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<figureCitation id="08C570BA172DFFDFFD79BAC8FCADE185" box="[743,844,700,726]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="3.[189,232,1963,1989]" captionStartId-1="4.[189,232,1882,1908]" captionStartId-2="5.[189,232,1963,1989]" captionStartId-3="6.[189,232,1963,1989]" captionTargetBox-0="[189,1398,1260,1918]" captionTargetBox-1="[189,1398,275,1817]" captionTargetBox-2="[189,1398,815,1914]" captionTargetBox-3="[202,1384,1143,1906]" captionTargetId-0="figure-178@3.[189,1398,1260,1918]" captionTargetId-1="figure-14@4.[189,1398,275,1817]" captionTargetId-2="figure-112@5.[189,1398,808,1917]" captionTargetId-3="figure-324@6.[189,1398,1130,1918]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="Fig. 1. Luberotenthredo cerestensis gen. et sp. nov., holotype (PNRL 2716). Photograph of habitus. Scale bar = 2 mm." captionText-1="Fig. 2. Luberotenthredo cerestensis gen. et sp. nov., holotype (PNRL 2716). Photographs. A. Right wings. B. Left wings. Scale bars = 2 mm." captionText-2="Fig. 3. Luberotenthredo cerestensis gen. et sp. nov., holotype (PNRL 2716). Photographs. A. Head and thorax. B. Head and antenna. C. Reconstruction of antenna. D. Apex of abdomen. Scale bars = 1 mm." captionText-3="Fig. 4. Luberotenthredo cerestensis gen. et sp. nov., holotype (PNRL 2716). Reconstructions of the wing venations with names of cells and veins labeled. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.10453408" figureDoi-1="http://doi.org/10.5281/zenodo.10453410" figureDoi-2="http://doi.org/10.5281/zenodo.10453414" figureDoi-3="http://doi.org/10.5281/zenodo.10453418" httpUri-0="https://zenodo.org/record/10453408/files/figure.png" httpUri-1="https://zenodo.org/record/10453410/files/figure.png" httpUri-2="https://zenodo.org/record/10453414/files/figure.png" httpUri-3="https://zenodo.org/record/10453418/files/figure.png" pageId="3" pageNumber="4">Figs 14</figureCitation>
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</heading>
</paragraph>
<paragraph id="90416C3F172DFFDFFF23BB58FD25E015" blockId="3.[189,708,812,838]" box="[189,708,812,838]" pageId="3" pageNumber="4">As for the genus, by monotypy (vide supra).</paragraph>
</subSubSection>
<subSubSection id="D8E43FB4172DFFDFFF23BB19FC0CE0E5" box="[189,1005,877,950]" pageId="3" pageNumber="4" type="etymology">
<paragraph id="90416C3F172DFFDFFF23BB19FEA5E0D4" blockId="3.[189,324,877,903]" box="[189,324,877,903]" pageId="3" pageNumber="4">
<heading id="CB09DB53172DFFDFFF23BB19FEA5E0D4" bold="true" box="[189,324,877,903]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="A28AB02D172DFFDFFF23BB19FEA5E0D4" bold="true" box="[189,324,877,903]" pageId="3" pageNumber="4">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="90416C3F172DFFDFFF23BBEFFC0CE0E5" blockId="3.[189,1005,923,950]" box="[189,1005,923,950]" pageId="3" pageNumber="4">
Named after the small village of Céreste (Luberon, Southern
<collectingCountry id="E8E92CAF172DFFDFFC15BBE8FC01E0E5" box="[907,992,924,950]" name="France" pageId="3" pageNumber="4">France</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="D8E43FB4172DFFD9FF23BBA9FDD6E227" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="90416C3F172DFFDFFF23BBA9FE8DE0A4" blockId="3.[189,364,989,1015]" box="[189,364,989,1015]" pageId="3" pageNumber="4">
<heading id="CB09DB53172DFFDFFF23BBA9FE8DE0A4" bold="true" box="[189,364,989,1015]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="A28AB02D172DFFDFFF23BBA9FE8DE0A4" bold="true" box="[189,364,989,1015]" pageId="3" pageNumber="4">Type material</emphasis>
</heading>
</paragraph>
<materialsCitation id="20966662172DFFDFFF74BC78FE5EE7E0" collectionCode="PNRL" country="France" county="Musee de Geologie" location="Cereste" municipality="Calcaire de Campagne-Calavon' Formation" pageId="3" pageNumber="4" specimenCode="PNRL 2716" specimenCount="1" specimenCount-female="1" stateProvince="Alpes-de-Haute-Provence" typeStatus="holotype">
<paragraph id="90416C3F172DFFDFFF74BC78FEBDE775" blockId="3.[188,1398,1036,1203]" box="[234,348,1036,1062]" pageId="3" pageNumber="4">
<emphasis id="A28AB02D172DFFDFFF74BC78FEBDE775" bold="true" box="[234,348,1036,1062]" pageId="3" pageNumber="4">
<typeStatus id="4F45D29D172DFFDFFF74BC78FEBDE775" box="[234,348,1036,1062]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="90416C3F172DFFDFFF23BC5BFE5EE7E0" blockId="3.[188,1398,1036,1203]" pageId="3" pageNumber="4">
<collectingCountry id="E8E92CAF172DFFDFFF23BC5BFED0E71A" box="[189,305,1071,1097]" name="France" pageId="3" pageNumber="4">FRANCE</collectingCountry>
<specimenCount id="86F8A7B6172DFFDFFED1BC5BFE84E71A" box="[335,357,1071,1097]" count="1" pageId="3" pageNumber="4" type="female"></specimenCount>
(imprint of a complete specimen fossilized in dorsal view);
<collectingRegion id="523AA2DD172DFFDFFBA0BC5BFA93E71A" box="[1086,1394,1071,1097]" pageId="3" pageNumber="4">Alpes-de-Haute-Provence</collectingRegion>
, South of the village of
<location id="95213AE4172DFFDFFE67BC26FDB4E73F" LSID="urn:lsid:plazi:treatment:1857DD29172DFFDBFDA0BA01FEC9E7EB:95213AE4172DFFDFFE67BC26FDB4E73F" box="[505,597,1106,1132]" country="France" county="Musee de Geologie" municipality="Calcaire de Campagne-Calavon' Formation" name="Cereste" pageId="3" pageNumber="4" stateProvince="Alpes-de-Haute-Provence">Céreste</location>
; Early Oligocene,
<location id="95213AE4172DFFDFFCD1BC26FA93E73F" LSID="urn:lsid:plazi:treatment:1857DD29172DFFDBFDA0BA01FEC9E7EB:95213AE4172DFFDFFCD1BC26FA93E73F" box="[847,1394,1106,1132]" country="France" county="Musee de Geologie" municipality="Calcaire de Campagne-Calavon' Formation" name="Calcaire de Campagne-Calavon' Formation" pageId="3" pageNumber="4" stateProvince="Alpes-de-Haute-Provence">Calcaire de Campagne-Calavon Formation</location>
;
<specimenCode id="C058C444172DFFDFFF23BC02FEAAE7DC" box="[189,331,1141,1168]" collectionCode="PNRL" pageId="3" pageNumber="4">PNRL 2716</specimenCode>
, stored in the Musée de Géologie, Réserve géologique du Luberon, Parc naturel régional du Luberon, Apt, France.
</paragraph>
</materialsCitation>
<caption id="C4813CB7172DFFDFFF23BFDFFE75E4BA" ID-DOI="http://doi.org/10.5281/zenodo.10453408" ID-Zenodo-Dep="10453408" httpUri="https://zenodo.org/record/10453408/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1963,1989]" targetBox="[189,1398,1260,1918]" targetPageId="3">
<paragraph id="90416C3F172DFFDFFF23BFDFFE75E4BA" blockId="3.[189,1399,1963,2025]" pageId="3" pageNumber="4">
<emphasis id="A28AB02D172DFFDFFF23BFDFFEEEE496" bold="true" box="[189,271,1963,1989]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
<taxonomicName id="57FE17BC172DFFDFFE84BFDFFD89E496" authority="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault, 2024" authorityName="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault" authorityYear="2024" box="[282,616,1963,1989]" class="Insecta" family="Tenthredinidae" genus="Luberotenthredo" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cerestensis" status="gen. et sp. nov.">
<emphasis id="A28AB02D172DFFDFFE84BFDFFD89E496" box="[282,616,1963,1989]" italics="true" pageId="3" pageNumber="4">Luberotenthredo cerestensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9B90D56172DFFDFFDECBFD8FCCEE495" box="[626,815,1964,1990]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype (PNRL 2716). Photograph of habitus. Scale bar = 2 mm.
</paragraph>
</caption>
<caption id="C4813CB7172AFFD8FF23BF2EFD42E4CB" ID-DOI="http://doi.org/10.5281/zenodo.10453410" ID-Zenodo-Dep="10453410" httpUri="https://zenodo.org/record/10453410/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1882,1908]" targetBox="[189,1398,275,1817]" targetPageId="4">
<paragraph id="90416C3F172AFFD8FF23BF2EFD42E4CB" blockId="4.[189,1398,1882,1944]" pageId="4" pageNumber="5">
<emphasis id="A28AB02D172AFFD8FF23BF2EFEEEE427" bold="true" box="[189,271,1882,1908]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
<taxonomicName id="57FE17BC172AFFD8FE84BF2FFD88E427" authority="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault, 2024" authorityName="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault" authorityYear="2024" box="[282,617,1883,1909]" class="Insecta" family="Tenthredinidae" genus="Luberotenthredo" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cerestensis" status="gen. et sp. nov.">
<emphasis id="A28AB02D172AFFD8FE84BF2FFD88E427" box="[282,617,1883,1909]" italics="true" pageId="4" pageNumber="5">Luberotenthredo cerestensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9B90D56172AFFD8FDEDBF2FFCD0E426" box="[627,817,1883,1909]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype (PNRL 2716). Photographs.
<emphasis id="A28AB02D172AFFD8FA90BF2EFAC2E427" bold="true" box="[1294,1315,1882,1908]" pageId="4" pageNumber="5">A</emphasis>
. Right wings.
<emphasis id="A28AB02D172AFFD8FE8CBF0AFEC7E4CB" bold="true" box="[274,294,1918,1944]" pageId="4" pageNumber="5">B</emphasis>
. Left wings. Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="90416C3F172BFFD9FF23B97DFE1AE271" blockId="5.[189,507,264,291]" box="[189,507,264,291]" pageId="5" pageNumber="6">
<heading id="CB09DB53172BFFD9FF23B97DFE1AE271" bold="true" box="[189,507,264,291]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="A28AB02D172BFFD9FF23B97DFE1AE271" bold="true" box="[189,507,264,291]" pageId="5" pageNumber="6">Type locality and horizon</emphasis>
</heading>
</paragraph>
<paragraph id="90416C3F172BFFD9FF23B94CFDD6E227" blockId="5.[189,1399,311,372]" pageId="5" pageNumber="6">
<collectingCountry id="E8E92CAF172BFFD9FF23B94CFEF1E201" box="[189,272,312,338]" name="France" pageId="5" pageNumber="6">France</collectingCountry>
, Alpes-de-Haute-Provence, South of the village of Céreste, Early Oligocene, Calcaire de Campagne-Calavon Formation.
</paragraph>
</subSubSection>
<subSubSection id="D8E43FB4172BFFDBFF23B9EFFD13E12D" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="90416C3F172BFFD9FF23B9EFFEAFE2E6" blockId="5.[189,334,411,437]" box="[189,334,411,437]" pageId="5" pageNumber="6">
<heading id="CB09DB53172BFFD9FF23B9EFFEAFE2E6" bold="true" box="[189,334,411,437]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="A28AB02D172BFFD9FF23B9EFFEAFE2E6" bold="true" box="[189,334,411,437]" pageId="5" pageNumber="6">Description</emphasis>
</heading>
</paragraph>
<paragraph id="90416C3F172BFFD9FF23B9BFFD97E2B6" blockId="5.[189,630,458,485]" box="[189,630,458,485]" pageId="5" pageNumber="6">
<smallCapsWord id="96A7FAE3172BFFD9FF23B9BFFE8FE2B0" baselines="478,478" box="[189,366,459,485]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="5" pageNumber="6">MEASUREMENTS</smallCapsWord>
. Body length
<quantity id="5706C1DA172BFFD9FD8BB9BEFD93E2B6" box="[533,626,458,485]" metricMagnitude="-3" metricUnit="m" metricValue="8.7" pageId="5" pageNumber="6" unit="mm" value="8.7">8.7 mm</quantity>
.
</paragraph>
<paragraph id="90416C3F172BFFD9FF23BA78FD72E11A" blockId="5.[189,1398,524,585]" pageId="5" pageNumber="6">
<smallCapsWord id="96A7FAE3172BFFD9FF23BA78FEE8E177" baselines="543,543" box="[189,265,524,550]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Color" pageId="5" pageNumber="6">COLOR</smallCapsWord>
. Body brown with darker head; compound eyes dark brown; wings hyaline, pterostigma dark brown anteriorly and lighter posteriorly.
</paragraph>
<paragraph id="90416C3F172BFFDAFF23BA05FD88E215" blockId="5.[189,1400,624,721]" lastBlockId="6.[189,1399,265,327]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<smallCapsWord id="96A7FAE3172BFFD9FF23BA05FF1DE1DA" baselines="644,644" box="[189,252,625,651]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Head" pageId="5" pageNumber="6">HEAD</smallCapsWord>
. Deformed and poorly preserved,
<quantity id="5706C1DA172BFFD9FD07BA04FD15E1D8" box="[665,756,624,651]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="6" unit="mm" value="1.4">1.4 mm</quantity>
long, 2.0 mm wide; compound eyes large, occupying most of head lateral surface, deformed; mouthparts not preserved except left mandible with at least one apical and two preapical teeth; antennae very long, slightly longer than abdomen; scape and pedicel very short; seven flagellomeres; third antennomere as long as fourth; preapical flagellomeres cylindrical, elongate, slightly truncate obliquely.
</paragraph>
<caption id="C4813CB7172BFFD9FF23BFDFFA84E4BA" ID-DOI="http://doi.org/10.5281/zenodo.10453414" ID-Zenodo-Dep="10453414" httpUri="https://zenodo.org/record/10453414/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1963,1989]" targetBox="[189,1398,815,1914]" targetPageId="5">
<paragraph id="90416C3F172BFFD9FF23BFDFFA84E4BA" blockId="5.[189,1398,1963,2025]" pageId="5" pageNumber="6">
<emphasis id="A28AB02D172BFFD9FF23BFDFFEEAE496" bold="true" box="[189,267,1963,1989]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
<taxonomicName id="57FE17BC172BFFD9FE8DBFDFFDBFE496" authority="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault, 2024" authorityName="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault" authorityYear="2024" box="[275,606,1963,1989]" class="Insecta" family="Tenthredinidae" genus="Luberotenthredo" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cerestensis" status="gen. et sp. nov.">
<emphasis id="A28AB02D172BFFD9FE8DBFDFFDBFE496" box="[275,606,1963,1989]" italics="true" pageId="5" pageNumber="6">Luberotenthredo cerestensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9B90D56172BFFD9FDFBBFD8FCF9E495" box="[613,792,1964,1990]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype (PNRL 2716). Photographs.
<emphasis id="A28AB02D172BFFD9FB7DBFDFFB19E496" bold="true" box="[1251,1272,1963,1989]" pageId="5" pageNumber="6">A</emphasis>
. Head and thorax.
<emphasis id="A28AB02D172BFFD9FE89BFBAFECAE4BB" bold="true" box="[279,299,1998,2024]" pageId="5" pageNumber="6">B</emphasis>
. Head and antenna.
<emphasis id="A28AB02D172BFFD9FD84BFBAFDCEE4BB" bold="true" box="[538,559,1998,2024]" pageId="5" pageNumber="6">C</emphasis>
. Reconstruction of antenna.
<emphasis id="A28AB02D172BFFD9FC1EBFBAFC7CE4BB" bold="true" box="[896,925,1998,2024]" pageId="5" pageNumber="6">D.</emphasis>
Apex of abdomen. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="90416C3F1728FFDAFF23B91AFAE7E2F8" blockId="6.[189,1399,366,427]" pageId="6" pageNumber="7">
<smallCapsWord id="96A7FAE31728FFDAFF23B91AFEF8E2D5" baselines="385,385" box="[189,281,366,392]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Thorax" pageId="6" pageNumber="7">THORAX</smallCapsWord>
. ca 3.0 mm long,
<quantity id="5706C1DA1728FFDAFD9AB91AFD83E2DB" box="[516,610,366,392]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="7" unit="mm" value="2.8">2.8 mm</quantity>
wide, apparently smooth (deformed by compression); pronotum markedly constricted medially; propleuron short and head close to thorax; legs not preserved.
</paragraph>
<paragraph id="90416C3F1728FFDAFF23B9A7FD6BE0AF" blockId="6.[188,1401,466,1021]" pageId="6" pageNumber="7">
<smallCapsWord id="96A7FAE31728FFDAFF23B9A7FED1E2B8" baselines="486,486" box="[189,304,467,493]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Forewing" pageId="6" pageNumber="7">FOREWING</smallCapsWord>
. Complete,
<quantity id="5706C1DA1728FFDAFE58B9A6FDC5E2BE" box="[454,548,466,493]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="6" pageNumber="7" unit="mm" value="7.5">7.5 mm</quantity>
long,
<quantity id="5706C1DA1728FFDAFDEAB9A6FD33E2BE" box="[628,722,466,493]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="6" pageNumber="7" unit="mm" value="2.9">2.9 mm</quantity>
wide; short anterior Sc branch located slightly anteriad Sc+R and 1-M meeting point; pterostigma
<quantity id="5706C1DA1728FFDAFD29B982FCF3E143" box="[695,786,502,528]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="6" pageNumber="7" unit="mm" value="1.3">1.3 mm</quantity>
long,
<quantity id="5706C1DA1728FFDAFCC2B981FC56E143" box="[860,951,501,528]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="mm" value="0.5">0.5 mm</quantity>
wide, strongly arched along cell 2R1, slightly arched along anterior wing margin; M+Cu slightly curved; 1-M slightly longer than 1-Cu, nearly straight; vein R clearly deviated between junctions of 1-M and Sc; fusion of M with R very long, longer than half length of vein 1-M; vein 2-M meeting R more than halfway between short anterior Sc branch and pterostigma; vein 2-M slightly longer than 1rs-m; Rs emerging at pterostigmal base; 2-Rs nearly straight, longer than 3-Rs; r-rs crossvein located in pterostigma distalmost part, meeting Rs anteriad 3rsm, enclosing cell 2R1; 4-Rs and 5-Rs nearly aligned with 4-Rs extremely short (more than twice shorter than 3-Rs); cell 1R1 trapezoidal,
<quantity id="5706C1DA1728FFDAFDD6BA98FD45E055" box="[584,676,748,775]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="mm" value="0.5">0.5 mm</quantity>
long; cell 1Rs
<quantity id="5706C1DA1728FFDAFCC4BA98FC54E055" box="[858,949,748,774]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="mm" value="1.2">1.2 mm</quantity>
long, enclosed distally by subvertical 2rs-m (the latter lightly longer than 1rs-m); 3-M shorter than 4-M; cell 2Rs
<quantity id="5706C1DA1728FFDAFBC0BB7BFB5AE079" box="[1118,1211,783,810]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="7" unit="mm" value="0.9">0.9 mm</quantity>
long, distinctly wider along sinusoidal 3rs-m; 1-Cu and 2-Cu nearly straight, 1-Cu longer than 2-Cu; lm-cu meeting Cu at an angle of ca 110°, 1-M and 1m-cu subparallel and of similar lengths; cell 1Cu elongate, longer than cell R; 1cu-a (nervulus) located slightly distad middle of cell 1M; cell 2Cu short trapezoidal; cell 2M fully enclosed distally by long slightly sinusoidal crossvein 2m-cu; vein 1-A long, nearly straight; vein 2A+3-A complete, strongly sinuate in first half, fused with to 1-A on a short distance; cell 1A shorter than cell 2A, ca 0.8× length of cell 2A.
</paragraph>
<caption id="C4813CB71728FFDAFF23BFDFFC2EE4BA" ID-DOI="http://doi.org/10.5281/zenodo.10453418" ID-Zenodo-Dep="10453418" httpUri="https://zenodo.org/record/10453418/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1963,1989]" targetBox="[202,1384,1143,1906]" targetPageId="6">
<paragraph id="90416C3F1728FFDAFF23BFDFFC2EE4BA" blockId="6.[188,1399,1963,2025]" pageId="6" pageNumber="7">
<emphasis id="A28AB02D1728FFDAFF23BFDFFEEBE496" bold="true" box="[189,266,1963,1989]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
<taxonomicName id="57FE17BC1728FFDAFE8EBFDFFDB8E496" authority="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault, 2024" authorityName="Nel &amp; Wei &amp; Niu &amp; Coster &amp; Boderau &amp; Josse &amp; Kundura &amp; Kundura &amp; Brisac &amp; Boudet &amp; Jouault" authorityYear="2024" box="[272,601,1963,1989]" class="Insecta" family="Tenthredinidae" genus="Luberotenthredo" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="cerestensis" status="gen. et sp. nov.">
<emphasis id="A28AB02D1728FFDAFE8EBFDFFDB8E496" box="[272,601,1963,1989]" italics="true" pageId="6" pageNumber="7">Luberotenthredo cerestensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9B90D561728FFDAFDC1BFD8FCEFE495" box="[607,782,1964,1990]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, holotype (PNRL 2716). Reconstructions of the wing venations with names of cells and veins labeled. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="90416C3F1729FFDBFF23B97DFB1EE14A" blockId="7.[188,1399,265,538]" pageId="7" pageNumber="8">
<smallCapsWord id="96A7FAE31729FFDBFF23B97DFF16E271" baselines="284,285" box="[189,247,265,291]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Hind" pageId="7" pageNumber="8">HIND</smallCapsWord>
<smallCapsWord id="96A7FAE31729FFDBFE9AB97AFEDDE272" baselines="284" box="[260,316,270,289]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="wing" pageId="7" pageNumber="8">WING</smallCapsWord>
. ca
<quantity id="5706C1DA1729FFDBFEE7B97DFE39E270" box="[377,472,265,291]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="7" pageNumber="8" unit="mm" value="6.3">6.3 mm</quantity>
long; thin costal cell present; thin space between C and R along anterior wing margin; M+Cu long, nearly straight, fork located anteriad Rs origin: 1M long, slightly arched, apparently meeting Rs (i.e., no rs-m crossvein present); abscissae of Rs distad meeting point with 1-M long and nearly straigth; cell R1 elongate and thin; cell Rs virtually fully enclosed (second abscissa of M difficult to interpret); crossvein 1rs-m located near middle of cell R1; cell 1M trapezoidal, rather broad, 1.4 times as long as wide; 1-Cu and 2-Cu of similar lengths, closed distally by 1m-cu; cell 1Cu elongate, rectangular, closed distally by cu-a; 1-A long slightly sinusoidal; crossvein a-a located at level of 1-M origin; 2-A long, slightly arched; cell 1A wider than 1Cu but shorter; vein 3-A present, short.
</paragraph>
<paragraph id="90416C3F1729FFDBFF23BA35FD13E12D" blockId="7.[189,1398,577,639]" pageId="7" pageNumber="8">
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(partly preserved). Without visible surface sculpture, virtually
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long,
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wide; ovipositor sheath short and broad,
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long.
</paragraph>
</subSubSection>
<subSubSection id="D8E43FB41729FFDBFF23BAD2FEC9E7EB" pageId="7" pageNumber="8" type="discussion">
<paragraph id="90416C3F1729FFDBFF23BAD2FECEE193" blockId="7.[189,303,678,704]" box="[189,303,678,704]" pageId="7" pageNumber="8">
<heading id="CB09DB531729FFDBFF23BAD2FECEE193" bold="true" box="[189,303,678,704]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="A28AB02D1729FFDBFF23BAD2FECEE193" bold="true" box="[189,303,678,704]" pageId="7" pageNumber="8">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="90416C3F1729FFDBFF23BAA0FEC9E7EB" blockId="7.[187,1399,724,1208]" pageId="7" pageNumber="8">
The new fossil differs from the genus
<taxonomicName id="57FE17BC1729FFDBFD1FBAA0FBFAE1BD" authority="Vilhelmsen &amp; Engel, 2012" authorityName="Vilhelmsen &amp; Engel" authorityYear="2012" box="[641,1051,724,750]" class="Insecta" family="Tenthredinidae" genus="Sambia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="A28AB02D1729FFDBFD1FBAA0FD38E1BD" box="[641,729,724,750]" italics="true" pageId="7" pageNumber="8">Sambia</emphasis>
<bibRefCitation id="F46F11CE1729FFDBFD7EBAA0FBFAE1BD" author="Vilhelmsen L. &amp; Engel M. S." box="[736,1051,724,750]" pageId="7" pageNumber="8" pagination="271 - 281" refId="ref10431" refString="Vilhelmsen L. &amp; Engel M. S. 2012. Sambia succinica, a crown group tenthredinid from Eocene Baltic amber (Hymenoptera: Tenthredinidae). Insect Systematics and Evolution 43: 271 - 281. https: // doi. org / 10.1163 / 1876312 X- 04303003" type="journal article" year="2012">Vilhelmsen &amp; Engel, 2012</bibRefCitation>
</taxonomicName>
(Eocene Baltic amber) in the forewing veins 2+3-A, which are fused with 1-A on a short distance (vs a long crossvein in-between) and its elongate flagellomeres (
<bibRefCitation id="F46F11CE1729FFDBFDB5BB6EFC82E067" author="Vilhelmsen L. &amp; Engel M. S." box="[555,867,794,820]" pageId="7" pageNumber="8" pagination="271 - 281" refId="ref10431" refString="Vilhelmsen L. &amp; Engel M. S. 2012. Sambia succinica, a crown group tenthredinid from Eocene Baltic amber (Hymenoptera: Tenthredinidae). Insect Systematics and Evolution 43: 271 - 281. https: // doi. org / 10.1163 / 1876312 X- 04303003" type="journal article" year="2012">Vilhelmsen &amp; Engel 2012</bibRefCitation>
). With the lower Oligocene genus
<taxonomicName id="57FE17BC1729FFDBFB69BB6EFE83E00B" authority="Rohwer, 1908" authorityName="Rohwer" authorityYear="1908" class="Arachnida" family="Gymnodamaeidae" genus="Nortonella" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="A28AB02D1729FFDBFB69BB6EFA97E067" box="[1271,1398,794,820]" italics="true" pageId="7" pageNumber="8">Nortonella</emphasis>
<bibRefCitation id="F46F11CE1729FFDBFF23BB49FE83E00B" author="Rohwer S. A." box="[189,354,829,856]" pageId="7" pageNumber="8" pagination="591 - 595" refId="ref9407" refString="Rohwer S. A. 1908. The Tertiary Tenthredinoidea of the expedition of 1908 to Florissant. Bulletin of the American Museum of Natural History 24: 591 - 595. Available from http: // hdl. handle. net / 2246 / 1721 [accessed 30 Nov. 2023]." type="journal article" year="1908">Rohwer, 1908</bibRefCitation>
</taxonomicName>
it shares the vein 2+3-A shortly fused with 1-A (
<bibRefCitation id="F46F11CE1729FFDBFC09BB49FBDAE00B" author="Rohwer S. A." box="[919,1083,829,856]" pageId="7" pageNumber="8" pagination="591 - 595" refId="ref9407" refString="Rohwer S. A. 1908. The Tertiary Tenthredinoidea of the expedition of 1908 to Florissant. Bulletin of the American Museum of Natural History 24: 591 - 595. Available from http: // hdl. handle. net / 2246 / 1721 [accessed 30 Nov. 2023]." type="journal article" year="1908">Rohwer 1908</bibRefCitation>
: fig. 1). But
<taxonomicName id="57FE17BC1729FFDBFB54BB49FAA8E004" authorityName="Rohwer" authorityYear="1908" box="[1226,1353,829,855]" class="Arachnida" family="Gymnodamaeidae" genus="Nortonella" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="A28AB02D1729FFDBFB54BB49FAA8E004" box="[1226,1353,829,855]" italics="true" pageId="7" pageNumber="8">Nortonella</emphasis>
</taxonomicName>
has the crossvein 1cu-a very close to the base of 1-M, versus about midway between the base of 1-M and 3-Cu in the new fossil and in several extant species of the genus
<taxonomicName id="57FE17BC1729FFDBFC04BBF1FB4AE0CD" authority="Hartig, 1837" authorityName="Hartig" authorityYear="1837" box="[922,1195,900,926]" class="Insecta" family="Tenthredinidae" genus="Perineura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="A28AB02D1729FFDBFC04BBF1FBF0E0CD" box="[922,1041,901,926]" italics="true" pageId="7" pageNumber="8">Perineura</emphasis>
Hartig, 1837
</taxonomicName>
(see photographs of the
<typeStatus id="4F45D29D1729FFDBFE94BBD3FE90E092" box="[266,369,935,961]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
UCM 4517: https://invertpaleosearch.colorado.edu/). The new fossil can be separated from the lower Oligocene genus
<taxonomicName id="57FE17BC1729FFDBFDDEBBBFFC67E0B7" authority="Cockerell, 1917" authorityName="Cockerell" authorityYear="1917" box="[576,902,970,996]" class="Insecta" family="Tenthredinidae" genus="Taeniurites" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="A28AB02D1729FFDBFDDEBBBFFD22E0B7" box="[576,707,971,996]" italics="true" pageId="7" pageNumber="8">Taeniurites</emphasis>
<bibRefCitation id="F46F11CE1729FFDBFD54BBBEFC67E0B7" author="Cockerell T. D. A." box="[714,902,970,996]" pageId="7" pageNumber="8" pagination="373 - 384" refId="ref7409" refString="Cockerell T. D. A. 1917. New Tertiary insects. Proceedings of the United States National Museum 52: 373 - 384. https: // doi. org / 10.5479 / si. 00963801.52 - 2181.373" type="journal article" year="1917">Cockerell, 1917</bibRefCitation>
</taxonomicName>
by the vein 2+3-A, which is shortly fused with 1-A (vs separated) and vein 1cu-a about midway between the base of 1-M and 3-Cu, vs very close to the base of 1-M (
<bibRefCitation id="F46F11CE1729FFDBFE38BC65FD80E778" author="Cockerell T. D. A." box="[422,609,1041,1067]" pageId="7" pageNumber="8" pagination="373 - 384" refId="ref7409" refString="Cockerell T. D. A. 1917. New Tertiary insects. Proceedings of the United States National Museum 52: 373 - 384. https: // doi. org / 10.5479 / si. 00963801.52 - 2181.373" type="journal article" year="1917">Cockerell 1917</bibRefCitation>
). The Uppermost Oligocene genus
<taxonomicName id="57FE17BC1729FFDBFB9CBC65FABAE77F" authority="Meunier, 1915" authorityName="Meunier" authorityYear="1915" box="[1026,1371,1041,1068]" class="Insecta" family="Pompilidae" genus="Tenthredinites" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="A28AB02D1729FFDBFB9CBC65FB49E778" box="[1026,1192,1041,1067]" italics="true" pageId="7" pageNumber="8">Tenthredinites</emphasis>
<bibRefCitation id="F46F11CE1729FFDBFB31BC65FABAE77F" author="Meunier F." box="[1199,1371,1041,1068]" pageId="7" pageNumber="8" pagination="1 - 17" refId="ref8745" refString="Meunier F. 1915. Nouvelles recherches sur quelques insectes des platrieres d'Aix-en-Provence. Verhandelingen der Koninklijke Akademie van Wetenschappen te Amsterdam (2) 18: 1 - 17." type="journal article" year="1915">Meunier, 1915</bibRefCitation>
</taxonomicName>
is based on a poorly preserved specimen (
<bibRefCitation id="F46F11CE1729FFDBFD11BC40FCDBE71C" author="Meunier F." box="[655,826,1076,1103]" pageId="7" pageNumber="8" pagination="1 - 17" refId="ref8745" refString="Meunier F. 1915. Nouvelles recherches sur quelques insectes des platrieres d'Aix-en-Provence. Verhandelingen der Koninklijke Akademie van Wetenschappen te Amsterdam (2) 18: 1 - 17." type="journal article" year="1915">Meunier 1915</bibRefCitation>
), which is probably lost.
<bibRefCitation id="F46F11CE1729FFDBFBFABC40FA97E71D" author="Rodriguez J. &amp; Waichert C. &amp; von Dohlen C. D. &amp; Pitts J. P." box="[1124,1398,1076,1102]" pageId="7" pageNumber="8" pagination="0185379" refId="ref9337" refString="Rodriguez J., Waichert C., von Dohlen C. D. &amp; Pitts J. P. 2017. The geological record and phylogeny of spider wasps (Hymenoptera: Pompilidae): a revision of fossil species and their phylogenetic placement. PLoS ONE 12 (10): e 0185379. https: // doi. org / 10.1371 / journal. pone. 0185379" type="journal article" year="2017">
Rodriguez
<emphasis id="A28AB02D1729FFDBFB78BC41FAC0E71D" box="[1254,1313,1076,1102]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2017)
</bibRefCitation>
suggested it could be a
<taxonomicName id="57FE17BC1729FFDBFE7DBC23FCC6E721" authority="Latreille, 1805" authorityName="Latreille" authorityYear="1805" box="[483,807,1111,1138]" class="Insecta" family="Pompilidae" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="family">Pompilidae Latreille, 1805</taxonomicName>
. The Miocene genus
<taxonomicName id="57FE17BC1729FFDBFBB1BC23FA96E722" authority="Zhang, 1989" authorityName="Zhang" authorityYear="1989" box="[1071,1399,1111,1137]" class="Insecta" family="Tenthredinidae" genus="Tenthredoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="A28AB02D1729FFDBFBB1BC23FB32E722" box="[1071,1235,1111,1137]" italics="true" pageId="7" pageNumber="8">Tenthredoides</emphasis>
<bibRefCitation id="F46F11CE1729FFDBFB40BC23FA96E722" author="Zhang J." box="[1246,1399,1111,1137]" pageId="7" pageNumber="8" refId="ref10795" refString="Zhang J. 1989. [Fossil insects from Shanwang, Shandong, China.] Shandong Science and Technology Publishing House, Jinan. [In Chinese with abstract in English.]" type="book" year="1989">Zhang, 1989</bibRefCitation>
</taxonomicName>
differs from the new fossil in the vein 1cu-a, which is very close to the base of 1-M (
<bibRefCitation id="F46F11CE1729FFDBFB3CBC0EFAD4E7C7" author="Zhang J." box="[1186,1333,1146,1172]" pageId="7" pageNumber="8" refId="ref10795" refString="Zhang J. 1989. [Fossil insects from Shanwang, Shandong, China.] Shandong Science and Technology Publishing House, Jinan. [In Chinese with abstract in English.]" type="book" year="1989">Zhang 1989</bibRefCitation>
: textfig. 224).
</paragraph>
</subSubSection>
</treatment>
</document>