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<document ID-DOI="10.5733/afin.053.0209" ID-ISSN="2305-2562" ID-Zenodo-Dep="7918274" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1683581517355" checkinUser="felipe" docAuthor="Herbert, D. G." docDate="2012" docId="0D38CE4BC27DF71EA2FB7402FDAEFBB2" docLanguage="en" docName="AfrInvertebr.53.2.381-502.pdf" docOrigin="African Invertebrates 53 (2)" docSource="http://www.bioone.org/doi/10.5733/afin.053.0209" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Pholidotrope gloriosa Herbert 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="480" masterDocId="F101B633C21FF77DA3357529FFF4FFB0" masterDocTitle="A revision of the Chilodontidae (Gastropoda: Vetigastropoda: Seguenzioidea) of southern Africa and the south-western Indian Ocean" masterLastPageNumber="381" masterPageNumber="381" pageNumber="479" updateTime="1683700561700" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A revision of the Chilodontidae (Gastropoda: Vetigastropoda: Seguenzioidea) of southern Africa and the south-western Indian Ocean</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Herbert, D. G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>African Invertebrates</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-11-06</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>381</mods:start>
<mods:end>381</mods:end>
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<mods:url>http://www.bioone.org/doi/10.5733/afin.053.0209</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5733/afin.053.0209</mods:identifier>
<mods:identifier type="ISSN">2305-2562</mods:identifier>
<mods:identifier type="Zenodo-Dep">7918274</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0D38CE4BC27DF71EA2FB7402FDAEFBB2" httpUri="http://treatment.plazi.org/id/0D38CE4BC27DF71EA2FB7402FDAEFBB2" lastPageId="99" lastPageNumber="480" pageId="98" pageNumber="479">
<subSubSection box="[462,786,299,326]" pageId="98" pageNumber="479" type="nomenclature">
<paragraph blockId="98.[462,786,299,326]" box="[462,786,299,326]" pageId="98" pageNumber="479">
<heading box="[462,786,299,326]" centered="true" fontSize="11" level="2" pageId="98" pageNumber="479" reason="2">
<taxonomicName ID-CoL="4GLVV" authority="Herbert, 2012" authorityName="Herbert" authorityYear="2012" box="[462,719,299,325]" class="Gastropoda" family="Chilodontaidae" genus="Pholidotrope" kingdom="Animalia" order="Seguenziida" pageId="98" pageNumber="479" phylum="Mollusca" rank="species" species="gloriosa" status="sp. n.">
<emphasis bold="true" box="[462,719,299,325]" pageId="98" pageNumber="479">Pholidotrope gloriosa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[726,786,300,326]" pageId="98" pageNumber="479" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="99" lastPageNumber="480" pageId="98" pageNumber="479" type="description">
<paragraph blockId="98.[144,1105,342,1225]" box="[561,686,342,368]" pageId="98" pageNumber="479">
<figureCitation box="[561,646,342,368]" captionStart="Fig" captionStartId="98.[144,177,1630,1652]" captionTargetBox="[147,1104,1262,1608]" captionTargetId="figure-344@98.[144,1104,1259,1608]" captionTargetPageId="98" captionText="Fig. 61. Pholidotrope gloriosa sp. n., holotype (MNHN): (A) apertural view, length 4.7 mm; (B) oblique basal view into aperture; (C) side view showing subterminal external varix (arrow)." figureDoi="http://doi.org/10.5281/zenodo.7918411" httpUri="https://zenodo.org/record/7918411/files/figure.png" pageId="98" pageNumber="479">Figs 61</figureCitation>
,
<figureCitation box="[658,686,342,368]" captionStart="Fig" captionStartId="99.[181,214,538,560]" captionTargetBox="[189,1134,207,519]" captionTargetId="figure-271@99.[189,1134,204,522]" captionTargetPageId="99" captionText="Fig. 62. Pholidotrope gloriosa sp. n., holotype: (A) granular juvenile microsculpture; (B) fine scratch-like microsculpture on last adult whorl; (C) protoconch. Bars all = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7918413" httpUri="https://zenodo.org/record/7918413/files/figure.png" pageId="98" pageNumber="479">62</figureCitation>
</paragraph>
<paragraph blockId="98.[144,1105,342,1225]" pageId="98" pageNumber="479">
Etymology: From Latin
<taxonomicName authorityName="Herbert" authorityYear="2012" box="[423,516,382,408]" class="Gastropoda" family="Chilodontaidae" genus="Pholidotrope" kingdom="Animalia" order="Seguenziida" pageId="98" pageNumber="479" phylum="Mollusca" rank="species" species="gloriosa">
<emphasis box="[423,516,382,408]" italics="true" pageId="98" pageNumber="479">gloriosa</emphasis>
</taxonomicName>
(famous, glorious); in reference to Îles Glorieuses, near which the
<typeStatus box="[315,363,415,441]" pageId="98" pageNumber="479">type</typeStatus>
material was collected.
</paragraph>
<paragraph blockId="98.[144,1105,342,1225]" box="[144,283,454,480]" pageId="98" pageNumber="479">Description:</paragraph>
<paragraph blockId="98.[144,1105,342,1225]" pageId="98" pageNumber="479">
<emphasis box="[144,199,494,520]" italics="true" pageId="98" pageNumber="479">Shell</emphasis>
: Trochiform, small, spire conical, base somewhat flattened (L/D=1.18); teleoconch of 5.5 whorls; initial whorls rounded, becoming more flat-sided with growth; suture narrowly indented; sculpture of spiral cords and lamellate axial pliculae; first whorl sculptured only by relatively widely spaced axial pliculae (13), second whorl with 16 axial pliculae and 2 developing spiral cords; third whorl with 17 axial pliculae and a third spiral cord developing at shoulder; fourth and fifth whorls with 3 strong spiral cords and 18 and 19 axial pliculae respectively; cords and pliculae subequal in strength, their intersections produced into scale-like nodules, those on shoulder cord somewhat more rounded; suture level with subperipheral cord; cord intervals wider than cords themselves, that between peripheral and subperipheral cord deep. Base with 4 spiral cords, progressively weaker toward centre; umbilicus narrowly patent in juvenile, but umbilical and parietal regions covered by a thin, smooth inductural callus spreading from aperture at maturity. Peristome markedly oblique, more or less in one tangential plane; aperture D-shaped; columella more or less straight, but with a distinct, roundly trigonal tooth near its base; outer lip internally thickened, its edge flaring outward; inner thickening with 10 ridge-like denticles, that nearest columella largest, creating a U-shaped notch between it and columella tooth; interior of aperture nacreous, labral denticles not extending internally as in-running spiral ridges; exterior of outer lip subterminally thickened by a low collabral varix.
</paragraph>
<paragraph blockId="98.[144,1105,342,1225]" lastBlockId="99.[181,1142,606,1027]" lastPageId="99" lastPageNumber="480" pageId="98" pageNumber="479">
Microsculpture (
<figureCitation box="[353,485,1102,1129]" captionStart="Fig" captionStartId="99.[181,214,538,560]" captionTargetBox="[189,1134,207,519]" captionTargetId="figure-271@99.[189,1134,204,522]" captionTargetPageId="99" captionText="Fig. 62. Pholidotrope gloriosa sp. n., holotype: (A) granular juvenile microsculpture; (B) fine scratch-like microsculpture on last adult whorl; (C) protoconch. Bars all = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7918413" httpUri="https://zenodo.org/record/7918413/files/figure.png" pageId="98" pageNumber="479">Fig. 62A, B</figureCitation>
): Early whorls with close-set, microscopic granules, vermiform spiral threads not evident; fine, prosocline, scratch-like marks on later whorls. Protoconch (
<figureCitation box="[315,420,1166,1192]" captionStart="Fig" captionStartId="99.[181,214,538,560]" captionTargetBox="[189,1134,207,519]" captionTargetId="figure-271@99.[189,1134,204,522]" captionTargetPageId="99" captionText="Fig. 62. Pholidotrope gloriosa sp. n., holotype: (A) granular juvenile microsculpture; (B) fine scratch-like microsculpture on last adult whorl; (C) protoconch. Bars all = 100 µm." figureDoi="http://doi.org/10.5281/zenodo.7918413" httpUri="https://zenodo.org/record/7918413/files/figure.png" pageId="98" pageNumber="479">Fig. 62C</figureCitation>
): White, globose and distinctly exsert, diameter
<emphasis box="[972,998,1167,1192]" italics="true" pageId="98" pageNumber="479">ca</emphasis>
280 µm; surface worn, but with evidence of a fine, flocculent microsculpture; terminal lip convex. Colour: White.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7918411" ID-Zenodo-Dep="7918411" httpUri="https://zenodo.org/record/7918411/files/figure.png" pageId="98" pageNumber="479" startId="98.[144,177,1630,1652]" targetBox="[147,1104,1262,1608]" targetPageId="98" targetType="figure">
<paragraph blockId="98.[144,1104,1630,1676]" pageId="98" pageNumber="479">
Fig. 61.
<taxonomicName authority="Herbert, 2012" authorityName="Herbert" authorityYear="2012" box="[221,422,1630,1651]" class="Gastropoda" family="Chilodontaidae" genus="Pholidotrope" kingdom="Animalia" order="Seguenziida" pageId="98" pageNumber="479" phylum="Mollusca" rank="species" species="gloriosa" status="sp. n.">
<emphasis box="[221,422,1630,1651]" italics="true" pageId="98" pageNumber="479">Pholidotrope gloriosa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[428,479,1631,1652]" pageId="98" pageNumber="479" rank="species">sp. n.</taxonomicNameLabel>
, holotype (MNHN): (A) apertural view, length 4.7 mm; (B) oblique basal view into aperture; (C) side view showing subterminal external varix (arrow).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7918413" ID-Zenodo-Dep="7918413" httpUri="https://zenodo.org/record/7918413/files/figure.png" pageId="99" pageNumber="480" startId="99.[181,214,538,560]" targetBox="[189,1134,207,519]" targetPageId="99" targetType="figure">
<paragraph blockId="99.[181,1142,538,584]" pageId="99" pageNumber="480">
Fig. 62.
<taxonomicName authority="Herbert, 2012" authorityName="Herbert" authorityYear="2012" box="[257,458,538,559]" class="Gastropoda" family="Chilodontaidae" genus="Pholidotrope" kingdom="Animalia" order="Seguenziida" pageId="99" pageNumber="480" phylum="Mollusca" rank="species" species="gloriosa" status="sp. n.">
<emphasis box="[257,458,538,559]" italics="true" pageId="99" pageNumber="480">Pholidotrope gloriosa</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[463,514,538,559]" pageId="99" pageNumber="480" rank="species">sp. n.</taxonomicNameLabel>
, holotype: (A) granular juvenile microsculpture; (B) fine scratch-like microsculpture on last adult whorl; (C) protoconch. Bars all = 100 µm.
</paragraph>
</caption>
<paragraph blockId="99.[181,1142,606,1027]" box="[208,850,638,664]" pageId="99" pageNumber="480">
Dimensions:
<typeStatus box="[357,464,638,664]" pageId="99" pageNumber="480">Holotype</typeStatus>
, length
<quantity box="[552,640,638,664]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="99" pageNumber="480" unit="mm" value="4.7">4.7 mm</quantity>
, diameter
<quantity box="[756,845,638,664]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="99" pageNumber="480" unit="mm" value="3.9">3.9 mm</quantity>
.
</paragraph>
<paragraph blockId="99.[181,1142,606,1027]" box="[181,779,678,704]" pageId="99" pageNumber="480">
<emphasis box="[181,646,678,704]" italics="true" pageId="99" pageNumber="480">Operculum, radula and external anatomy</emphasis>
: Unknown.
</paragraph>
<paragraph blockId="99.[181,1142,606,1027]" pageId="99" pageNumber="480">
<materialsCitation collectionCode="MNHN" country="Madagascar" elevation="93" latitude="-12.53" location="Banc du Leven" longLatPrecision="776" longitude="47.67" pageId="99" pageNumber="480" specimenCode="MNHN 24656" specimenCount="1" typeStatus="holotype">
<typeStatus box="[181,269,716,738]" pageId="99" pageNumber="480">Holotype</typeStatus>
:
<collectingCountry box="[278,443,716,737]" name="Madagascar" pageId="99" pageNumber="480">MADAGASCAR</collectingCountry>
:
<locationDeviation box="[451,631,716,738]" location="Madagascar" pageId="99" pageNumber="480">NW of Madagascar</locationDeviation>
, west of
<location LSID="urn:lsid:plazi:treatment:0D38CE4BC27DF71EA2FB7402FDAEFBB2:804E2986C27CF71EA1FD77E4FCA6FD52" box="[712,850,716,738]" country="Madagascar" latitude="-12.53" longLatPrecision="776" longitude="47.67" name="Banc du Leven" pageId="99" pageNumber="480">Banc du Leven</location>
(
<geoCoordinate box="[862,936,716,738]" degrees="12.53" direction="south" orientation="latitude" pageId="99" pageNumber="480" precision="555" value="-12.53">12.53°S</geoCoordinate>
<geoCoordinate box="[942,1019,716,738]" degrees="47.67" direction="east" orientation="longitude" pageId="99" pageNumber="480" precision="555" value="47.67">47.67°E</geoCoordinate>
), -
<quantity box="[1041,1138,716,738]" metricMagnitude="1" metricUnit="m" metricValue="9.25" metricValueMax="15.0" metricValueMin="3.5" pageId="99" pageNumber="480" unit="m" value="92.5" valueMax="150.0" valueMin="35.0">
<elevation box="[1041,1138,716,738]" metricMagnitude="1" metricUnit="m" metricValue="9.25" metricValueMax="15.0" metricValueMin="3.5" pageId="99" pageNumber="480" unit="m" value="92.5" valueMax="150.0" valueMin="35.0">35150 m</elevation>
</quantity>
,
<smallCapsWord baselines="757,756" box="[181,266,741,762]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Benthedi" pageId="99" pageNumber="480">BENTHEDI</smallCapsWord>
Expedn, stn 5, dredged, 1977 (
<specimenCode box="[563,700,740,762]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="99" pageNumber="480">MNHN 24656</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="99.[181,1142,606,1027]" pageId="99" pageNumber="480">
<materialsCitation collectingDate="1977-03-19" collectionCode="MNHN" country="Madagascar" elevation="250" latitude="-11.487" location="Iles Glorieuses" longLatPrecision="77" longitude="47.303" municipality="Exped'n" pageId="99" pageNumber="480" specimenCode="MNHN 24657" specimenCount="1" typeStatus="paratype">
<typeStatus box="[181,264,773,794]" pageId="99" pageNumber="480">Paratype</typeStatus>
:
<collectingCountry box="[271,436,772,793]" name="Madagascar" pageId="99" pageNumber="480">MADAGASCAR</collectingCountry>
:
<locationDeviation box="[442,618,772,794]" location="Madagascar" pageId="99" pageNumber="480">NW of Madagascar</locationDeviation>
, west of
<location LSID="urn:lsid:plazi:treatment:0D38CE4BC27DF71EA2FB7402FDAEFBB2:804E2986C27CF71EA183762CFCCFFCAA" box="[694,827,772,795]" country="Madagascar" latitude="-11.487" longLatPrecision="77" longitude="47.303" municipality="Exped'n" name="Iles Glorieuses" pageId="99" pageNumber="480">Îles Glorieuses</location>
(
<geoCoordinate box="[838,921,772,793]" degrees="11.487" direction="south" orientation="latitude" pageId="99" pageNumber="480" precision="55" value="-11.487">11.487°S</geoCoordinate>
<geoCoordinate box="[926,1014,772,793]" degrees="47.303" direction="east" orientation="longitude" pageId="99" pageNumber="480" precision="55" value="47.303">47.303°E</geoCoordinate>
), -
<quantity box="[1034,1094,772,794]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="99" pageNumber="480" unit="m" value="250.0">
<elevation box="[1034,1094,772,794]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="99" pageNumber="480" unit="m" value="250.0">250 m</elevation>
</quantity>
, rock dredge,
<smallCapsWord baselines="813,812" box="[254,339,797,818]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Benthedi" pageId="99" pageNumber="480">BENTHEDI</smallCapsWord>
<collectingMunicipality box="[344,423,796,818]" pageId="99" pageNumber="480">Expedn</collectingMunicipality>
, stn 8, dredged,
<date box="[579,677,796,818]" pageId="99" pageNumber="480" value="1977-03-19">
<collectingDate box="[579,677,796,818]" pageId="99" pageNumber="480" value="1977-03-19">19.iii.1977</collectingDate>
</date>
(
<specimenCode box="[689,826,796,818]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="99" pageNumber="480">MNHN 24657</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="99.[181,1142,606,1027]" pageId="99" pageNumber="480">
Distribution and habitat: Known only from off north-western
<collectingCountry box="[864,1001,832,858]" name="Madagascar" pageId="99" pageNumber="480">Madagascar</collectingCountry>
, -
<quantity box="[1019,1137,832,859]" metricMagnitude="2" metricUnit="m" metricValue="1.425" metricValueMax="2.5" metricValueMin="0.35" pageId="99" pageNumber="480" unit="m" value="142.5" valueMax="250.0" valueMin="35.0">35250 m</quantity>
, presumably on rocky substrata.
</paragraph>
<paragraph blockId="99.[181,1142,606,1027]" pageId="99" pageNumber="480">
Remarks: The small size and conical profile of
<taxonomicName authorityName="Herbert" authorityYear="2012" box="[726,974,904,930]" class="Gastropoda" family="Chilodontaidae" genus="Pholidotrope" kingdom="Animalia" order="Seguenziida" pageId="99" pageNumber="480" phylum="Mollusca" rank="species" species="gloriosa">
<emphasis box="[726,974,904,930]" italics="true" pageId="99" pageNumber="480">Pholidotrope gloriosa</emphasis>
</taxonomicName>
renders it distinctive amongst south-western Indian Ocean chilodontids. The only similar species,
<taxonomicName authorityName="Herbert" authorityYear="2012" baseAuthorityName="Barnard" baseAuthorityYear="1964" box="[181,348,968,994]" class="Gastropoda" family="Chilodontaidae" genus="Perrinia" kingdom="Animalia" order="Seguenziida" pageId="99" pageNumber="480" phylum="Mollusca" rank="species" species="konos">
<emphasis box="[181,348,968,994]" italics="true" pageId="99" pageNumber="480">Perrinia konos</emphasis>
</taxonomicName>
, is more elevated, less coarsely sculptured and has less well-developed dentition at the base of the columella.
</paragraph>
</subSubSection>
</treatment>
</document>