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<document id="A7353CECE11894AEAE60C3762C715538" ID-CLB-Dataset="2443" ID-DOI="10.5281/zenodo.5740035" ID-GBIF-Dataset="6874a62b-f2be-4314-b3ef-fd91600ffceb" ID-ISSN="0374-1036" ID-Zenodo-Dep="5740035" ID-ZooBank="9A396426-08FE-4E2A-A4EF-CD0DA819D8AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638253787490" checkinUser="marcus" docAuthor="Bezděk, Jan" docDate="2013" docId="166387BCFFFCFFD39A0BFE17FDB76BA1" docLanguage="en" docName="ActaEntMusNatPra.2013.53.2.715-746.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (2)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Hesperopenna persimilis Bezděk 2013, comb. nov." docType="treatment" docVersion="11" lastPageNumber="726" masterDocId="EA5AFFC4FFF6FFD89B56FFEBFF8F6A15" masterDocTitle="Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group" masterLastPageNumber="746" masterPageNumber="715" pageNumber="725" updateTime="1699471224025" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="97F28C3E148EED60DD67261FE2AADAB0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="7A753BC3976984DA687F65D6D6D46866">Revision of the genus Hesperopenna (Coleoptera: Chrysomelidae: Galerucinae). I. Generic redescription, definition of species groups and taxonomy of H. medvedevi species group</mods:title>
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<mods:name id="72A7259B44E9AAEF95B942AA1F575606" type="personal">
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<mods:roleTerm id="21278D7DD31AD30BEF846E14BDA2A041">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C3DFB368039ED0D2EE17C7ADB65010D1">Bezděk, Jan</mods:namePart>
<mods:affiliation id="5F86A079277FAB97D5F4CB1CB849E946">Mendel University, Department of Zoology, Zemědělská 1, CZ – 613 00 Brno, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="52C993E48DE94B40F2DC6C4F3BB086E9" type="email">bezdek@mendelu.cz</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="7468CCA266F9DE08CC4C55DF74565A4C">text</mods:typeOfResource>
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<mods:title id="5D63F684F327CEA3661D3CDB8EE2072A">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="14BA49C1BF72C13795E73E65343F2C63">
<mods:date id="524B0321DC1A4D75BEC9BA73FC6FE7E2">2013</mods:date>
<mods:detail id="F85B76B40A987FA33E0809C94FACE493" type="pubDate">
<mods:number id="B22CD600B4E55B8721522486116C5C14">2013-11-15</mods:number>
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<mods:detail id="1E0EB1A5F9D7FF2F7159AEBDB2889A2B" type="volume">
<mods:number id="739A56835090C61246BC9733DCB5DC3B">53</mods:number>
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<mods:classification id="8ED0DFA104592C0193F662F0EC81869F">journal article</mods:classification>
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<mods:identifier id="8D7E81C1E6AF867794A15581BDB1D506" type="DOI">10.5281/zenodo.5740035</mods:identifier>
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<treatment id="166387BCFFFCFFD39A0BFE17FDB76BA1" ID-DOI="http://doi.org/10.5281/zenodo.5898426" ID-GBIF-Taxon="192572522" ID-Zenodo-Dep="5898426" LSID="urn:lsid:plazi:treatment:166387BCFFFCFFD39A0BFE17FDB76BA1" httpUri="http://treatment.plazi.org/id/166387BCFFFCFFD39A0BFE17FDB76BA1" lastPageId="11" lastPageNumber="726" pageId="10" pageNumber="725">
<subSubSection id="D6D06521FFFCFFD29A0BFE17FC2E6801" box="[349,929,506,532]" pageId="10" pageNumber="725" type="nomenclature">
<paragraph id="9E7536AAFFFCFFD29A0BFE17FC2E6801" blockId="10.[349,929,506,532]" box="[349,929,506,532]" pageId="10" pageNumber="725">
<heading id="C53D81C6FFFCFFD29A0BFE17FC2E6801" bold="true" box="[349,929,506,532]" centered="true" fontSize="8" level="2" pageId="10" pageNumber="725" reason="3">
<emphasis id="ACBEEAB8FFFCFFD29A0BFE17FC2E6801" bold="true" box="[349,929,506,532]" pageId="10" pageNumber="725">
<taxonomicName id="59CA4D29FFFCFFD29A0BFE17FC906806" authority="(Kimoto, 1989)" authorityName="Bezděk" authorityYear="2013" baseAuthorityName="Kimoto" baseAuthorityYear="1989" box="[349,799,506,531]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="persimilis" status="comb. nov.">
<emphasis id="ACBEEAB8FFFCFFD29A0BFE17FDE06806" bold="true" box="[349,623,506,531]" italics="true" pageId="10" pageNumber="725">Hesperopenna persimilis</emphasis>
(
<bibRefCitation id="FA5B4B5BFFFCFFD2992AFE10FC976806" author="KIMOTO S." box="[636,792,507,531]" pageId="10" pageNumber="725" pagination="1 - 241" refId="ref15334" refString="KIMOTO S. 1989: Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. IV. Galerucinae. Esakia 27: 1 - 241." type="journal article" year="1989">Kimoto, 1989</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFCFFD29870FE17FC2E6801" box="[806,929,508,532]" pageId="10" pageNumber="725" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D6D06521FFFCFFD29ABEFDF1FC996838" box="[488,790,538,557]" pageId="10" pageNumber="725" type="description">
<paragraph id="9E7536AAFFFCFFD29ABEFDF1FC996838" blockId="10.[151,1127,538,645]" box="[488,790,538,557]" pageId="10" pageNumber="725">
(
<figureCitation id="06F12A2FFFFCFFD29AB8FDF1FDBF6838" box="[494,560,538,557]" captionStart="Figs 11–14" captionStartId="12.[151,188,1557,1576]" captionTargetBox="[158,1115,231,1431]" captionTargetId="figure-10@12.[151,1126,219,1436]" captionTargetPageId="12" captionText="Figs 11–14.Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 11 – Hesperopenna medvedevi nom.nov.; 12 – H. persimilis (Kimoto, 1989); 13 – H. tibialis (Kimoto, 1989); 14 – H. arnoldi sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740043" httpUri="https://zenodo.org/record/5740043/files/figure.png" pageId="10" pageNumber="725">Figs 12</figureCitation>
,
<figureCitation id="06F12A2FFFFCFFD2996FFDF1FDDF6838" box="[569,592,538,557]" captionStart="Figs 19–26" captionStartId="15.[112,149,1058,1077]" captionTargetBox="[124,1078,218,1029]" captionTargetId="figure-153@15.[113,1088,207,1034]" captionTargetPageId="15" captionText="Figs 19–26. Last abdominal ventrite of males.19 – Hesperopenna medvedevi nom. nov.; 20 – H. persimilis (Kimoto, 1989); 21 – H. tibialis (Kimoto, 1989); 22 – H. arnoldi sp. nov.; 23 – H. bonifaci sp. nov.; 24 – H. helferi sp. nov.; 25 – H. sipekorum sp. nov.; 26 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740049" httpUri="https://zenodo.org/record/5740049/files/figure.png" pageId="10" pageNumber="725">20</figureCitation>
,
<figureCitation id="06F12A2FFFFCFFD2990FFDF1FDFF6838" box="[601,624,538,557]" captionStart="Figs 27–33" captionStartId="17.[113,149,1518,1537]" captionTargetBox="[138,1066,210,1470]" captionTargetId="figure-14@17.[135,1067,206,1474]" captionTargetPageId="17" captionText="Figs 27–33.Last abdominal ventrite of females. 27 – Hesperopenna medvedevi nom.nov.; 28 – H.persimilis (Kimoto, 1989); 29 – H. tibialis (Kimoto, 1989); 30 – H. arnoldi sp. nov.; 31 – H. helferi sp. nov.; 32 – H. sipekorum sp. nov.; 33 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740051" httpUri="https://zenodo.org/record/5740051/files/figure.png" pageId="10" pageNumber="725">28</figureCitation>
,
<figureCitation id="06F12A2FFFFCFFD2992FFDF1FD1F6838" box="[633,656,538,557]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="10" pageNumber="725">35</figureCitation>
,
<figureCitation id="06F12A2FFFFCFFD299CFFDF1FD3F6838" box="[665,688,538,557]" captionStart="Figs 41–47" captionStartId="19.[112,149,1518,1537]" captionTargetBox="[208,996,760,1504]" captionTargetId="figure-215@19.[204,1000,749,1508]" captionTargetPageId="19" captionText="Figs 41–47. Spermatheca. 41 – Hesperopenna medvedevi nom. nov.; 42 – H. persimilis (Kimoto, 1989); 43 – H. tibialis (Kimoto, 1989); 44 – H. arnoldi sp. nov.; 45 – H. helferi sp. nov.; 46 – H. sipekorum sp. nov.; 47 – H. zofka sp. nov. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5740059" httpUri="https://zenodo.org/record/5740059/files/figure.png" pageId="10" pageNumber="725">42</figureCitation>
,
<figureCitation id="06F12A2FFFFCFFD299EFFDF1FD5F6838" box="[697,720,538,557]" captionStart="Figs 48–51" captionStartId="20.[151,188,1519,1538]" captionTargetBox="[202,1081,214,1486]" captionTargetId="figure-10@20.[196,1082,208,1487]" captionTargetPageId="20" captionText="Figs 48–51.Habitus. 48 – Hesperopenna medvedevi nom. nov.(male, Vietnam, 8.2 mm); 49 – H. persimilis (Kimoto, 1989) (male, Thailand, 6.2 mm); 50 – H. tibialis (Kimoto, 1989) (male, Laos, 9.5 mm); 51 – H. arnoldi sp. nov. (male, holotype, 8.4 mm)." figureDoi="http://doi.org/10.5281/zenodo.5740061" httpUri="https://zenodo.org/record/5740061/files/figure.png" pageId="10" pageNumber="725">49</figureCitation>
,
<figureCitation id="06F12A2FFFFCFFD2998FFDF1FC9F6838" box="[729,784,538,557]" captionStart="Figs 56–65" captionStartId="22.[152,188,1493,1512]" captionTargetBox="[193,1088,224,1464]" captionTargetId="figure-10@22.[186,1091,218,1471]" captionTargetPageId="22" captionText="Figs 56–65.Type specimens.56–57 – Hesperopenna flava Medvedev &amp; Dang, 1981 (male, holotype, 7.6 mm); 58–59 – Calomicrus persimilis Kimoto, 1989 (male, paratype, 7.1 mm); 60–61 – Martinella merkli Medvedev, 2000 (male, paratype, 8.5 mm); 62–63 – Liroetiella englerae Medvedev, 1995 (female, paratype, 5.8 mm); 64–65 – Liroetiella warisan Mohamedsaid, 1998 (female, paratype, 4.1 mm). 56, 58, 60, 62, 64 – habitus; 57, 59, 61, 63, 65 – labels." figureDoi="http://doi.org/10.5281/zenodo.5740067" httpUri="https://zenodo.org/record/5740067/files/figure.png" pageId="10" pageNumber="725">58–59</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D6D06521FFFCFFD29BC1FDD6FDDC6891" pageId="10" pageNumber="725" type="reference_group">
<paragraph id="9E7536AAFFFCFFD29BC1FDD6FE51687F" blockId="10.[151,1127,538,645]" pageId="10" pageNumber="725">
<treatmentCitationGroup id="BEDA1184FFFCFFD29BC1FDD6FE51687F" pageId="10" pageNumber="725">
<treatmentCitation id="1F6B10BBFFFCFFD29BC1FDD6FE76685A" author="KIMOTO S." box="[151,505,572,591]" page="102" pageId="10" pageNumber="725" year="1989">
<taxonomicName id="59CA4D29FFFCFFD29BC1FDD6FE76685A" authority="Kimoto, 1989: 102" authorityName="Kimoto" authorityPageNumber="102" authorityYear="1989" box="[151,505,572,591]" class="Insecta" family="Chrysomelidae" genus="Calomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="ACBEEAB8FFFCFFD29BC1FDD6FEDB685A" box="[151,340,573,591]" italics="true" pageId="10" pageNumber="725">Calomicrus persimilis</emphasis>
<bibRefCitation id="FA5B4B5BFFFCFFD29A0FFDD7FE76685A" author="KIMOTO S." box="[345,505,572,591]" pageId="10" pageNumber="725" pagination="1 - 241" refId="ref15334" refString="KIMOTO S. 1989: Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. IV. Galerucinae. Esakia 27: 1 - 241." type="journal article" year="1989">Kimoto, 1989: 102</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description);
<treatmentCitation id="1F6B10BBFFFCFFD299EBFDD6FC12685A" author="WARCHALOWSKI A." box="[701,925,572,592]" page="50" pageId="10" pageNumber="725" year="1991">
<bibRefCitation id="FA5B4B5BFFFCFFD299EBFDD6FC0F685A" author="WARCHALOWSKI A." box="[701,896,572,592]" pageId="10" pageNumber="725" pagination="41 - 55" refId="ref16366" refString="WARCHALOWSKI A. 1991: Kurze Ubersicht der oberseits hellen Calomicrus Steph. - Arten (Coleptera: Chrysomelidae: Galerucinae). Genus 2: 41 - 55." type="journal article" year="1991">WARCHAŁOWSKI (1991)</bibRefCitation>
: 50
</treatmentCitation>
(key);
<treatmentCitation id="1F6B10BBFFFCFFD2988CFDD7FEFD687F" author="MOHAMEDSAID M. S. &amp; CONSTANT J." page="167" pageId="10" pageNumber="725" year="2007">
<bibRefCitation id="FA5B4B5BFFFCFFD2988CFDD7FEC6687F" author="MOHAMEDSAID M. S. &amp; CONSTANT J." pageId="10" pageNumber="725" pagination="163 - 177" refId="ref15946" refString="MOHAMEDSAID M. S. &amp; CONSTANT J. 2007: Chrysomelid beetles of the subfamily Galerucinae from Thailand and Cambodia in the collections of the Royal Belgian Institute of Natural Sciences (Coleoptera: Chrysomelidae). Bulletin de l'Institut Royal des Sciences Naturelles de Belgique (Entomologie) 77: 163 - 177." type="journal article" year="2007">MOHAMEDSAID &amp; CONSTANT (2007)</bibRefCitation>
: 167
</treatmentCitation>
(faunistics).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9E7536AAFFFCFFD29BC1FD99FDDC6891" blockId="10.[151,1127,538,645]" box="[151,595,625,645]" pageId="10" pageNumber="725">
<treatmentCitationGroup id="BEDA1184FFFCFFD29BC1FD99FDDC6891" box="[151,595,625,645]" pageId="10" pageNumber="725">
<treatmentCitation id="1F6B10BBFFFCFFD29BC1FD99FD956891" author="MEDVEDEV L. N." box="[151,538,625,645]" page="169" pageId="10" pageNumber="725" year="2000">
<taxonomicName id="59CA4D29FFFCFFD29BC1FD99FE7D6891" authority=": MEDVEDEV (2000)" authorityName="MEDVEDEV" authorityYear="2000" box="[151,498,625,645]" class="Insecta" family="Bignoniaceae" genus="Martinella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="ACBEEAB8FFFCFFD29BC1FD99FEC36891" box="[151,332,626,644]" italics="true" pageId="10" pageNumber="725">Martinella persimilis</emphasis>
:
<bibRefCitation id="FA5B4B5BFFFCFFD29A01FD99FE7D6891" author="MEDVEDEV L. N." box="[343,498,625,645]" pageId="10" pageNumber="725" pagination="161 - 182" refId="ref15477" refString="MEDVEDEV L. N. 2000: Chrysomelidae (Coleoptera) of Laos from the collection of the Hungarian Natural History Museum. Annales Historico-Naturales Musei Nationalis Hungarici 92: 161 - 182." type="journal article" year="2000">MEDVEDEV (2000)</bibRefCitation>
</taxonomicName>
: 169
</treatmentCitation>
(key).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D6D06521FFFCFFD29BC1FD70FBB8698B" pageId="10" pageNumber="725" type="materials_examined">
<paragraph id="9E7536AAFFFCFFD29BC1FD70FD8468BB" blockId="10.[151,1127,666,1583]" box="[151,523,666,687]" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD29BC1FD70FE9F68BB" bold="true" box="[151,272,666,686]" pageId="10" pageNumber="725">
<typeStatus id="41718808FFFCFFD29BC1FD70FF4B68BB" box="[151,196,667,686]" pageId="10" pageNumber="725">Type</typeStatus>
locality.
</emphasis>
‘
<collectingCountry id="E6DD763AFFFCFFD29A4FFD70FEC668BB" box="[281,329,667,686]" name="Laos" pageId="10" pageNumber="725">Laos</collectingCountry>
: Sedone Prov., Pakse’.
</paragraph>
<paragraph id="9E7536AAFFFCFFD29BC1FD5DFCDB695B" blockId="10.[151,1127,666,1583]" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD29BC1FD5DFEF668DC" bold="true" box="[151,377,693,713]" pageId="10" pageNumber="725">Type material examined.</emphasis>
<materialsCitation id="2EA23CF7FFFCFFD29A28FD5DFE4068EB" ID-GBIF-Occurrence="3451920328" collectingDate="1996-05-31" collectionCode="BPBM" country="Laos" location="Sedone Prov." municipality="Pakse" pageId="10" pageNumber="725" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="41718808FFFCFFD29A28FD5DFE5E68DD" box="[382,465,694,713]" pageId="10" pageNumber="725" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="88CCFD23FFFCFFD29A8BFD5AFE6268DF" box="[477,493,689,714]" pageId="10" pageNumber="725" type="male">♂</specimenCount>
, ‘
<collectingCountry id="E6DD763AFFFCFFD29AADFD5DFDB568DC" box="[507,570,694,713]" name="Laos" pageId="10" pageNumber="725">LAOS</collectingCountry>
: /
<location id="9B156071FFFCFFD2991BFD5DFD3168DC" LSID="urn:lsid:plazi:treatment:166387BCFFFCFFD39A0BFE17FDB76BA1:9B156071FFFCFFD2991BFD5DFD3168DC" box="[589,702,694,713]" country="Laos" county="Native Collector" municipality="Pakse" name="Sedone Prov." pageId="10" pageNumber="725">Sedone Prov.</location>
/
<collectingMunicipality id="7E11ACD0FFFCFFD29998FD5DFC8F68DC" box="[718,768,694,713]" pageId="10" pageNumber="725">Pakse</collectingMunicipality>
/ 31.V.196 [p] 5 [w, h] // Native Collector / BISHOP MUS. [w, p] // HOLOTYPE [red label, p] //
<taxonomicName id="59CA4D29FFFCFFD29939FD3BFD5E68F6" authorityName="Dillwyn" authorityYear="1829" box="[623,721,720,739]" class="Insecta" family="Chrysomelidae" genus="Calomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus">Calomicrus</taxonomicName>
/ persimilis / n. sp. [w, h] // PHOTO [red label, p] // 14964 [pink label, h]’ (
<collectionCode id="F8DBAE6FFFFCFFD29AD3FD00FE4A68EB" box="[389,453,747,766]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="10" pageNumber="725" type="Museum">BPBM</collectionCode>
).
</materialsCitation>
<materialsCitation id="2EA23CF7FFFCFFD29A82FD07FF706921" ID-GBIF-Occurrence="3451920310" collectionCode="BPBM" country="Laos" county="Native Collector" location="Sedone Prov." municipality="Pakse" pageId="10" pageNumber="725" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="41718808FFFCFFD29A82FD07FDA468E8" box="[468,555,748,767]" pageId="10" pageNumber="725" type="paratype">PARATYPES</typeStatus>
: 1 ♂
<specimenCount id="88CCFD23FFFCFFD2990AFD00FDF368EB" box="[604,636,741,766]" pageId="10" pageNumber="725" type="female">1 ♀</specimenCount>
, ‘
<collectingCountry id="E6DD763AFFFCFFD299DCFD00FD4668EB" box="[650,713,747,766]" name="Laos" pageId="10" pageNumber="725">LAOS</collectingCountry>
: /
<location id="9B156071FFFCFFD2998BFD00FCC068EA" LSID="urn:lsid:plazi:treatment:166387BCFFFCFFD39A0BFE17FDB76BA1:9B156071FFFCFFD2998BFD00FCC068EA" box="[733,847,747,767]" country="Laos" county="Native Collector" municipality="Pakse" name="Sedone Prov." pageId="10" pageNumber="725">Sedone Prov.</location>
/
<collectingMunicipality id="7E11ACD0FFFCFFD29809FD00FC1E68EB" box="[863,913,747,766]" pageId="10" pageNumber="725">Pakse</collectingMunicipality>
/ 31.V.196 [p] 5 [w, h] //
<collectingCounty id="77144E26FFFCFFD29BC1FCEDFEAA690C" box="[151,293,774,793]" pageId="10" pageNumber="725">Native Collector</collectingCounty>
/ BISHOP MUS. [w, p] // PARATOPOTYPE [blue label, p] //
<taxonomicName id="59CA4D29FFFCFFD29869FCEDFC2E690C" authorityName="Dillwyn" authorityYear="1829" box="[831,929,774,793]" class="Insecta" family="Chrysomelidae" genus="Calomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus">Calomicrus</taxonomicName>
/ persimilis / n. sp. [w, h]’ (
<collectionCode id="F8DBAE6FFFFCFFD29BECFCCAFF756921" box="[186,250,801,820]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="10" pageNumber="725" type="Museum">BPBM</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="2EA23CF7FFFCFFD29A5EFCCAFCDB695B" ID-GBIF-Occurrence="3451920347" collectingDate="1965-04-17" collectorName="de Khong &amp; J. L. Gressitt" country="Laos" location="Ile de Khong" pageId="10" pageNumber="725" specimenCount="1" typeStatus="paratype">
2 ♂♂, ‘
<collectingCountry id="E6DD763AFFFCFFD29A12FCCAFE0C6921" box="[324,387,801,820]" name="Laos" pageId="10" pageNumber="725">LAOS</collectingCountry>
: /
<location id="9B156071FFFCFFD29AC3FCCBFD8B6926" LSID="urn:lsid:plazi:treatment:166387BCFFFCFFD39A0BFE17FDB76BA1:9B156071FFFCFFD29AC3FCCBFD8B6926" box="[405,516,800,819]" country="Laos" name="Ile de Khong" pageId="10" pageNumber="725">
Ile
<collectorName id="333F537CFFFCFFD29AE7FCCBFD8B6926" box="[433,516,800,819]" pageId="10" pageNumber="725">de Khong</collectorName>
</location>
/
<date id="EA74106AFFFCFFD29942FCCBFDFD6926" box="[532,626,800,819]" pageId="10" pageNumber="725" value="1965-04-17">
<collectingDate id="FA30E982FFFCFFD29942FCCBFDFD6926" box="[532,626,800,819]" pageId="10" pageNumber="725" value="1965-04-17">17.IV.1965</collectingDate>
</date>
[w, p] //
<collectorName id="333F537CFFFCFFD299E8FCCAFCA56926" box="[702,810,800,820]" pageId="10" pageNumber="725">J. L. Gressitt</collectorName>
/ Collector / BISHOP MUSEUM [w, p] //
<typeStatus id="41718808FFFCFFD29BE9FCD0FEA6695B" box="[191,297,827,846]" pageId="10" pageNumber="725" type="paratype">PARATYPE</typeStatus>
[blue label, p] //
<taxonomicName id="59CA4D29FFFCFFD29AEAFCD0FD91695B" authorityName="Dillwyn" authorityYear="1829" box="[444,542,827,846]" class="Insecta" family="Chrysomelidae" genus="Calomicrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="741" phylum="Arthropoda" rank="genus">Calomicrus</taxonomicName>
/ persimilis / n. sp. [w, h]’ (
<collectionCode id="F8DBAE6FFFFCFFD2985CFCD0FCC5695B" box="[778,842,827,846]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" lsid="urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="10" pageNumber="725" type="Museum">BPBM</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="9E7536AAFFFCFFD29BC1FCBEFBB8698B" blockId="10.[151,1127,666,1583]" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD29BC1FCBEFE2A697C" bold="true" box="[151,421,853,873]" pageId="10" pageNumber="725">Additional material examined</emphasis>
(
<specimenCount id="88CCFD23FFFCFFD29AE6FCBDFE77697C" box="[432,504,854,873]" pageId="10" pageNumber="725" type="generic">11 spec.</specimenCount>
).
<materialsCitation id="2EA23CF7FFFCFFD29952FCBEFD9C6991" ID-GBIF-Occurrence="3451920342" latitude="16.866667" longitude="101.816666" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD29952FCBEFD65697C" bold="true" box="[516,746,853,873]" pageId="10" pageNumber="725">
<collectingCountry id="E6DD763AFFFCFFD29952FCBEFDF5697C" box="[516,634,853,873]" name="Thailand" pageId="10" pageNumber="725">THAILAND</collectingCountry>
:
<collectingRegion id="5C0EF848FFFCFFD299D4FCBDFD24697D" box="[642,683,854,873]" country="Thailand" name="Loei" pageId="10" pageNumber="725">LOEI</collectingRegion>
PROV.:
</emphasis>
Phu Kradung N. P.,
<geoCoordinate id="FBFE506DFFFCFFD298C5FCBDFC53697C" box="[915,988,854,873]" degrees="16" direction="north" minutes="52" orientation="latitude" pageId="10" pageNumber="725" precision="925" value="16.866667">16°52´N</geoCoordinate>
<geoCoordinate id="FBFE506DFFFCFFD298B6FCBDFBBA697C" box="[992,1077,854,873]" degrees="101" direction="east" minutes="49" orientation="longitude" pageId="10" pageNumber="725" precision="925" value="101.816666">101°49´E</geoCoordinate>
,
<quantity id="59329B4FFFFCFFD29F6AFCBDFF256991" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="725" unit="m" value="1000.0">1000 m</quantity>
,
<date id="EA74106AFFFCFFD29BE4FC9BFEA56996" box="[178,298,880,899]" pageId="10" pageNumber="725" value="1999-05-16" valueMax="1999-05-17" valueMin="1999-05-16">16.-17.v.1999</date>
,
<specimenCount id="88CCFD23FFFCFFD29A64FC9AFEDD6991" box="[306,338,875,900]" pageId="10" pageNumber="725" type="female">1 ♀</specimenCount>
, M. Říha leg. (
<collectionCode id="F8DBAE6FFFFCFFD29A84FC9AFD866991" box="[466,521,881,900]" pageId="10" pageNumber="725">JBCB</collectionCode>
).
</materialsCitation>
<materialsCitation id="2EA23CF7FFFCFFD29941FC9AFD9A698B" ID-GBIF-Occurrence="3451920305" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD29941FC9AFD316991" bold="true" box="[535,702,881,900]" pageId="10" pageNumber="725">
<collectingRegion id="5C0EF848FFFCFFD29941FC9AFD316991" box="[535,702,881,900]" country="Thailand" name="Phetchabun" pageId="10" pageNumber="725">PHETCHABUN PROV.</collectingRegion>
</emphasis>
: Lom Sak – Dan Sai,
<date id="EA74106AFFFCFFD2982CFC9AFC796991" box="[890,1014,881,900]" pageId="10" pageNumber="725" value="1993-05-17" valueMax="1993-05-19" valueMin="1993-05-17">17.–19.v.1993</date>
, 6 ♂♂
<specimenCount id="88CCFD23FFFCFFD29F64FC9AFBEE6991" box="[1074,1121,875,900]" pageId="10" pageNumber="725" type="female">3 ♀♀</specimenCount>
, Pacholátko &amp; Dembický leg. (
<collectionCode id="F8DBAE6FFFFCFFD29ACCFC60FE5F698B" box="[410,464,907,926]" pageId="10" pageNumber="725">JBCB</collectionCode>
,
<collectionCode id="F8DBAE6FFFFCFFD29A8FFC60FD84698B" box="[473,523,907,926]" pageId="10" pageNumber="725">JVCJ</collectionCode>
).
</materialsCitation>
<materialsCitation id="2EA23CF7FFFCFFD2994FFC61FBB8698B" ID-GBIF-Occurrence="3451920351" box="[537,1079,902,927]" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD2994FFC61FD77698B" bold="true" box="[537,760,906,926]" pageId="10" pageNumber="725">
<collectingCountry id="E6DD763AFFFCFFD2994FFC61FDD9698B" box="[537,598,906,926]" name="Laos" pageId="10" pageNumber="725">LAOS</collectingCountry>
:
<collectingRegion id="5C0EF848FFFCFFD29909FC60FD366988" box="[607,697,907,926]" country="Laos" name="Viangchan" pageId="10" pageNumber="725">VIENTIANE</collectingRegion>
PROV.:
</emphasis>
Vientiane, 1 ♂, Vitalis leg. (
<collectionCode id="F8DBAE6FFFFCFFD298BDFC60FBA3698B" box="[1003,1068,907,926]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="10" pageNumber="725">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D6D06521FFFCFFD29BC1FC5DFEEB6F9A" pageId="10" pageNumber="725" type="description">
<paragraph id="9E7536AAFFFCFFD29BC1FC5DFCA869DB" blockId="10.[151,1127,666,1583]" box="[151,807,944,975]" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD29BC1FC5DFEAD69DB" bold="true" box="[151,290,950,974]" pageId="10" pageNumber="725">Description.</emphasis>
Body length: ♂♂ 6.6–7.0 mm,
<specimenCount id="88CCFD23FFFCFFD29925FC5BFD1769DA" box="[627,664,944,975]" pageId="10" pageNumber="725" type="female">♀♀</specimenCount>
<quantity id="59329B4FFFFCFFD299C9FC5DFCAC69DB" box="[671,803,950,974]" metricMagnitude="-3" metricUnit="m" metricValue="7.35" metricValueMax="7.5" metricValueMin="7.2" pageId="10" pageNumber="725" unit="mm" value="7.35" valueMax="7.5" valueMin="7.2">7.2–7.5 mm</quantity>
.
</paragraph>
<paragraph id="9E7536AAFFFCFFD29BE3FC3CFF576E1A" blockId="10.[151,1127,666,1583]" pageId="10" pageNumber="725">
Male (
<figureCitation id="06F12A2FFFFCFFD29BA9FC3CFEDE69FA" box="[255,337,983,1007]" captionStart="Figs 48–51" captionStartId="20.[151,188,1519,1538]" captionTargetBox="[202,1081,214,1486]" captionTargetId="figure-10@20.[196,1082,208,1487]" captionTargetPageId="20" captionText="Figs 48–51.Habitus. 48 – Hesperopenna medvedevi nom. nov.(male, Vietnam, 8.2 mm); 49 – H. persimilis (Kimoto, 1989) (male, Thailand, 6.2 mm); 50 – H. tibialis (Kimoto, 1989) (male, Laos, 9.5 mm); 51 – H. arnoldi sp. nov. (male, holotype, 8.4 mm)." figureDoi="http://doi.org/10.5281/zenodo.5740061" httpUri="https://zenodo.org/record/5740061/files/figure.png" pageId="10" pageNumber="725">Fig. 49</figureCitation>
,
<typeStatus id="41718808FFFCFFD29A36FC3CFE3269FA" box="[352,445,983,1007]" pageId="10" pageNumber="725" type="paratype">paratype</typeStatus>
<figureCitation id="06F12A2FFFFCFFD29A91FC3CFD9569FB" box="[455,538,982,1007]" captionStart="Figs 56–65" captionStartId="22.[152,188,1493,1512]" captionTargetBox="[193,1088,224,1464]" captionTargetId="figure-10@22.[186,1091,218,1471]" captionTargetPageId="22" captionText="Figs 56–65.Type specimens.56–57 – Hesperopenna flava Medvedev &amp; Dang, 1981 (male, holotype, 7.6 mm); 58–59 – Calomicrus persimilis Kimoto, 1989 (male, paratype, 7.1 mm); 60–61 – Martinella merkli Medvedev, 2000 (male, paratype, 8.5 mm); 62–63 – Liroetiella englerae Medvedev, 1995 (female, paratype, 5.8 mm); 64–65 – Liroetiella warisan Mohamedsaid, 1998 (female, paratype, 4.1 mm). 56, 58, 60, 62, 64 – habitus; 57, 59, 61, 63, 65 – labels." figureDoi="http://doi.org/10.5281/zenodo.5740067" httpUri="https://zenodo.org/record/5740067/files/figure.png" pageId="10" pageNumber="725">Fig. 58</figureCitation>
). Body completely orange, only apices of mandibles black.
</paragraph>
<paragraph id="9E7536AAFFFCFFD29BE3FBFCFD466EDA" blockId="10.[151,1127,666,1583]" pageId="10" pageNumber="725">
Interocular space 1.5 times as wide as transverse diameter of eye. Antennae 0.80 times as long as body, length ratio of antennomeres I to XI equals 11-4-5-13-12-12-12-13-12-12- 13. Pronotum 1.42
<emphasis id="ACBEEAB8FFFCFFD29A08FBBCFEE46E7A" bold="true" box="[350,363,1111,1135]" pageId="10" pageNumber="725">–</emphasis>
1.54 times as broad as long. Elytra glabrous, 1.75 times as long as wide (measured at humeral calli), 0.70 times as long as body. Length ratio of metatarsomeres I–IV equals 9-4-4-7. Last ventrite with two very small incisions posteriorly, depression on the disc anteriorly surrounded with U-shaped ridge (
<figureCitation id="06F12A2FFFFCFFD2993BFB5CFD336EDA" box="[621,700,1207,1231]" captionStart="Figs 19–26" captionStartId="15.[112,149,1058,1077]" captionTargetBox="[124,1078,218,1029]" captionTargetId="figure-153@15.[113,1088,207,1034]" captionTargetPageId="15" captionText="Figs 19–26. Last abdominal ventrite of males.19 – Hesperopenna medvedevi nom. nov.; 20 – H. persimilis (Kimoto, 1989); 21 – H. tibialis (Kimoto, 1989); 22 – H. arnoldi sp. nov.; 23 – H. bonifaci sp. nov.; 24 – H. helferi sp. nov.; 25 – H. sipekorum sp. nov.; 26 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740049" httpUri="https://zenodo.org/record/5740049/files/figure.png" pageId="10" pageNumber="725">Fig. 20</figureCitation>
).
</paragraph>
<paragraph id="9E7536AAFFFCFFD29BE3FB3CFDF96F3A" blockId="10.[151,1127,666,1583]" pageId="10" pageNumber="725">
Aedeagus: dorsal side with deep asymmetrical incision on left side, on right side with short rounded process and wide shallow subtriangular incision; ventral side gradually tapering to triangular apex, ventral surface flat (
<figureCitation id="06F12A2FFFFCFFD2994CFAFCFDE56F3A" box="[538,618,1303,1327]" captionStart="Figs 11–14" captionStartId="12.[151,188,1557,1576]" captionTargetBox="[158,1115,231,1431]" captionTargetId="figure-10@12.[151,1126,219,1436]" captionTargetPageId="12" captionText="Figs 11–14.Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 11 – Hesperopenna medvedevi nom.nov.; 12 – H. persimilis (Kimoto, 1989); 13 – H. tibialis (Kimoto, 1989); 14 – H. arnoldi sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740043" httpUri="https://zenodo.org/record/5740043/files/figure.png" pageId="10" pageNumber="725">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="9E7536AAFFFCFFD29BE3FADCFEEB6F9A" blockId="10.[151,1127,666,1583]" pageId="10" pageNumber="725">
Female. Apex of last ventrite widely rounded (
<figureCitation id="06F12A2FFFFCFFD299CDFADCFD666F5A" box="[667,745,1335,1359]" captionStart="Figs 27–33" captionStartId="17.[113,149,1518,1537]" captionTargetBox="[138,1066,210,1470]" captionTargetId="figure-14@17.[135,1067,206,1474]" captionTargetPageId="17" captionText="Figs 27–33.Last abdominal ventrite of females. 27 – Hesperopenna medvedevi nom.nov.; 28 – H.persimilis (Kimoto, 1989); 29 – H. tibialis (Kimoto, 1989); 30 – H. arnoldi sp. nov.; 31 – H. helferi sp. nov.; 32 – H. sipekorum sp. nov.; 33 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740051" httpUri="https://zenodo.org/record/5740051/files/figure.png" pageId="10" pageNumber="725">Fig. 28</figureCitation>
), pygidium with obtusangulate apex (
<figureCitation id="06F12A2FFFFCFFD29BC9FABCFF606F7B" box="[159,239,1366,1391]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="10" pageNumber="725">Fig. 35</figureCitation>
). Spermatheca: nodulus subglobular, cornu regularly rounded with very long apical appendix (
<figureCitation id="06F12A2FFFFCFFD29A5EFA9CFED76F9A" box="[264,344,1399,1423]" captionStart="Figs 41–47" captionStartId="19.[112,149,1518,1537]" captionTargetBox="[208,996,760,1504]" captionTargetId="figure-215@19.[204,1000,749,1508]" captionTargetPageId="19" captionText="Figs 41–47. Spermatheca. 41 – Hesperopenna medvedevi nom. nov.; 42 – H. persimilis (Kimoto, 1989); 43 – H. tibialis (Kimoto, 1989); 44 – H. arnoldi sp. nov.; 45 – H. helferi sp. nov.; 46 – H. sipekorum sp. nov.; 47 – H. zofka sp. nov. Scale bar 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.5740059" httpUri="https://zenodo.org/record/5740059/files/figure.png" pageId="10" pageNumber="725">Fig. 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D6D06521FFFCFFD39BC1FA7DFE296B61" lastPageId="11" lastPageNumber="726" pageId="10" pageNumber="725" type="diagnosis">
<paragraph id="9E7536AAFFFCFFD39BC1FA7DFE296B61" blockId="10.[151,1127,666,1583]" lastBlockId="11.[113,1089,220,436]" lastPageId="11" lastPageNumber="726" pageId="10" pageNumber="725">
<emphasis id="ACBEEAB8FFFCFFD29BC1FA7DFE196FBB" bold="true" box="[151,406,1430,1454]" pageId="10" pageNumber="725">Differential diagnosis.</emphasis>
Due to glabrous elytra and uniformly brown legs
<taxonomicName id="59CA4D29FFFCFFD2989DFA73FE8F6FDA" authorityName="Bezděk" authorityYear="2013" baseAuthorityName="Kimoto" baseAuthorityYear="1989" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="ACBEEAB8FFFCFFD2989DFA73FE8F6FDA" italics="true" pageId="10" pageNumber="725">Hesperopenna persimilis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="59CA4D29FFFCFFD29ACEFA53FD8D6FDA" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[408,514,1464,1487]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFCFFD29ACEFA53FD8D6FDA" box="[408,514,1464,1487]" italics="true" pageId="10" pageNumber="725">H. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFCFFD2995DFA5CFDEE6FDB" box="[523,609,1463,1486]" pageId="10" pageNumber="725" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="59CA4D29FFFCFFD299CDFA53FC9A6FDA" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[667,789,1464,1487]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="bonifaci" status="sp. nov.">
<emphasis id="ACBEEAB8FFFCFFD299CDFA53FC9A6FDA" box="[667,789,1464,1487]" italics="true" pageId="10" pageNumber="725">H. bonifaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFCFFD29876FA5CFCFA6FDB" box="[800,885,1463,1486]" pageId="10" pageNumber="725" rank="species">sp. nov.</taxonomicNameLabel>
but differs from both species in the subapical impression on the last ventrite in male, anteriorly surrounded with an U-shaped ridge (without ridge in
<taxonomicName id="59CA4D29FFFCFFD29973FA13FD026C1A" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[549,653,1528,1551]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFCFFD29973FA13FD026C1A" box="[549,653,1528,1551]" italics="true" pageId="10" pageNumber="725">H. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFCFFD299C3FA1CFD666C1B" box="[661,745,1527,1550]" pageId="10" pageNumber="725" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="59CA4D29FFFCFFD29849FA13FC166C1A" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[799,921,1528,1551]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="725" phylum="Arthropoda" rank="species" species="bonifaci" status="sp. nov.">
<emphasis id="ACBEEAB8FFFCFFD29849FA13FC166C1A" box="[799,921,1528,1551]" italics="true" pageId="10" pageNumber="725">H. bonifaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFCFFD298F7FA1CFC786C1A" box="[929,1015,1527,1551]" pageId="10" pageNumber="725" rank="species">sp. nov.</taxonomicNameLabel>
) (
<figureCitation id="06F12A2FFFFCFFD29F5BFA1CFF536C3A" captionStart="Figs 19–26" captionStartId="15.[112,149,1058,1077]" captionTargetBox="[124,1078,218,1029]" captionTargetId="figure-153@15.[113,1088,207,1034]" captionTargetPageId="15" captionText="Figs 19–26. Last abdominal ventrite of males.19 – Hesperopenna medvedevi nom. nov.; 20 – H. persimilis (Kimoto, 1989); 21 – H. tibialis (Kimoto, 1989); 22 – H. arnoldi sp. nov.; 23 – H. bonifaci sp. nov.; 24 – H. helferi sp. nov.; 25 – H. sipekorum sp. nov.; 26 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740049" httpUri="https://zenodo.org/record/5740049/files/figure.png" pageId="10" pageNumber="725">Figs 20, 23, 24</figureCitation>
) and simple triangular apex of the ventral side of aedeagus (ventral side of aedeagus asymmetrical with a deep U-shaped incision in
<taxonomicName id="59CA4D29FFFDFFD39922FF36FD536AE1" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[628,732,221,244]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFDFFD39922FF36FD536AE1" box="[628,732,221,244]" italics="true" pageId="11" pageNumber="726">H. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFDFFD399B5FF37FCB86AE6" box="[739,823,220,243]" pageId="11" pageNumber="726" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="59CA4D29FFFDFFD3983BFF36FC6A6AE1" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[877,997,221,244]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="bonifaci" status="sp. nov.">
<emphasis id="ACBEEAB8FFFDFFD3983BFF36FC6A6AE1" box="[877,997,221,244]" italics="true" pageId="11" pageNumber="726">H. bonifaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFDFFD398BBFF37FBCE6AE6" box="[1005,1089,220,243]" pageId="11" pageNumber="726" rank="species">sp. nov.</taxonomicNameLabel>
– incision on the right side in
<taxonomicName id="59CA4D29FFFDFFD39AFFFF16FD806B01" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[425,527,253,276]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFDFFD39AFFFF16FD806B01" box="[425,527,253,276]" italics="true" pageId="11" pageNumber="726">H. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFDFFD39942FF17FDE76B06" box="[532,616,252,275]" pageId="11" pageNumber="726" rank="species">sp. nov.</taxonomicNameLabel>
, on the left side in
<taxonomicName id="59CA4D29FFFDFFD3987BFF16FC2B6B01" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[813,932,253,276]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="bonifaci" status="sp. nov.">
<emphasis id="ACBEEAB8FFFDFFD3987BFF16FC2B6B01" box="[813,932,253,276]" italics="true" pageId="11" pageNumber="726">H. bonifaci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFDFFD398FCFF17FC726B01" box="[938,1021,252,276]" pageId="11" pageNumber="726" rank="species">sp. nov.</taxonomicNameLabel>
) (
<figureCitation id="06F12A2FFFFDFFD39F47FF17FF016B21" captionStart="Figs 11–14" captionStartId="12.[151,188,1557,1576]" captionTargetBox="[158,1115,231,1431]" captionTargetId="figure-10@12.[151,1126,219,1436]" captionTargetPageId="12" captionText="Figs 11–14.Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 11 – Hesperopenna medvedevi nom.nov.; 12 – H. persimilis (Kimoto, 1989); 13 – H. tibialis (Kimoto, 1989); 14 – H. arnoldi sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740043" httpUri="https://zenodo.org/record/5740043/files/figure.png" pageId="11" pageNumber="726">Figs 12</figureCitation>
,
<figureCitation id="06F12A2FFFFDFFD39BCCFEF0FF6D6B26" box="[154,226,283,307]" captionStart="Figs 15–18" captionStartId="13.[115,152,1542,1561]" captionTargetBox="[126,1079,226,1362]" captionTargetId="figure-14@13.[113,1090,218,1370]" captionTargetPageId="13" captionText="Figs 15–18. Aedeagus (a – dorsal view; b – lateral view; c – ventral view). 15 – H. bonifaci sp. nov.; 16 – H. helferi sp. nov.; 17 – H. sipekorum sp. nov.; 18 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740045" httpUri="https://zenodo.org/record/5740045/files/figure.png" pageId="11" pageNumber="726">15, 16</figureCitation>
). Females of
<taxonomicName id="59CA4D29FFFDFFD39A22FEF6FE706B21" authorityName="Bezděk" authorityYear="2013" baseAuthorityName="Kimoto" baseAuthorityYear="1989" box="[372,511,285,308]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="ACBEEAB8FFFDFFD39A22FEF6FE706B21" box="[372,511,285,308]" italics="true" pageId="11" pageNumber="726">H. persimilis</emphasis>
</taxonomicName>
and
<taxonomicName id="59CA4D29FFFDFFD39963FEF6FD126B21" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" box="[565,669,285,308]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFDFFD39963FEF6FD126B21" box="[565,669,285,308]" italics="true" pageId="11" pageNumber="726">H. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFDFFD399F2FEF7FD776B26" box="[676,760,284,307]" pageId="11" pageNumber="726" rank="species">sp. nov.</taxonomicNameLabel>
can be separated by the shape of pygidium: apex obtusangulate in
<taxonomicName id="59CA4D29FFFDFFD39AB8FED6FDF86B41" authorityName="Bezděk" authorityYear="2013" baseAuthorityName="Kimoto" baseAuthorityYear="1989" box="[494,631,317,340]" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="persimilis">
<emphasis id="ACBEEAB8FFFDFFD39AB8FED6FDF86B41" box="[494,631,317,340]" italics="true" pageId="11" pageNumber="726">H. persimilis</emphasis>
</taxonomicName>
while with a small U-shaped incision in
<taxonomicName id="59CA4D29FFFDFFD39F71FED6FF386B61" authority="Bezděk, 2013" authorityName="Bezděk" authorityYear="2013" class="Insecta" family="Chrysomelidae" genus="Hesperopenna" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="726" phylum="Arthropoda" rank="species" species="helferi" status="sp. nov.">
<emphasis id="ACBEEAB8FFFDFFD39F71FED6FF386B61" italics="true" pageId="11" pageNumber="726">H. helferi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B78D57C3FFFDFFD39BE8FEB7FE9F6B66" box="[190,272,348,371]" pageId="11" pageNumber="726" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="06F12A2FFFFDFFD39A48FEB7FE156B61" box="[286,410,347,372]" captionStart="Figs 34–40" captionStartId="18.[151,188,1519,1538]" captionTargetBox="[231,1053,221,1476]" captionTargetId="figure-10@18.[218,1059,210,1480]" captionTargetPageId="18" captionText="Figs 34–40. Female pygidium. 34 – Hesperopenna medvedevi nom. nov.; 35 – H. persimilis (Kimoto, 1989); 36 – H. tibialis (Kimoto, 1989); 37 – H. arnoldi sp. nov.; 38 – H. helferi sp. nov.; 39 – H. sipekorum sp. nov.; 40 – H. zofka sp. nov. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5740055" httpUri="https://zenodo.org/record/5740055/files/figure.png" pageId="11" pageNumber="726">Figs 35, 38</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D6D06521FFFDFFD39B27FE90FDB76BA1" pageId="11" pageNumber="726" type="distribution">
<paragraph id="9E7536AAFFFDFFD39B27FE90FDB76BA1" blockId="11.[113,1089,220,436]" pageId="11" pageNumber="726">
<emphasis id="ACBEEAB8FFFDFFD39B27FE90FE8B6B86" bold="true" box="[113,260,379,403]" pageId="11" pageNumber="726">Distribution.</emphasis>
<collectingCountry id="E6DD763AFFFDFFD39A5FFE97FEB26B86" box="[265,317,380,403]" name="Laos" pageId="11" pageNumber="726">Laos</collectingCountry>
(
<bibRefCitation id="FA5B4B5BFFFDFFD39A1FFE97FE576B81" author="KIMOTO S." box="[329,472,380,404]" pageId="11" pageNumber="726" pagination="1 - 241" refId="ref15334" refString="KIMOTO S. 1989: Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. IV. Galerucinae. Esakia 27: 1 - 241." type="journal article" year="1989">KIMOTO 1989</bibRefCitation>
, present paper),
<collectingCountry id="E6DD763AFFFDFFD399D5FE97FD526B81" box="[643,733,380,404]" name="Vietnam" pageId="11" pageNumber="726">Vietnam</collectingCountry>
(
<bibRefCitation id="FA5B4B5BFFFDFFD399BCFE97FCF56B81" author="KIMOTO S." box="[746,890,380,404]" pageId="11" pageNumber="726" pagination="1 - 241" refId="ref15334" refString="KIMOTO S. 1989: Chrysomelidae (Coleoptera) of Thailand, Cambodia, Laos and Vietnam. IV. Galerucinae. Esakia 27: 1 - 241." type="journal article" year="1989">KIMOTO 1989</bibRefCitation>
),
<collectingCountry id="E6DD763AFFFDFFD398DDFE97FC656B81" box="[907,1002,380,404]" name="Thailand" pageId="11" pageNumber="726">Thailand</collectingCountry>
(MOHAMEDSAID &amp; CONSTANT 2007, present paper).
</paragraph>
</subSubSection>
</treatment>
</document>
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