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<document ID-DOI="10.5733/afin.053.0209" ID-ISSN="2305-2562" ID-Zenodo-Dep="7918274" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="valdenar" IM.treatmentCitations_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1683581517355" checkinUser="felipe" docAuthor="Herbert, D. G." docDate="2012" docId="0D38CE4BC200F75FA12770F8FF2BFED6" docLanguage="en" docName="AfrInvertebr.53.2.381-502.pdf" docOrigin="African Invertebrates 53 (2)" docSource="http://www.bioone.org/doi/10.5733/afin.053.0209" docStyle="DocumentStyle:B78C63EC8CEBBB75364A82B35BB1F83D.4:AfrInvertebr.2008-2015.journal_article.0cover" docStyleId="B78C63EC8CEBBB75364A82B35BB1F83D" docStyleName="AfrInvertebr.2008-2015.journal_article.0cover" docStyleVersion="4" docTitle="Danilia boucheti Herbert 2012, sp. n." docType="treatment" docVersion="3" lastPageNumber="414" masterDocId="F101B633C21FF77DA3357529FFF4FFB0" masterDocTitle="A revision of the Chilodontidae (Gastropoda: Vetigastropoda: Seguenzioidea) of southern Africa and the south-western Indian Ocean" masterLastPageNumber="381" masterPageNumber="381" pageNumber="412" updateTime="1683700561700" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title>A revision of the Chilodontidae (Gastropoda: Vetigastropoda: Seguenzioidea) of southern Africa and the south-western Indian Ocean</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Herbert, D. G.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>African Invertebrates</mods:title>
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<mods:date>2012</mods:date>
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<mods:number>2012-11-06</mods:number>
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<mods:detail type="volume">
<mods:number>53</mods:number>
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<mods:number>2</mods:number>
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<mods:start>381</mods:start>
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<mods:url>http://www.bioone.org/doi/10.5733/afin.053.0209</mods:url>
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<mods:identifier type="ISSN">2305-2562</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:0D38CE4BC200F75FA12770F8FF2BFED6" httpUri="http://treatment.plazi.org/id/0D38CE4BC200F75FA12770F8FF2BFED6" lastPageId="34" lastPageNumber="414" pageId="31" pageNumber="412">
<subSubSection box="[530,793,1489,1516]" pageId="31" pageNumber="412" type="nomenclature">
<paragraph blockId="31.[530,793,1489,1516]" box="[530,793,1489,1516]" pageId="31" pageNumber="412">
<heading box="[530,793,1489,1516]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="412" reason="2">
<taxonomicName ID-CoL="343YN" authority="Herbert, 2012" authorityName="Herbert" authorityYear="2012" box="[530,727,1489,1515]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="31" pageNumber="412" phylum="Mollusca" rank="species" species="boucheti" status="sp. n.">
<emphasis bold="true" box="[530,727,1489,1515]" pageId="31" pageNumber="412">Danilia boucheti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[733,793,1490,1516]" pageId="31" pageNumber="412" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="34" lastPageNumber="415" pageId="31" pageNumber="412" type="description">
<paragraph blockId="31.[599,724,1532,1558]" box="[599,724,1532,1558]" pageId="31" pageNumber="412">
<figureCitation box="[599,684,1532,1558]" captionStart="Fig" captionStartId="32.[144,177,834,856]" captionTargetBox="[151,1096,204,813]" captionTargetId="figure-312@32.[151,1096,204,813]" captionTargetPageId="32" captionText="Fig. 13. Distribution of Danilia species in the south-western Indian Ocean: D. boucheti (square), D. textilis (triangles). Each symbol represents one or more records." figureDoi="http://doi.org/10.5281/zenodo.7918304" httpUri="https://zenodo.org/record/7918304/files/figure.png" pageId="31" pageNumber="412">Figs 13</figureCitation>
,
<figureCitation box="[696,724,1532,1558]" captionStart="Fig" captionStartId="33.[181,214,503,525]" captionTargetBox="[181,1141,204,479]" captionTargetId="figure-393@33.[181,1141,204,479]" captionTargetPageId="33" captionText="Fig. 14. Danilia boucheti sp. n., south-west of Cap dAmbre, Antsiranana, Madagascar: (AC) holotype (apertural features subadult), length 5.3 mm (MNHN 24648); (D) paratype (apertural features adult, apex missing), diameter 4.46 mm (MNHN 24649)." figureDoi="http://doi.org/10.5281/zenodo.7918306" httpUri="https://zenodo.org/record/7918306/files/figure.png" pageId="31" pageNumber="412">14</figureCitation>
</paragraph>
<paragraph blockId="31.[181,1141,1574,1633]" pageId="31" pageNumber="412">Etymology: Named for Prof. Philippe Bouchet (MNHN), well-known malacologist and co-ordinator of numerous important dredging expeditions.</paragraph>
<paragraph blockId="31.[181,321,1649,1675]" lastBlockId="33.[181,1142,603,1460]" lastPageId="33" lastPageNumber="414" pageId="31" pageNumber="412">
Description:
<emphasis box="[144,200,914,940]" italics="true" pageId="32" pageNumber="413">Shell</emphasis>
: Turbiniform with moderately elevated spire (L/D=1.091.14); teleoconch of 4.254.5 whorls; apical angle approx. 75°; whorls convex and suture indented, level with periphery on spire, but descending below this just prior to outer lip; protoconch sunken and apex thus appearing truncated; outer lip with well-developed subterminal varix. First teleoconch whorl sculptured with approx. 17 orthocline axial pliculae; 4 spiral cords develop during second whorl (P1P4); additional secondary and tertiary cords arising during subsequent whorls; abapical cord (P4) strongest; axial pliculae narrow, becoming strongly prosocline on later whorls, the crest crisp and leaning forward; spiral cords beaded where crossed by axial pliculae, beads on primary cords scale-like; last adult whorl with P2P4 (particularly P3 and P4) stronger than other cords; P3 and P4 delimiting an almost cylindrical peripheral band; subsutural cord (P1) also slightly larger and with angular beads that intermesh in a zip-like manner with beads of peripheral cord (P4) of preceding whorl, delimiting a narrow sutural channel; number of intermediary cords variable, generally only 12 between P3 and P4; 56 between P1 and P3 (including P2); beading of intermediary cords weaker; cords generally somewhat narrower than their intervals and slightly stronger than axial pliculae; interaction of cords and pliculae producing a fine, regular, oblique cancellation with rhomboidal interstices. Base with approx. 10 spiral cords of alternating size; the second one below peripheral cord generally the strongest; axial pliculae continue onto base rendering basal cords finely beaded. Peristome markedly oblique, more or less in one near-tangential plane; aperture roundly D-shaped and flattened in parietal region; columella lip a thickened pillar set with 2 teeth, lower one a well-developed, rounded peg, the upper one smaller; interval between teeth shallowly concave; umbilical region medial to thickened columella pillar sunken, forming an elongate curved pit, broadening basally; umbilical/parietal region covered by a thin, extensive inductural callus; callus translucent, smooth and glossy, raised somewhat basally and confluent with flaring margin of outer lip; interior of outer lip with an in-set varix lying beneath external outer lip varix; internal varix swollen to form prominent bulge near junction of basal and columella lips, creating deep, U-shaped notch below lower columella tooth; apertural varix set with numerous small, ridge-like denticles, these restricted to varix, not running into deeper portions of aperture; interior of aperture with traces of nacre (specimens not fresh), not spirally lirate, though spiral cords of shell exterior visible by translucence; margin of outer lip flaring; external varix
<emphasis box="[181,207,859,884]" italics="true" pageId="33" pageNumber="414">ca</emphasis>
<quantity box="[215,315,858,885]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="33" pageNumber="414" unit="mm" value="0.25">0.25 mm</quantity>
wide and set back a similar distance behind lip edge.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7918304" ID-Zenodo-Dep="7918304" httpUri="https://zenodo.org/record/7918304/files/figure.png" pageId="32" pageNumber="413" startId="32.[144,177,834,856]" targetBox="[151,1096,204,813]" targetPageId="32" targetType="figure">
<paragraph blockId="32.[144,1103,834,880]" pageId="32" pageNumber="413">
Fig. 13. Distribution of
<taxonomicName box="[356,496,834,856]" pageId="32" pageNumber="413">
<emphasis box="[356,425,834,855]" italics="true" pageId="32" pageNumber="413">Danilia</emphasis>
species
</taxonomicName>
in the south-western Indian Ocean:
<taxonomicName authorityName="Herbert" authorityYear="2012" box="[821,926,834,855]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="32" pageNumber="413" phylum="Mollusca" rank="species" species="boucheti">
<emphasis box="[821,926,834,855]" italics="true" pageId="32" pageNumber="413">D. boucheti</emphasis>
</taxonomicName>
(square),
<taxonomicName authorityName="Herbert" authorityYear="2012" box="[1016,1103,834,855]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="32" pageNumber="413" phylum="Mollusca" rank="species" species="textilis">
<emphasis box="[1016,1103,834,855]" italics="true" pageId="32" pageNumber="413">D. textilis</emphasis>
</taxonomicName>
(triangles). Each symbol represents one or more records.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7918306" ID-Zenodo-Dep="7918306" httpUri="https://zenodo.org/record/7918306/files/figure.png" pageId="33" pageNumber="414" startId="33.[181,214,503,525]" targetBox="[181,1141,204,479]" targetPageId="33" targetType="figure">
<paragraph blockId="33.[181,1142,503,573]" pageId="33" pageNumber="414">
Fig. 14.
<taxonomicName authority="Herbert, 2012" authorityName="Herbert" authorityYear="2012" box="[260,414,503,524]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="33" pageNumber="414" phylum="Mollusca" rank="species" species="boucheti" status="sp. n.">
<emphasis box="[260,414,503,524]" italics="true" pageId="33" pageNumber="414">Danilia boucheti</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[421,473,503,524]" pageId="33" pageNumber="414" rank="species">sp. n.</taxonomicNameLabel>
, south-west of Cap dAmbre, Antsiranana, Madagascar: (AC) holotype (apertural features subadult), length 5.3 mm (MNHN 24648); (D) paratype (apertural features adult, apex missing), diameter 4.46 mm (MNHN 24649).
</paragraph>
</caption>
<paragraph blockId="33.[181,1142,603,1460]" pageId="33" pageNumber="414">Microsculpture: Irregular, vermiform spiral threads present on early whorls; later whorls with evidence of scratch-like sculpture, but sculptural details largely obscured by dirty intritacalx deposit, particularly in interstices.</paragraph>
<paragraph blockId="33.[181,1142,603,1460]" pageId="33" pageNumber="414">
Protoconch: Present only in
<typeStatus box="[530,632,987,1013]" pageId="33" pageNumber="414">holotype</typeStatus>
; translucent white; diameter 320 µm; sunken into first teleoconch whorl and somewhat tilted; terminal lip evidently more or less straight.
</paragraph>
<paragraph blockId="33.[181,1142,603,1460]" pageId="33" pageNumber="414">
Colour: First teleoconch whorl uniform white, later whorls pale buff with faint brown spots on primary spiral cords; flared outer lip and external varix with brown marks in intervals between paler primary spiral cords. One
<typeStatus box="[721,816,1147,1173]" pageId="33" pageNumber="414">paratype</typeStatus>
with darker brown subsutural blotches, stronger spots on the primary peripheral cords and irregular brownish mottling on the base.
</paragraph>
<paragraph blockId="33.[181,1142,603,1460]" box="[208,1090,1243,1270]" pageId="33" pageNumber="414">
Dimensions:
<typeStatus box="[357,461,1243,1269]" pageId="33" pageNumber="414">Holotype</typeStatus>
(largest specimen), length
<quantity box="[764,866,1243,1270]" metricMagnitude="-3" metricUnit="m" metricValue="5.32" pageId="33" pageNumber="414" unit="mm" value="5.32">5.32 mm</quantity>
, diameter
<quantity box="[983,1086,1243,1269]" metricMagnitude="-3" metricUnit="m" metricValue="4.66" pageId="33" pageNumber="414" unit="mm" value="4.66">4.66 mm</quantity>
.
</paragraph>
<paragraph blockId="33.[181,1142,603,1460]" box="[181,779,1285,1312]" pageId="33" pageNumber="414">
<emphasis box="[181,646,1285,1311]" italics="true" pageId="33" pageNumber="414">Operculum, radula and external anatomy</emphasis>
: Unknown.
</paragraph>
<paragraph blockId="33.[181,1142,603,1460]" pageId="33" pageNumber="414">
<materialsCitation collectingDate="2009-06-28" collectionCode="MNHN" country="Madagascar" elevation="244" latitude="-12.133" location="Campagne Miriky" longLatPrecision="77" longitude="48.933" municipality="Cap d'Ambre" pageId="33" pageNumber="414" specimenCode="MNHN 24648" specimenCount="1" stateProvince="Antsiranana" typeStatus="holotype">
<typeStatus box="[181,265,1324,1346]" pageId="33" pageNumber="414">Holotype</typeStatus>
(
<figureCitation box="[277,388,1324,1346]" captionStart="Fig" captionStartId="33.[181,214,503,525]" captionTargetBox="[181,1141,204,479]" captionTargetId="figure-393@33.[181,1141,204,479]" captionTargetPageId="33" captionText="Fig. 14. Danilia boucheti sp. n., south-west of Cap dAmbre, Antsiranana, Madagascar: (AC) holotype (apertural features subadult), length 5.3 mm (MNHN 24648); (D) paratype (apertural features adult, apex missing), diameter 4.46 mm (MNHN 24649)." figureDoi="http://doi.org/10.5281/zenodo.7918306" httpUri="https://zenodo.org/record/7918306/files/figure.png" pageId="33" pageNumber="414">Fig. 14AC</figureCitation>
):
<collectingCountry box="[402,567,1324,1345]" name="Madagascar" pageId="33" pageNumber="414">MADAGASCAR</collectingCountry>
:
<collectingRegion box="[573,685,1324,1346]" country="Madagascar" name="Antsiranana" pageId="33" pageNumber="414">Antsiranana</collectingRegion>
, south-west of
<collectingMunicipality box="[819,942,1324,1346]" pageId="33" pageNumber="414">Cap dAmbre</collectingMunicipality>
(
<geoCoordinate box="[953,1039,1324,1345]" degrees="12.133" direction="south" orientation="latitude" pageId="33" pageNumber="414" precision="55" value="-12.133">12.133°S</geoCoordinate>
<geoCoordinate box="[1043,1132,1324,1345]" degrees="48.933" direction="east" orientation="longitude" pageId="33" pageNumber="414" precision="55" value="48.933">48.933°E</geoCoordinate>
), -
<quantity box="[188,296,1348,1369]" metricMagnitude="2" metricUnit="m" metricValue="2.435" metricValueMax="2.4899999999999998" metricValueMin="2.38" pageId="33" pageNumber="414" unit="m" value="243.5" valueMax="249.0" valueMin="238.0">
<elevation box="[188,296,1348,1369]" metricMagnitude="2" metricUnit="m" metricValue="2.435" metricValueMax="2.4899999999999998" metricValueMin="2.38" pageId="33" pageNumber="414" unit="m" value="243.5" valueMax="249.0" valueMin="238.0">238249 m</elevation>
</quantity>
,
<location LSID="urn:lsid:plazi:treatment:0D38CE4BC200F75FA12770F8FF2BFED6:804E2986C23EF75CA204706DFE27FAE9" box="[305,467,1348,1369]" country="Madagascar" latitude="-12.133" longLatPrecision="77" longitude="48.933" municipality="Cap d'Ambre" name="Campagne Miriky" pageId="33" pageNumber="414" stateProvince="Antsiranana">
Campagne
<emphasis box="[407,467,1348,1369]" italics="true" pageId="33" pageNumber="414">Miriky</emphasis>
</location>
, stn DW3196, dredged,
<date box="[697,794,1348,1370]" pageId="33" pageNumber="414" value="2009-06-28">
<collectingDate box="[697,794,1348,1370]" pageId="33" pageNumber="414" value="2009-06-28">28.vi.2009</collectingDate>
</date>
(
<specimenCode box="[806,944,1348,1370]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="414">MNHN 24648</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="33.[181,1142,603,1460]" box="[181,767,1380,1402]" pageId="33" pageNumber="414">
<materialsCitation box="[181,763,1380,1402]" collectingDate="2009-06-28" collectionCode="MNHN" country="Madagascar" elevation="244" latitude="-12.133" location="Campagne Miriky" longLatPrecision="77" longitude="48.933" municipality="Cap d'Ambre" pageId="33" pageNumber="414" specimenCode="MNHN 24649" specimenCount="2" stateProvince="Antsiranana" typeStatus="paratype">
<typeStatus box="[181,273,1380,1401]" pageId="33" pageNumber="414">Paratypes</typeStatus>
: Same data as holotype (
<specimenCode box="[497,632,1380,1402]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="33" pageNumber="414">MNHN 24649</specimenCode>
,
<specimenCount box="[641,757,1380,1402]" pageId="33" pageNumber="414" type="generic">2 specimens</specimenCount>
)
</materialsCitation>
.
</paragraph>
<paragraph blockId="33.[181,1142,603,1460]" pageId="33" pageNumber="414">
Note: The
<typeStatus box="[274,352,1414,1436]" pageId="33" pageNumber="414">holotype</typeStatus>
is the most complete specimen, but its apertural dentition is subadult; one of the
<typeStatus box="[1055,1141,1415,1436]" pageId="33" pageNumber="414">paratypes</typeStatus>
has more mature apertural features (
<figureCitation box="[509,592,1438,1460]" captionStart="Fig" captionStartId="33.[181,214,503,525]" captionTargetBox="[181,1141,204,479]" captionTargetId="figure-393@33.[181,1141,204,479]" captionTargetPageId="33" captionText="Fig. 14. Danilia boucheti sp. n., south-west of Cap dAmbre, Antsiranana, Madagascar: (AC) holotype (apertural features subadult), length 5.3 mm (MNHN 24648); (D) paratype (apertural features adult, apex missing), diameter 4.46 mm (MNHN 24649)." figureDoi="http://doi.org/10.5281/zenodo.7918306" httpUri="https://zenodo.org/record/7918306/files/figure.png" pageId="33" pageNumber="414">Fig. 14D</figureCitation>
), but is missing the apical whorls.
</paragraph>
<paragraph blockId="33.[181,1141,1477,1536]" pageId="33" pageNumber="414">
Distribution (
<figureCitation box="[332,416,1477,1503]" captionStart="Fig" captionStartId="32.[144,177,834,856]" captionTargetBox="[151,1096,204,813]" captionTargetId="figure-312@32.[151,1096,204,813]" captionTargetPageId="32" captionText="Fig. 13. Distribution of Danilia species in the south-western Indian Ocean: D. boucheti (square), D. textilis (triangles). Each symbol represents one or more records." figureDoi="http://doi.org/10.5281/zenodo.7918304" httpUri="https://zenodo.org/record/7918304/files/figure.png" pageId="33" pageNumber="414">Fig. 13</figureCitation>
): Known only from off the coast of the far north of
<collectingCountry box="[1000,1137,1477,1503]" name="Madagascar" pageId="33" pageNumber="414">Madagascar</collectingCountry>
, -
<quantity box="[189,323,1509,1536]" metricMagnitude="2" metricUnit="m" metricValue="2.435" metricValueMax="2.4899999999999998" metricValueMin="2.38" pageId="33" pageNumber="414" unit="m" value="243.5" valueMax="249.0" valueMin="238.0">
238
<specimenCount box="[247,323,1509,1536]" pageId="33" pageNumber="414" type="generic">249 m</specimenCount>
</quantity>
.
</paragraph>
<paragraph blockId="33.[181,1142,1552,1675]" lastBlockId="34.[144,1104,204,358]" lastPageId="34" lastPageNumber="415" pageId="33" pageNumber="414">
Remarks:
<taxonomicName authorityName="Herbert" authorityYear="2012" box="[293,480,1552,1578]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="33" pageNumber="414" phylum="Mollusca" rank="species" species="boucheti">
<emphasis box="[293,480,1552,1578]" italics="true" pageId="33" pageNumber="414">Danilia boucheti</emphasis>
</taxonomicName>
is smaller than any of the other described species of
<taxonomicName authorityName=": Brusina" authorityYear="1865" box="[1049,1134,1552,1578]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="33" pageNumber="414" phylum="Mollusca" rank="genus">
<emphasis box="[1049,1134,1552,1578]" italics="true" pageId="33" pageNumber="414">Danilia</emphasis>
</taxonomicName>
, most of which attain or exceed
<quantity box="[534,614,1584,1611]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="414" unit="mm" value="10.0">10 mm</quantity>
in length (Vilvens &amp; Héros 2005). Perhaps the most similar species is
<taxonomicName authority="Vilvens &amp; Heros, 2005" authorityName="Vilvens &amp; Heros" authorityYear="2005" box="[449,861,1616,1643]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="33" pageNumber="414" phylum="Mollusca" rank="species" species="angulosa">
<emphasis box="[449,589,1616,1642]" italics="true" pageId="33" pageNumber="414">D. angulosa</emphasis>
Vilvens &amp; Héros, 2005
</taxonomicName>
from Melanesia and the
<collectingCountry box="[181,310,1648,1674]" name="Philippines" pageId="33" pageNumber="414">Philippines</collectingCountry>
, which is also small (length up to
<quantity box="[708,797,1648,1674]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="33" pageNumber="414" unit="mm" value="7.9">7.9 mm</quantity>
) and has stronger spiral cords associated with the periphery, creating a cylindrical peripheral band. In that species, however, the cords are stronger than in
<taxonomicName authorityName="Herbert" authorityYear="2012" box="[569,697,236,262]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="34" pageNumber="415" phylum="Mollusca" rank="species" species="boucheti">
<emphasis box="[569,697,236,262]" italics="true" pageId="34" pageNumber="415">D. boucheti</emphasis>
</taxonomicName>
, the whorls more distinctly shoulder- ed, and there is only one intermediary cord between the shoulder cord (upper peripheral cord) and the subsutural cord. In
<taxonomicName authorityName="Herbert" authorityYear="2012" box="[510,639,300,326]" class="Gastropoda" family="Chilodontaidae" genus="Danilia" kingdom="Animalia" order="Seguenziida" pageId="34" pageNumber="415" phylum="Mollusca" rank="species" species="boucheti">
<emphasis box="[510,639,300,326]" italics="true" pageId="34" pageNumber="415">D. boucheti</emphasis>
</taxonomicName>
there are 5 or 6 intermediary cords in this region.
</paragraph>
</subSubSection>
</treatment>
</document>