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<document id="B439F12DBF8754A210387F9A2951DB26" ID-CLB-Dataset="2443" ID-DOI="10.5281/zenodo.5740035" ID-GBIF-Dataset="6874a62b-f2be-4314-b3ef-fd91600ffceb" ID-ISSN="0374-1036" ID-Zenodo-Dep="5740035" ID-ZooBank="9A396426-08FE-4E2A-A4EF-CD0DA819D8AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638253787490" checkinUser="marcus" docAuthor="Bezděk, Jan" docDate="2013" docId="166387BCFFFEFFD19A00FDA3FBCF6BC1" docLanguage="en" docName="ActaEntMusNatPra.2013.53.2.715-746.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Hesperopenna medvedevi Bezděk 2013" docType="key" docVersion="8" lastPageNumber="724" masterDocId="EA5AFFC4FFF6FFD89B56FFEBFF8F6A15" masterDocTitle="Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group" masterLastPageNumber="746" masterPageNumber="715" pageNumber="723" updateTime="1699471224025" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A8E1ED70F4483C739BACB6F20C28622F">Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group</mods:title>
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<mods:namePart id="70246E8C04EA1A90AE9960B605E9CBC0">Bezděk, Jan</mods:namePart>
<mods:affiliation id="7299B4843EE2016257A50E52A36AD531">Mendel University, Department of Zoology, Zemědělská 1, CZ – 613 00 Brno, Czech Republic</mods:affiliation>
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<mods:title id="F781129B37C578835972509E04416D2B">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="CCD7D20A994A8FC7BBA1F144F840A02F">2013</mods:date>
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<mods:number id="76B19831B63587FCA3F56423B4AF1ED0">2013-11-15</mods:number>
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<treatment id="166387BCFFFEFFD19A00FDA3FBCF6BC1" ID-DOI="http://doi.org/10.5281/zenodo.5740308" ID-GBIF-Taxon="192572520" ID-Zenodo-Dep="5740308" LSID="urn:lsid:plazi:treatment:166387BCFFFEFFD19A00FDA3FBCF6BC1" httpUri="http://treatment.plazi.org/id/166387BCFFFEFFD19A00FDA3FBCF6BC1" lastPageId="9" lastPageNumber="724" pageId="8" pageNumber="723">
<subSubSection id="D6D06521FFFEFFD09A00FDA3FC286875" box="[342,935,583,608]" pageId="8" pageNumber="723" type="nomenclature">
<paragraph id="9E7536AAFFFEFFD09A00FDA3FC286875" blockId="8.[342,935,583,608]" box="[342,935,583,608]" pageId="8" pageNumber="723">
<heading id="C53D81C6FFFEFFD09A00FDA3FC286875" bold="true" box="[342,935,583,608]" centered="true" fontSize="8" level="2" pageId="8" pageNumber="723" reason="3">
<emphasis id="ACBEEAB8FFFEFFD09A00FDA3FC286875" bold="true" box="[342,935,583,608]" pageId="8" pageNumber="723">
Key to the species of
<taxonomicName id="59CA4D29FFFEFFD09912FDA3FCD26875" authorityName="Bezděk" authorityYear="2013" box="[580,861,583,608]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="723" phylum="Arthropoda" rank="species" species="medvedevi">
<emphasis id="ACBEEAB8FFFEFFD09912FDA3FCD26875" bold="true" box="[580,861,583,608]" italics="true" pageId="8" pageNumber="723">Hesperopenna medvedevi</emphasis>
</taxonomicName>
group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D6D06521FFFEFFD19BC1FD9CFBCF6BC1" lastPageId="9" lastPageNumber="724" pageId="8" pageNumber="723" type="key">
<keyStep id="253E2E0FFFFEFFD09BC1FD9CFBE968DA" pageId="8" pageNumber="723">
<paragraph id="9E7536AAFFFEFFD09BC1FD9CFBE9689A" blockId="8.[151,1127,631,1583]" box="[151,1126,631,655]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FD9CFBE9689A" box="[151,1126,631,655]" pageId="8" pageNumber="723">1 Tibiae and tarsi black, elytra yellowish, glabrous. ......................................................... 2</keyLead>
</paragraph>
<paragraph id="9E7536AAFFFEFFD09BC1FD7CFBE968DA" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FD7CFBE968DA" pageId="8" pageNumber="723">
<emphasis id="ACBEEAB8FFFEFFD09BC1FD7CFF2B68BA" bold="true" box="[151,164,663,687]" pageId="8" pageNumber="723">–</emphasis>
Legs completely brownish or orange, elytra brown, glabrous or partly covered with fine setae. ............................................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="253E2E0FFFFEFFD09BC1FD3CFBE969DA" pageId="8" pageNumber="723">
<paragraph id="9E7536AAFFFEFFD09BC1FD3CFBE9695A" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FD3CFBE9695A" pageId="8" pageNumber="723">
2 Antennae completely yellow. Extreme margin of epipleura in apical third yellow or at most brownish. Aedeagus ventrally with long thin incision placed on right side (
<figureCitation id="06F12A2FFFFEFFD09F6DFD1CFF55693A" captionStart="Figs 11–14" captionStartId="12.[151,188,1557,1576]" captionTargetBox="[158,1115,231,1431]" captionTargetId="figure-10@12.[151,1126,219,1436]" captionTargetPageId="12" captionText="Figs 11–14.Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 11 – Hesperopenna medvedevi nom.nov.; 12 – H. persimilis (Kimoto, 1989); 13 – H. tibialis (Kimoto, 1989); 14 – H. arnoldi sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740043" httpUri="https://zenodo.org/record/5740043/files/figure.png" pageId="8" pageNumber="723">Fig. 13</figureCitation>
). Female pygidium with apex distinctly pointed (
<figureCitation id="06F12A2FFFFEFFD099B1FCFCFCB8693A" box="[743,823,791,815]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="8" pageNumber="723">Fig. 36</figureCitation>
).
<collectingCountry id="E6DD763AFFFEFFD0981FFCF3FCF0693A" box="[841,895,792,815]" name="Laos" pageId="8" pageNumber="723">Laos</collectingCountry>
,
<collectingCountry id="E6DD763AFFFEFFD098DCFCFCFC63693A" box="[906,1004,791,815]" name="Thailand" pageId="8" pageNumber="723">Thailand</collectingCountry>
, Peninsular
<collectingCountry id="E6DD763AFFFEFFD09BEBFCDCFEAD695A" box="[189,290,823,847]" name="Malaysia" pageId="8" pageNumber="723">Malaysia</collectingCountry>
. ................................................................................
<taxonomicName id="59CA4D29FFFEFFD0981BFCDCFBE9695A" authority="(Kimoto, 1989)" baseAuthorityName="Kimoto" baseAuthorityYear="1989" box="[845,1126,822,847]" class="Insecta" family="Chrysomelidae" genus="Liroetiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="723" phylum="Arthropoda" rank="species" species="tibialis">
<emphasis id="ACBEEAB8FFFEFFD0981BFCDCFC36695A" bold="true" box="[845,953,822,847]" italics="true" pageId="8" pageNumber="723">H. tibialis</emphasis>
(
<bibRefCitation id="FA5B4B5BFFFEFFD09891FCDCFBD0695A" author="KIMOTO S." box="[967,1119,823,847]" pageId="8" pageNumber="723" pagination="1 - 241" refId="ref15334" refString="KIMOTO S. 1989: Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. IV. Galerucinae. Esakia 27: 1 - 241." type="journal article" year="1989">Kimoto, 1989</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="9E7536AAFFFEFFD09BC1FCBCFBE969DA" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FCBCFBE969DA" pageId="8" pageNumber="723">
<emphasis id="ACBEEAB8FFFEFFD09BC1FCBCFF2B697A" bold="true" box="[151,164,855,879]" pageId="8" pageNumber="723">–</emphasis>
Antennae black with first two antennomeres pale. Extreme margin of epipleura in apical third black. Aedeagus ventrally with long thin incision placed medially (
<figureCitation id="06F12A2FFFFEFFD098E1FC9CFB8A699A" box="[951,1029,887,911]" captionStart="Figs 15–18" captionStartId="13.[115,152,1542,1561]" captionTargetBox="[126,1079,226,1362]" captionTargetId="figure-14@13.[113,1090,218,1370]" captionTargetPageId="13" captionText="Figs 15–18. Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 15 – H. bonifaci sp. nov.; 16 – H. helferi sp. nov.; 17 – H. sipekorum sp. nov.; 18 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740045" httpUri="https://zenodo.org/record/5740045/files/figure.png" pageId="8" pageNumber="723">Fig. 18</figureCitation>
). Female pygidium with apex simply triangular (
<figureCitation id="06F12A2FFFFEFFD09909FC7CFD2069BA" box="[607,687,919,943]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="8" pageNumber="723">Fig. 40</figureCitation>
).
<collectingCountry id="E6DD763AFFFEFFD09997FC7CFCA669BA" box="[705,809,919,943]" name="Indonesia" pageId="8" pageNumber="723">Indonesia</collectingCountry>
(
<collectingRegion id="5C0EF848FFFEFFD09861FC7CFCE569BA" box="[823,874,919,943]" country="Indonesia" name="Jawa Timur" pageId="8" pageNumber="723">Java</collectingRegion>
,
<collectingRegion id="5C0EF848FFFEFFD09823FC7CFC2969BA" box="[885,934,919,943]" country="Indonesia" name="Bali" pageId="8" pageNumber="723">Bali</collectingRegion>
). ......................... ...............................................................................................................
<emphasis id="ACBEEAB8FFFEFFD098FBFC5CFBE969DA" bold="true" box="[941,1126,950,975]" pageId="8" pageNumber="723">
<taxonomicName id="59CA4D29FFFEFFD098FBFC5CFB8569DA" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[941,1034,950,975]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="723" phylum="Arthropoda" rank="species" species="zofka" status="sp. nov.">
<emphasis id="ACBEEAB8FFFEFFD098FBFC5CFB8569DA" bold="true" box="[941,1034,950,975]" italics="true" pageId="8" pageNumber="723">H. zofka</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFEFFD09F47FC5CFBE969DA" box="[1041,1126,951,975]" pageId="8" pageNumber="723" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="253E2E0FFFFEFFD09BC1FC3CFBE96E1A" pageId="8" pageNumber="723">
<paragraph id="9E7536AAFFFEFFD09BC1FC3CFBE969FA" blockId="8.[151,1127,631,1583]" box="[151,1126,983,1007]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FC3CFBE969FA" box="[151,1126,983,1007]" pageId="8" pageNumber="723">3 Elytra glabrous. .............................................................................................................. 4</keyLead>
</paragraph>
<paragraph id="9E7536AAFFFEFFD09BC1FC1CFBE96E1A" blockId="8.[151,1127,631,1583]" box="[151,1126,1015,1039]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FC1CFBE96E1A" box="[151,1126,1015,1039]" pageId="8" pageNumber="723">
<emphasis id="ACBEEAB8FFFEFFD09BC1FC1CFF2B6E1A" bold="true" box="[151,164,1015,1039]" pageId="8" pageNumber="723">–</emphasis>
Elytra densely covered with fine setae at least in apical half of elytra, .......................... 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="253E2E0FFFFEFFD09BC1FBFCFBE96EDA" pageId="8" pageNumber="723">
<paragraph id="9E7536AAFFFEFFD09BC1FBFCFBE96E9A" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FBFCFBE96E9A" pageId="8" pageNumber="723">
4 Last ventrite in male with subapical impression anteriorly surrounded with U-shaped ridge (
<figureCitation id="06F12A2FFFFEFFD09A54FBDCFEDE6E5A" box="[258,337,1079,1103]" captionStart="Figs 19–26" captionStartId="15.[112,149,1058,1077]" captionTargetBox="[124,1078,218,1029]" captionTargetId="figure-153@15.[113,1088,207,1034]" captionTargetPageId="15" captionText="Figs 19–26. Last abdominal ventrite of males.19 – Hesperopenna medvedevi nom. nov.; 20 – H. persimilis (Kimoto, 1989); 21 – H. tibialis (Kimoto, 1989); 22 – H. arnoldi sp. nov.; 23 – H. bonifaci sp. nov.; 24 – H. helferi sp. nov.; 25 – H. sipekorum sp. nov.; 26 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740049" httpUri="https://zenodo.org/record/5740049/files/figure.png" pageId="8" pageNumber="723">Fig. 20</figureCitation>
). Aedeagus ventraly flat, with gradually tapering triangular apex (
<figureCitation id="06F12A2FFFFEFFD09F5DFBDCFBD56E5A" box="[1035,1114,1079,1103]" captionStart="Figs 11–14" captionStartId="12.[151,188,1557,1576]" captionTargetBox="[158,1115,231,1431]" captionTargetId="figure-10@12.[151,1126,219,1436]" captionTargetPageId="12" captionText="Figs 11–14.Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 11 – Hesperopenna medvedevi nom.nov.; 12 – H. persimilis (Kimoto, 1989); 13 – H. tibialis (Kimoto, 1989); 14 – H. arnoldi sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740043" httpUri="https://zenodo.org/record/5740043/files/figure.png" pageId="8" pageNumber="723">Fig. 12</figureCitation>
). Female pygidium with apex obtusangulate (
<figureCitation id="06F12A2FFFFEFFD099C7FBBCFD6E6E7A" box="[657,737,1111,1135]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="8" pageNumber="723">Fig. 35</figureCitation>
).
<collectingCountry id="E6DD763AFFFEFFD099A5FBBCFCDF6E7A" box="[755,848,1111,1135]" name="Vietnam" pageId="8" pageNumber="723">Vietnam</collectingCountry>
,
<collectingCountry id="E6DD763AFFFEFFD0980DFBB3FC1E6E7A" box="[859,913,1112,1135]" name="Laos" pageId="8" pageNumber="723">Laos</collectingCountry>
,
<collectingCountry id="E6DD763AFFFEFFD098CDFBBCFC726E7A" box="[923,1021,1111,1135]" name="Thailand" pageId="8" pageNumber="723">Thailand</collectingCountry>
. .............. ............................................................................................
<taxonomicName id="59CA4D29FFFEFFD0987BFB9CFBE96E9A" authority="(Kimoto, 1989)" baseAuthorityName="Kimoto" baseAuthorityYear="1989" box="[813,1126,1142,1167]" class="Insecta" family="Chrysomelidae" genus="Calomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="723" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="ACBEEAB8FFFEFFD0987BFB9CFC356E9A" bold="true" box="[813,954,1142,1167]" italics="true" pageId="8" pageNumber="723">H. persimilis</emphasis>
(
<bibRefCitation id="FA5B4B5BFFFEFFD09891FB9CFBD06E9A" author="KIMOTO S." box="[967,1119,1143,1167]" pageId="8" pageNumber="723" pagination="1 - 241" refId="ref15334" refString="KIMOTO S. 1989: Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. IV. Galerucinae. Esakia 27: 1 - 241." type="journal article" year="1989">Kimoto, 1989</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="9E7536AAFFFEFFD09BC1FB7CFBE96EDA" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FB7CFBE96EDA" pageId="8" pageNumber="723">
<emphasis id="ACBEEAB8FFFEFFD09BC1FB7CFF2B6EBA" bold="true" box="[151,164,1175,1199]" pageId="8" pageNumber="723">–</emphasis>
Last ventrite in male without U-shaped ridge. Female pygidium with small U-shaped incision at apex (
<figureCitation id="06F12A2FFFFEFFD09A25FB5CFE4C6EDA" box="[371,451,1207,1231]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="8" pageNumber="723">Fig. 38</figureCitation>
). ............................................................................................... 5
</keyLead>
</paragraph>
</keyStep>
<keyStep id="253E2E0FFFFEFFD09BC1FB3CFBE96F9A" pageId="8" pageNumber="723">
<paragraph id="9E7536AAFFFEFFD09BC1FB3CFBE96F3A" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FB3CFBE96F3A" pageId="8" pageNumber="723">
5 Aedeagus dorsally with two thin touching processes, ventrally with deep incision on left side, ventral surface not keeled in the middle (
<figureCitation id="06F12A2FFFFEFFD099FCFB1CFD766F1A" box="[682,761,1271,1295]" captionStart="Figs 15–18" captionStartId="13.[115,152,1542,1561]" captionTargetBox="[126,1079,226,1362]" captionTargetId="figure-14@13.[113,1090,218,1370]" captionTargetPageId="13" captionText="Figs 15–18. Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 15 – H. bonifaci sp. nov.; 16 – H. helferi sp. nov.; 17 – H. sipekorum sp. nov.; 18 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740045" httpUri="https://zenodo.org/record/5740045/files/figure.png" pageId="8" pageNumber="723">Fig. 15</figureCitation>
). North
<collectingCountry id="E6DD763AFFFEFFD09804FB13FC336F05" box="[850,956,1272,1296]" name="Myanmar" pageId="8" pageNumber="723">Myanmar</collectingCountry>
. ....................... ..........................................................................................................
<emphasis id="ACBEEAB8FFFEFFD098D8FAFCFBE96F3A" bold="true" box="[910,1126,1302,1327]" pageId="8" pageNumber="723">
<taxonomicName id="59CA4D29FFFEFFD098D8FAFCFB856F3A" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[910,1034,1302,1327]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="723" phylum="Arthropoda" rank="species" species="bonifaci" status="sp. nov.">
<emphasis id="ACBEEAB8FFFEFFD098D8FAFCFB856F3A" bold="true" box="[910,1034,1302,1327]" italics="true" pageId="8" pageNumber="723">H. bonifaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFEFFD09F47FAFCFBE96F3A" box="[1041,1126,1303,1327]" pageId="8" pageNumber="723" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="9E7536AAFFFEFFD09BC1FADCFBE96F9A" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FADCFBE96F9A" pageId="8" pageNumber="723">
<emphasis id="ACBEEAB8FFFEFFD09BC1FADCFF2B6F5A" bold="true" box="[151,164,1335,1359]" pageId="8" pageNumber="723">–</emphasis>
Aedeagus dorsally forms two wider touching processes, ventrally with deep incision on right side, ventral surface with high median keel (
<figureCitation id="06F12A2FFFFEFFD09987FABCFCAE6F7A" box="[721,801,1367,1391]" captionStart="Figs 15–18" captionStartId="13.[115,152,1542,1561]" captionTargetBox="[126,1079,226,1362]" captionTargetId="figure-14@13.[113,1090,218,1370]" captionTargetPageId="13" captionText="Figs 15–18. Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 15 – H. bonifaci sp. nov.; 16 – H. helferi sp. nov.; 17 – H. sipekorum sp. nov.; 18 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740045" httpUri="https://zenodo.org/record/5740045/files/figure.png" pageId="8" pageNumber="723">Fig. 16</figureCitation>
). Female pygidium with small U-shaped incision at apex (
<figureCitation id="06F12A2FFFFEFFD09AB6FA9CFDBF6F9A" box="[480,560,1399,1423]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="8" pageNumber="723">Fig. 38</figureCitation>
). South
<collectingCountry id="E6DD763AFFFEFFD099D1FA93FD7E6F85" box="[647,753,1400,1424]" name="Myanmar" pageId="8" pageNumber="723">Myanmar</collectingCountry>
. .......................
<emphasis id="ACBEEAB8FFFEFFD098C9FA9CFBE96F9A" bold="true" box="[927,1126,1398,1423]" pageId="8" pageNumber="723">
<taxonomicName id="59CA4D29FFFEFFD098C9FA9CFB856F9A" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[927,1034,1398,1423]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="723" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFEFFD098C9FA9CFB856F9A" bold="true" box="[927,1034,1398,1423]" italics="true" pageId="8" pageNumber="723">H. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFEFFD09F47FA9CFBE96F9A" box="[1041,1126,1399,1423]" pageId="8" pageNumber="723" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="253E2E0FFFFEFFD09BC1FA7CFBE96C3A" pageId="8" pageNumber="723">
<paragraph id="9E7536AAFFFEFFD09BC1FA7CFBE96FBA" blockId="8.[151,1127,631,1583]" box="[151,1126,1431,1455]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FA7CFBE96FBA" box="[151,1126,1431,1455]" pageId="8" pageNumber="723">6 Elytra densely covered with fine setae only in apical half. ............................................ 7</keyLead>
</paragraph>
<paragraph id="9E7536AAFFFEFFD09BC1FA5CFBE96C3A" blockId="8.[151,1127,631,1583]" pageId="8" pageNumber="723">
<keyLead id="253B959FFFFEFFD09BC1FA5CFBE96C3A" pageId="8" pageNumber="723">
<emphasis id="ACBEEAB8FFFEFFD09BC1FA5CFF2B6FDA" bold="true" box="[151,164,1463,1487]" pageId="8" pageNumber="723">–</emphasis>
Elytra almost completely covered with fine setae (at least in apical two thirds). Aedeagus ventrally with deep U-shaped incision on right side (
<figureCitation id="06F12A2FFFFEFFD0998BFA3CFCA66FFA" box="[733,809,1495,1519]" captionStart="Figs 11–14" captionStartId="12.[151,188,1557,1576]" captionTargetBox="[158,1115,231,1431]" captionTargetId="figure-10@12.[151,1126,219,1436]" captionTargetPageId="12" captionText="Figs 11–14.Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 11 – Hesperopenna medvedevi nom.nov.; 12 – H. persimilis (Kimoto, 1989); 13 – H. tibialis (Kimoto, 1989); 14 – H. arnoldi sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740043" httpUri="https://zenodo.org/record/5740043/files/figure.png" pageId="8" pageNumber="723">Fig. 14</figureCitation>
). Female pygidium with small semicircular incision at apex (
<figureCitation id="06F12A2FFFFEFFD09AABFA1CFDC26C1A" box="[509,589,1527,1551]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="8" pageNumber="723">Fig. 37</figureCitation>
).
<collectingCountry id="E6DD763AFFFEFFD09936FA13FD196C1A" box="[608,662,1528,1551]" name="Laos" pageId="8" pageNumber="723">Laos</collectingCountry>
,
<collectingCountry id="E6DD763AFFFEFFD099F7FA1CFC8C6C1A" box="[673,771,1527,1551]" name="Thailand" pageId="8" pageNumber="723">Thailand</collectingCountry>
,
<collectingCountry id="E6DD763AFFFEFFD0985BFA1CFCC26C1A" box="[781,845,1527,1551]" name="China" pageId="8" pageNumber="723">China</collectingCountry>
(
<collectingRegion id="5C0EF848FFFEFFD0980DFA1CFC3C6C1A" box="[859,947,1527,1551]" country="China" name="Yunnan" pageId="8" pageNumber="723">Yunnan</collectingRegion>
). ........................ ............................................................................................................
<emphasis id="ACBEEAB8FFFEFFD098CEF9FCFBE96C3A" bold="true" box="[920,1126,1558,1583]" pageId="8" pageNumber="723">
<taxonomicName id="59CA4D29FFFEFFD098CEF9FCFB856C3A" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[920,1034,1558,1583]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="723" phylum="Arthropoda" rank="species" species="arnoldi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFEFFD098CEF9FCFB856C3A" bold="true" box="[920,1034,1558,1583]" italics="true" pageId="8" pageNumber="723">H. arnoldi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFEFFD09F47F9FCFBE96C3A" box="[1041,1126,1559,1583]" pageId="8" pageNumber="723" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="253E2E0FFFFFFFD19B27FF37FBCF6BC1" pageId="9" pageNumber="724">
<paragraph id="9E7536AAFFFFFFD19B27FF37FBCF6B41" blockId="9.[113,1089,219,468]" pageId="9" pageNumber="724">
<keyLead id="253B959FFFFFFFD19B27FF37FBCF6B41" pageId="9" pageNumber="724">
7 Aedeagus ventrally with symmetrical subtriangular apex, lateral incisions short and of the same depth, ventral surface flat with thin median keel (
<figureCitation id="06F12A2FFFFFFFD1984BFF10FCE26B01" box="[797,877,251,276]" captionStart="Figs 11–14" captionStartId="12.[151,188,1557,1576]" captionTargetBox="[158,1115,231,1431]" captionTargetId="figure-10@12.[151,1126,219,1436]" captionTargetPageId="12" captionText="Figs 11–14.Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 11 – Hesperopenna medvedevi nom.nov.; 12 – H. persimilis (Kimoto, 1989); 13 – H. tibialis (Kimoto, 1989); 14 – H. arnoldi sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740043" httpUri="https://zenodo.org/record/5740043/files/figure.png" pageId="9" pageNumber="724">Fig. 11</figureCitation>
). Female pygidium with U-shaped incision and slightly concave posterior margins near incision (
<figureCitation id="06F12A2FFFFFFFD198B5FEF0FBBB6B21" box="[995,1076,283,308]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="9" pageNumber="724">Fig. 37</figureCitation>
).
<collectingCountry id="E6DD763AFFFFFFD19BC1FED0FF7A6B46" box="[151,245,315,339]" name="Vietnam" pageId="9" pageNumber="724">Vietnam</collectingCountry>
. ...................................................................................
<emphasis id="ACBEEAB8FFFFFFD19861FED7FBCF6B41" bold="true" box="[823,1088,314,340]" pageId="9" pageNumber="724">
<taxonomicName id="59CA4D29FFFFFFD19861FED7FC446B46" authorityName="Bezděk" authorityYear="2013" box="[823,971,314,339]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="724" phylum="Arthropoda" rank="species" species="medvedevi" status="nom. nov.">
<emphasis id="ACBEEAB8FFFFFFD19861FED7FC446B46" bold="true" box="[823,971,314,339]" italics="true" pageId="9" pageNumber="724">H. medvedevi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFFFFD19884FED7FBCF6B41" box="[978,1088,316,340]" pageId="9" pageNumber="724">nom. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="9E7536AAFFFFFFD19B27FEB7FBCF6BC1" blockId="9.[113,1089,219,468]" pageId="9" pageNumber="724">
<keyLead id="253B959FFFFFFFD19B27FEB7FBCF6BC1" pageId="9" pageNumber="724">
<emphasis id="ACBEEAB8FFFFFFD19B27FEB7FFF16B61" bold="true" box="[113,126,348,372]" pageId="9" pageNumber="724">–</emphasis>
Aedeagus ventrally asymmetrical, with very deep and wide subtriangular incision on right side, ventral surface with high median keel apically forming robust process bent downwards (
<figureCitation id="06F12A2FFFFFFFD19A76FE70FEE16BA1" box="[288,366,411,436]" captionStart="Figs 15–18" captionStartId="13.[115,152,1542,1561]" captionTargetBox="[126,1079,226,1362]" captionTargetId="figure-14@13.[113,1090,218,1370]" captionTargetPageId="13" captionText="Figs 15–18. Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 15 – H. bonifaci sp. nov.; 16 – H. helferi sp. nov.; 17 – H. sipekorum sp. nov.; 18 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740045" httpUri="https://zenodo.org/record/5740045/files/figure.png" pageId="9" pageNumber="724">Fig. 17</figureCitation>
). Female pygidium with deep U-shaped incision with oblique apical margins straight (
<figureCitation id="06F12A2FFFFFFFD19A04FE50FE2D6BC1" box="[338,418,443,468]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="9" pageNumber="724">Fig. 39</figureCitation>
).
<collectingCountry id="E6DD763AFFFFFFD19AE3FE50FE636BC6" box="[437,492,443,467]" name="India" pageId="9" pageNumber="724">India</collectingCountry>
(
<collectingRegion id="5C0EF848FFFFFFD19AAFFE50FDF96BC6" box="[505,630,443,467]" country="India" name="Meghalaya" pageId="9" pageNumber="724">Meghalaya</collectingRegion>
). ............................
<emphasis id="ACBEEAB8FFFFFFD19819FE57FBCF6BC1" bold="true" box="[847,1088,442,468]" pageId="9" pageNumber="724">
<taxonomicName id="59CA4D29FFFFFFD19819FE57FC6A6BC6" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[847,997,442,467]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="724" phylum="Arthropoda" rank="species" species="sipekorum" status="sp. nov.">
<emphasis id="ACBEEAB8FFFFFFD19819FE57FC6A6BC6" bold="true" box="[847,997,442,467]" italics="true" pageId="9" pageNumber="724">H. sipekorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFFFFD198BDFE57FBCF6BC1" box="[1003,1088,444,468]" pageId="9" pageNumber="724" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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