short setae. Fourth abdominal tergite with appressed setae. Dorsal setae on fourth abdominal tergite directed posteriorly.</paragraph>
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<paragraph id="F571F6AD24200F84BB5D88F1BD7F2E5C" pageNumber="28">Body color black.</paragraph>
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</subSubSection>
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<subSubSection id="DF5B64C1C6B2F389864EC0A6A23E273D" type="distribution">
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<paragraph id="CF7710AE42818C5B1E0451B3E55A9DC4" pageNumber="28">
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Distribution.
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<taxonomicName id="743EA1BE0666F75E72363750ECE3A7CA" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228741" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Paracrema">Crematogaster (P.) sewardi</taxonomicName>
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is the most wide-ranging species of the subgenus
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<taxonomicName id="088881993643E8C39F224819F2541577" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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(Fig. 29). This species ranges from Thailand to Malaysia (Peninsula and Borneo) and to Indonesia (Kalimantan, Sumatra, Krakatau Island, Java, Bali and Lombok).
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</paragraph>
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</subSubSection>
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<subSubSection id="0C76B9E8911DDAF2459D62B96B38E9C2" type="discussion">
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<paragraph id="BB1DAF7DDBE1A571A98233C96F4B78D2" pageNumber="28">
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Remarks. This species is very similar to
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<taxonomicName id="54DAC904C7F09A34385F5A8435B785F6" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ampullaris" subGenus="Physocrema">C. (P.) ampullaris</taxonomicName>
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, but can be distinguished by the dorsal outline of the mesosoma and the depth of the metanotal groove.
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</paragraph>
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<paragraph id="9090503C8A16C8848B4179478A652250" pageNumber="28">
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The
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<collectionCode id="31959D32A55640A6AEF117077538A9CA" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle">
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<collectionCode id="1E4DA7456900584106C56417DC838D19">MHNG</collectionCode>
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</collectionCode>
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syntypes were lighter colored than normal, but otherwise match the characters of the specimens examined.
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</paragraph>
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<paragraph id="75F3A70118F8E4E3E1B8C90CEDC7995C" lastPageNumber="29" pageNumber="28">
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Among the
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<taxonomicName id="89CBBD419768A57F7F060CCFB742B9C9" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="subGenus" subGenus="Physocrema">Physocrema</taxonomicName>
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species,
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<taxonomicName id="A5C8C00057949EAA4C973693ABDC649D" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ampullaris" subGenus="Physocrema">C. (P.) ampullaris</taxonomicName>
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,
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<taxonomicName id="5FCAAD12E7514CAD935377AB4EEA8A40" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="difformis" subGenus="Physocrema">C. (P.) difformis</taxonomicName>
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, and
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<taxonomicName id="650874056E505191A1512FEAAE996894" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228741" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Physocrema">C. (P.) sewardi</taxonomicName>
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are very similar to each other.
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<taxonomicName id="C88DF669B68812C729C23DF1A0E85E61" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228741" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Paracrema">Crematogaster (P.) sewardi</taxonomicName>
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is widely distributed in South East Asia, but almost all have been referred to as
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<taxonomicName id="E92EDB405C3D097A5CF97FE24AD19793" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="difformis" subGenus="Physocrema">C. (P.) difformis</taxonomicName>
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in the past. It is also likely that
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<taxonomicName id="F4123909C27FA067E2B33B163DE7692C" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228741" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Physocrema">C. (P.) sewardi</taxonomicName>
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has been misidentified as
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<taxonomicName id="527BEDB3A0114F75174651A5A13E9A70" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="difformis" subGenus="Physocrema">C. (P.) difformis</taxonomicName>
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in the literature (Attygalle et al. 1989; Gay & Hensen, 1992; Ito et al. 2001; Jaitrong & Nabhitabhata, 2005; Jones et al. 2005). Based on the key above,
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<taxonomicName id="1A48C5E470B3F667A7FD8B5B81A4AA54" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="difformis" subGenus="Physocrema">C. (P.) difformis</taxonomicName>
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can be easily distinguished from
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<taxonomicName id="34576DC84B20ECC7C56BCF03A6190C74" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ampullaris">ampullaris</taxonomicName>
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and
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<taxonomicName id="0C9576AF360174EF8D8A024E62EB25B8" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:149437" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi">sewardi</taxonomicName>
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, but as revealed above, the latter two species are very similar to each other and have been frequently confused.
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<taxonomicName id="5BD773A2F83E13D3D6B80C018E77A6D8" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="difformis" subGenus="Physocrema">Crematogaster (P.) difformis</taxonomicName>
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and
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<taxonomicName id="8B20C2871EA8995810BE7FE6A1994172" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228741" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Physocrema">C. (P.) sewardi</taxonomicName>
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are geographically sympartric in the Malay Peninsula and Borneo, whereas
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<taxonomicName id="E9B29C78BB9F0337F7DA0BF00A7955D8" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ampullaris" subGenus="Physocrema">C. (P.) ampullaris</taxonomicName>
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and
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<taxonomicName id="CF05C8592F8D22D5C06899107CC160AC" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228741" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Physocrema">C. (P.) sewardi</taxonomicName>
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are allopatric.
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<taxonomicName id="75AF3B98FCAD087DED0191FF6502EE9D" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="ampullaris" subGenus="Physocrema">Crematogaster (P.) ampullaris</taxonomicName>
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is found in Sulawesi, Timor and Philippines, and
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<taxonomicName id="825553BAF32C3DFE2DD16A069BDD4860" LSID-HNS="urn:lsid:biosci.ohio-state.edu:osuc_concepts:228741" class="Insecta" family="Formicidae" genus="Crematogaster" kingdom="Animalia" lsidName-HNS="Crematogaster (Physocrema) difformis sewardi Forel" order="Hymenoptera" pageNumber="28" phylum="Arthropoda" rank="species" species="sewardi" subGenus="Physocrema">C. (P.) sewardi</taxonomicName>
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in Indochina,
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<pageBreakToken id="8824868E9335B87B2B3AFBB9572247DE" pageNumber="29">Malay</pageBreakToken>
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Peninsula, Borneo, Sumatra, Krakatau Island, Java, Bali and Lombok. The slight morphological differences and allopatric distribution may imply that the two species are relatively new taxa.
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</paragraph>
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</subSubSection>
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<subSubSection id="5C232E0C4AF4179B9D6F044A67315CCB" type="materials_examined">
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<paragraph id="A556D19F19DA6BEC2B4765EEE295BD6E" pageNumber="29">
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Specimens examined. THAILAND: 6 workers, Sakaerat lowland forest (DEF), Nakornratchasima, 10. vii. 1999 (Sk. Yamane); 6 workers, Nam Tok Philio NP (300-500m alt.), Chanthaburi Prov., 22. xi. 2003 (Sk. Yamane); 4 workers, Khao Soi Dao, W. S., Chanthaburi Prov., 19. vii. 1997 (H. Okido); 4 workers, Khao Soi Dao, W. S., Chanthaburi Prov., 19. vii. 1997 (Sk. Yamane); 2 workers, Khao Chong (river side), Trang Prov., 24. viii. 1998 (Sk. Yamane) MALAYSIA: 11 workers, 7miles, Cameron Highland, 9. iii. 2005 (S. Hosoishi); 4 workers, 7miles, Cameron Highland, 12. iii. 2005 (S. Hosoishi); 3 workers, Poring, Kinabalu (HQ, 550m), Sabah, 17. iii. 1995 (Sk. Yamane); 2 workers, Kota Kinabalu, Sabah, 22. vii. 1996 (K. Eguchi). SINGAPORE: 3 workers, 4. xii. 1995 (Sk. Yamane); 2 workers, 23. vii. 1996 (Sk. Yamane); BRUNEI: 2 workers, Mixed Dipt. For.,
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<taxonomicName id="A424B9F53522505F90834E2852C548FC" class="Insecta" family="Formicidae" genus="Dryobalanops" kingdom="Animalia" order="Hymenoptera" phylum="Arthropoda" rank="species" species="beccarii">Dryobalanops beccarii</taxonomicName>
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, Site 7. 50m alt., Fog 11, Andalau, viii. 1991 (N. Mawdsley); INDONESIA: 3 workers, Kutai (Kal. Tim.), Kalimantan, 17. viii. 1986 (T. Yajima); 2 workers, Teluk Kabung nr Padang, Sumatra, 14. viii. 1985 (Sk. Yamane); 1 worker, Sako nr Tapan, Sumatra, 4-5. ix. 1985 (Sk. Yamane); 1 worker, Limau Manis nr Padang, Sumatra, 3. ix. 1985 (Sk. Yamane); 2 workers, P. Rakata Kecil, Krakatau Is., 31. vii. 1982 (Sk. Yamane); 2 workers, P. Peucang nr Ujung Kulon, 5. viii. 1982 (Sk. Yamane); 1 worker, P. Sertung, Krakatau Is., 30. vii. 1982 (Sk. Yamane); 1 worker, P. Sertung, Krakatau Is., 6. vii. 1982 (Sk. Yamane); 3 workers, Bogor, Java, 4-8. xi. 1985 (Sk. Yamane);3 workers, 1 male, Pulau Rakata, Krakatau Is., Sunda Strait, 1. i. 2007 (RK07-SKY-04) (Sk. Yamane);1 queen, Pulau Rakata, Krakatau Is., Sunda Strait, 1. i. 2007 (RK07-SKY-15) (Sk. Yamane); 1 worker, 1 queen, Pulau Sertung, Krakatau Is., Sunda Strait, 30. xii. 2006 (RK06-SKY-35) (Sk. Yamane); 2 workers, Pulau Sebesi, Lampung Prov., Sunda Strait, 12. viii. 2005 (Syaukani); 2 workers, P. Anak Krakatau, Krakatau Is., Sunda Strait, 17. viii. 2005 (Rk05-SKY-113) (Sk. Yamane); 2 workers, P. Anak Krakatau, Krakatau Is., Sunda Strait, 17. viii. 2005 (RK05-SKY-117) (Sk. Yamane); 3 workers, 1 male, Surisura, Pulau Siberut, Mentawai Is., 20. ii. 2007 (SU07-SKY-041) (Sk. Yamane); 2 workers, Carita, W. Java, 24. xii. 2006 (JV06-SKY-08) (Sk. Yamane); 2 workers, Bogor, Java, 9. xi. 1985 (Sk. Yamane); 1 worker, Campus of Gajah Mada Univ. (100m alt.), Yogyakarta, Java, 19. xii. 2002 (F. Yamane); 8 workers, Campus of Gajah Mada Univ. (100m alt.), Yogyakarta, Java, 29. xii. 2002 (F. Yamane); 6 workers, Dusun PK, Jelati Mendaya, Bali, 5-6. v. 1998 (Sk. Yamane); 20 workers, Senaru, Lombok, 30. ix. 2000 (H. Simbolon); 7 workers, Senaru, Lombok, 30. ix. 2000 (J. Abe); 4 workers, Senaru, Lombok, 30. ix. 2000 (K. Ogata); 2 workers, Kopi house nr Semaya, Lombok, 29. x. 1998 (K. Eguchi).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |