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<document id="888492938D5E63B1703AA4E9CC27D764" ID-DOI="10.5281/zenodo.213360" ID-GBIF-Dataset="b1a33963-aa99-478f-8b76-50c13ada801c" ID-ISSN="1175-5326" ID-Zenodo-Dep="213360" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460549395632" checkinUser="plazi" docAuthor="Sathiandran, Nithya &amp; Sabu, Thomas K." docDate="2012" docId="0D07755309340F5DFF3706C3F9C4ABF6" docLanguage="en" docName="zt03526p058.pdf" docOrigin="Zootaxa 3526" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus sahai Biswas &amp; Chatterjee 1986" docType="treatment" docVersion="9" lastPageNumber="57" masterDocId="F13E0D2B09370F59FFA0036DFD7CAA5A" masterDocTitle="New species, new synonym, and redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India" masterLastPageNumber="58" masterPageNumber="53" pageNumber="56" updateTime="1698318061882" updateUser="plazi">
<mods:mods id="AAD9629C397FE6F2F88149ADC7600AB3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BCE81625612B510D9CEEF6A0382BE52F">
<mods:title id="7FDEEE3C849B589181BED0FF44CA3A4D">New species, new synonym, and redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India</mods:title>
</mods:titleInfo>
<mods:name id="E97ED96B4C48D1A9F3F5DB9F1FFC3F14" type="personal">
<mods:role id="EBED8BCFDC5729F11DFA67B9E072B12D">
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</mods:role>
<mods:namePart id="B23C96E1ECD710C868FEC710A762007C">Sathiandran, Nithya</mods:namePart>
</mods:name>
<mods:name id="CBAE5094B3D5D02B05F9BB04247909FD" type="personal">
<mods:role id="E1818954B430D2614B36A33F3C8861D4">
<mods:roleTerm id="46E586C6A97B00A6AD7B2D0F5BBDAC33">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5E8BA30D7CDF73B61F429EE046F13044">Sabu, Thomas K.</mods:namePart>
</mods:name>
<mods:typeOfResource id="420246D1F6FF0A0FBECF22AACDAC3326">text</mods:typeOfResource>
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<mods:title id="0625508C7669B2F9C7A1458B8DF8CFE3">Zootaxa</mods:title>
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<mods:part id="F0884AA87ADE93C4736C895388F0E1C7">
<mods:date id="41F13C93178162E919D0252F39FB676A">2012</mods:date>
<mods:detail id="E49CD783EA6188E3ED5725C0CC0FF4B8" type="volume">
<mods:number id="EDFE9086D4F2A916F04E66CA8AE384B3">3526</mods:number>
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<mods:start id="30C987310E2B0BF3CC96BA60659218FC">53</mods:start>
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<mods:classification id="EEFB99CF46A61117CE99925B4D9362EA">journal article</mods:classification>
<mods:identifier id="3C92BE980BAF36902041A7A7855AA05D" type="DOI">10.5281/zenodo.213360</mods:identifier>
<mods:identifier id="1CCF1489A20E2B93C1E6758243B74D88" type="GBIF-Dataset">b1a33963-aa99-478f-8b76-50c13ada801c</mods:identifier>
<mods:identifier id="A107FF1A135BD69104FD12A8C316AC2F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7B510C3CE24B3732FFE2027CD1EEC662" type="Zenodo-Dep">213360</mods:identifier>
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<treatment id="0D07755309340F5DFF3706C3F9C4ABF6" ID-DOI="http://doi.org/10.5281/zenodo.6175216" ID-GBIF-Taxon="119529155" ID-Zenodo-Dep="6175216" LSID="urn:lsid:plazi:treatment:0D07755309340F5DFF3706C3F9C4ABF6" httpUri="http://treatment.plazi.org/id/0D07755309340F5DFF3706C3F9C4ABF6" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
<subSubSection id="CDB497CE09340F5AFF3706C3FC6EAFB3" pageId="3" pageNumber="56" type="nomenclature">
<paragraph id="8511C44509340F5AFF3706C3FFAAAF92" blockId="3.[151,726,1454,1513]" box="[151,726,1454,1480]" pageId="3" pageNumber="56">
<heading id="DE59732909340F5AFF3706C3FFAAAF92" bold="true" box="[151,726,1454,1480]" fontSize="11" level="1" pageId="3" pageNumber="56" reason="1">
<taxonomicName id="42AEBFC609340F5AFF3706C3FFAAAF92" ID-CoL="7XP4M" authority="Biswas &amp; Chatterjee, 1986" authorityName="Biswas &amp; Chatterjee" authorityYear="1986" box="[151,726,1454,1480]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="56" phylum="Arthropoda" rank="species" species="sahai">
<emphasis id="B7DA185709340F5AFF3706C3FFAAAF92" bold="true" box="[151,726,1454,1480]" pageId="3" pageNumber="56">
<emphasis id="B7DA185709340F5AFF3706C3FCF8AF92" bold="true" box="[151,388,1454,1480]" italics="true" pageId="3" pageNumber="56">Onthophagus sahai</emphasis>
<bibRefCitation id="E13FB9B409340F5AFE2B06C3FFAAAF92" author="Biswas" box="[395,726,1454,1480]" pageId="3" pageNumber="56" refString="Biswas, S. &amp; Chatterjee, S. K. (1986) Scarabaeidae (Coleoptera) of Silent Valley, Kerala, India, with description of three new species. Silent Valley special issue. Record of Zoological Survey of India, 84, 79 - 96." type="journal article" year="1986">Biswas &amp; Chatterjee, 1986</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8511C44509340F5AFF3706BCFC6EAFB3" blockId="3.[151,726,1454,1513]" box="[151,274,1489,1513]" pageId="3" pageNumber="56">
(
<figureCitation id="1D95D8C009340F5AFF3F06BCFC75AFB3" box="[159,265,1489,1513]" captionStart="FIGURES 4 – 6" captionStartId="4.[151,264,1349,1372]" captionTargetBox="[449,1136,473,1329]" captionTargetId="figure@4.[442,1145,461,1338]" captionTargetPageId="4" captionText="FIGURES 4 – 6. Onthophagus sahai: dorsal habitus—male (4), female (5), aedeagus, lateral view (6). Scale bar = 1 mm." httpUri="https://zenodo.org/record/213362/files/figure.png" pageId="3" pageNumber="56">Figs. 4–6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CDB497CE09340F5DFF370574FCD3AAAD" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="description">
<paragraph id="8511C44509340F5AFF370574FF8FAD2F" blockId="3.[151,1436,1561,2017]" pageId="3" pageNumber="56">
<emphasis id="B7DA185709340F5AFF370574FC55AC68" bold="true" box="[151,297,1561,1586]" pageId="3" pageNumber="56">Description.</emphasis>
Shiny black, with the tarsi, antennae, and the lower surface red, body smooth and without setae (
<figureCitation id="1D95D8C009340F5AFACE0574FDB9AC0F" captionStart="FIGURES 4 – 6" captionStartId="4.[151,264,1349,1372]" captionTargetBox="[449,1136,473,1329]" captionTargetId="figure@4.[442,1145,461,1338]" captionTargetPageId="4" captionText="FIGURES 4 – 6. Onthophagus sahai: dorsal habitus—male (4), female (5), aedeagus, lateral view (6). Scale bar = 1 mm." httpUri="https://zenodo.org/record/213362/files/figure.png" pageId="3" pageNumber="56">Fig. 4–5</figureCitation>
).
<emphasis id="B7DA185709340F5AFF780553FC63AC0C" bold="true" box="[216,287,1598,1622]" pageId="3" pageNumber="56">Head.</emphasis>
Broad; ocular lobes rounded; clypeus separated from the frons by a strong, straight carina; vertex with a small, transverse elevation, area behind elevation smooth.
<emphasis id="B7DA185709340F5AFCE0050FFEBCAC20" bold="true" box="[832,960,1634,1658]" pageId="3" pageNumber="56">Pronotum.</emphasis>
Convex with fine punctures. Front angles blunt, sides nearly straight in front and sinuate behind, base gently rounded.
<emphasis id="B7DA185709340F5AFC4C05EBF943ACC4" bold="true" box="[1004,1087,1670,1694]" pageId="3" pageNumber="56">Elytra.</emphasis>
Black and shiny; finely striate, striae lightly punctate, intervals feebly convex, finely but not closely punctate. Elytron with orange spot at base.
<emphasis id="B7DA185709340F5AFF3705A0FD96ACBC" bold="true" box="[151,234,1741,1766]" pageId="3" pageNumber="56">Wings.</emphasis>
<collectingCountry id="FDB984D509340F5AFF5005A0FC54ACBF" box="[240,296,1741,1765]" name="India" pageId="3" pageNumber="56">Hind</collectingCountry>
wings present.
<emphasis id="B7DA185709340F5AFE7605A3FFF5ACBC" bold="true" box="[470,649,1741,1766]" pageId="3" pageNumber="56">Lower surface.</emphasis>
Pygidium in male with an incised line near the outer margin (absent in female). Metasternal shield smooth and shiny with a large, rounded pit anteriorly; with a longitudinal groove in the middle and the sides of the metasternal shield with setiferous punctures.
<emphasis id="B7DA185709340F5AFC49047BF954AD74" bold="true" box="[1001,1064,1814,1838]" pageId="3" pageNumber="56">Legs.</emphasis>
All femora and tibiae with few setiferous punctures; protibia quadridentate externally.
<emphasis id="B7DA185709340F5AFCB70457FEAEAD08" bold="true" box="[791,978,1849,1874]" pageId="3" pageNumber="56">Male genetalia.</emphasis>
Parameres angled relative to the basal piece, symmetrical, pointed towards the apex (
<figureCitation id="1D95D8C009340F5AFD000430FF99AD2F" box="[672,741,1885,1909]" captionStart="FIGURES 4 – 6" captionStartId="4.[151,264,1349,1372]" captionTargetBox="[449,1136,473,1329]" captionTargetId="figure@4.[442,1145,461,1338]" captionTargetPageId="4" captionText="FIGURES 4 – 6. Onthophagus sahai: dorsal habitus—male (4), female (5), aedeagus, lateral view (6). Scale bar = 1 mm." httpUri="https://zenodo.org/record/213362/files/figure.png" pageId="3" pageNumber="56">Fig. 6</figureCitation>
).
</paragraph>
<paragraph id="8511C44509340F5DFF6704EFFCD3AAAD" blockId="3.[151,1436,1561,2017]" lastBlockId="4.[151,1436,151,429]" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56">
<emphasis id="B7DA185709340F5AFF6704EFFC0AADC0" bold="true" box="[199,374,1922,1946]" pageId="3" pageNumber="56">Measurements</emphasis>
(mm; n = 47): TL = 4.5–6.0, BW = 2.7–3.3, PL = 1.50–1.95, PW = 2.4–3.0, EL = 2.25–3.00.
<emphasis id="B7DA185709340F5AFF6704C8FCD5ADE4" bold="true" box="[199,425,1957,1982]" pageId="3" pageNumber="56">Material examined</emphasis>
:
<typeStatus id="5A157AE709340F5AFE1704C8FF63ADE7" box="[439,543,1957,1981]" pageId="3" pageNumber="56" type="holotype">Holotype</typeStatus>
(male, ZSI- Calcutta): “
<collectingCountry id="FDB984D509340F5AFC8D04C8FE17ADE7" box="[813,875,1957,1981]" name="India" pageId="3" pageNumber="56">India</collectingCountry>
, Kerala, Silent valley, No.10793/H4A”.
<typeStatus id="5A157AE709340F5AFA8F04CBF8E0ADE4" box="[1327,1436,1958,1982]" pageId="3" pageNumber="56" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="93A80FCC09340F5AFF3F04A4FC79ADBB" box="[159,261,1993,2017]" pageId="3" pageNumber="56" type="female">2 female</specimenCount>
, ZSI- Calcutta): Kerala, Silent valley, No. 10794/H4A; 10795/H4A. Additional material:
<specimenCount id="93A80FCC09340F5AFB5D04A4F829ADBB" box="[1277,1365,1993,2017]" pageId="3" pageNumber="56" type="male">2 males</specimenCount>
and
<specimenCount id="93A80FCC09340F5DFA2E04A4FD92AAF5" lastPageId="4" lastPageNumber="57" pageId="3" pageNumber="56" type="female">2 females</specimenCount>
in NPC labeled “
<collectingCountry id="FDB984D509330F5DFE1003FAFC92AAF5" box="[432,494,151,175]" name="India" pageId="4" pageNumber="57">India</collectingCountry>
, Kerala, Silent valley National Park, Walakkad,
<quantity id="425669A009330F5DFBA803FAF922AAF5" box="[1032,1118,151,175]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="57" unit="m" value="1100.0">1100 m</quantity>
,
<date id="F110E28509330F5DFBC903FAF9AAAAF5" box="[1129,1238,151,175]" pageId="4" pageNumber="57" value="2009-06-05">5.vi.2009</date>
, Vinod, K.V.”;
<specimenCount id="93A80FCC09330F5DFA2003FAFDA4AA8E" pageId="4" pageNumber="57" type="male">21 males</specimenCount>
and
<specimenCount id="93A80FCC09330F5DFEAF03D1FC00AA8E" box="[271,380,188,212]" pageId="4" pageNumber="57" type="female">8 females</specimenCount>
in SJC with same data,
<specimenCount id="93A80FCC09330F5DFD2C03D1FF9FAA8E" box="[652,739,188,212]" pageId="4" pageNumber="57" type="male">8 males</specimenCount>
and
<specimenCount id="93A80FCC09330F5DFCBA03D1FEFBAA8E" box="[794,903,188,212]" pageId="4" pageNumber="57" type="female">6 females</specimenCount>
in ZSI-Ca labeled “Kerala: Thirunelly,
<quantity id="425669A009330F5DFAE703D1F8E4AA8E" box="[1351,1432,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.395" pageId="4" pageNumber="57" unit="m" value="1395.0">1395m</quantity>
,
<date id="F110E28509330F5DFF3703B2FC6AAAAD" box="[151,278,223,247]" pageId="4" pageNumber="57" value="2009-05-18">18. v. 2009</date>
, Nirdev, P.M.
</paragraph>
</subSubSection>
<subSubSection id="CDB497CE09330F5DFF67026EF973AB46" box="[199,1039,259,284]" pageId="4" pageNumber="57" type="distribution">
<paragraph id="8511C44509330F5DFF67026EF973AB46" blockId="4.[151,1436,151,429]" box="[199,1039,259,284]" pageId="4" pageNumber="57">
<emphasis id="B7DA185709330F5DFF67026EFC1CAB46" bold="true" box="[199,352,259,284]" pageId="4" pageNumber="57">Distribution.</emphasis>
<collectingCountry id="FDB984D509330F5DFEC70269FCDDAB46" box="[359,417,260,284]" name="India" pageId="4" pageNumber="57">India</collectingCountry>
(South Western Ghats montane rain forests ecoregion).
</paragraph>
</subSubSection>
<subSubSection id="CDB497CE09330F5DFF670245F9C4ABF6" pageId="4" pageNumber="57" type="discussion">
<paragraph id="8511C44509330F5DFF670245F9C4ABF6" blockId="4.[151,1436,151,429]" pageId="4" pageNumber="57">
<emphasis id="B7DA185709330F5DFF670245FC47AB1A" bold="true" box="[199,315,296,320]" pageId="4" pageNumber="57">Remarks.</emphasis>
<taxonomicName id="42AEBFC609330F5DFEE20245FF61AB1A" box="[322,541,296,320]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="sahai">
<emphasis id="B7DA185709330F5DFEE20245FF61AB1A" box="[322,541,296,320]" italics="true" pageId="4" pageNumber="57">Onthophagus sahai</emphasis>
</taxonomicName>
differs from
<taxonomicName id="42AEBFC609330F5DFD120245FE47AB1A" box="[690,827,296,320]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="refulgens">
<emphasis id="B7DA185709330F5DFD120245FE47AB1A" box="[690,827,296,320]" italics="true" pageId="4" pageNumber="57">O. refulgens</emphasis>
</taxonomicName>
by the presence of an orange spot on the base of each elytron in contrast with the absence of orange spots in
<taxonomicName id="42AEBFC609330F5DFC960220FEB9AB3E" box="[822,965,332,356]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="refulgens">
<emphasis id="B7DA185709330F5DFC960220FEB9AB3E" box="[822,965,332,356]" italics="true" pageId="4" pageNumber="57">O. refulgens</emphasis>
</taxonomicName>
. Males and females of
<taxonomicName id="42AEBFC609330F5DFB490221F83DAB3E" box="[1257,1345,332,356]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="57" phylum="Arthropoda" rank="species" species="sahai">
<emphasis id="B7DA185709330F5DFB490221F83DAB3E" box="[1257,1345,332,356]" italics="true" pageId="4" pageNumber="57">O.sahai</emphasis>
</taxonomicName>
can be differentiated on the basis of variation in the punctures on clypeus and the additional carina on the clypeus of the female. The front part of the clypeus strongly punctate in males and closely rugose in females.
</paragraph>
</subSubSection>
</treatment>
</document>
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