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<document id="0879F8D6033251AAC9A93C6995A97C47" ID-DOI="10.11646/zootaxa.2046.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5321288" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629617332214" checkinUser="felipe" docAuthor="Zettel, H." docDate="2009" docId="082CE471FFF4871175A930BC4BDE948B" docLanguage="en" docName="zt02046p042.pdf" docOrigin="Zootaxa 2046 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2046.1.2" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Archaeoptilomera Zettel, 2009, gen. n." docType="treatment" docVersion="4" lastPageNumber="39" masterDocId="F4159C09FFFF871C753E34184F6F9306" masterDocTitle="Three new genera of Ptilomerinae (Hemiptera: Heteroptera: Gerridae) from Southeast Asia" masterLastPageNumber="42" masterPageNumber="26" pageNumber="37" updateTime="1699083916582" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="70A0F3ECA3426DECC675AE7761CED32A">Three new genera of Ptilomerinae (Hemiptera: Heteroptera: Gerridae) from Southeast Asia</mods:title>
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<mods:namePart id="2E84BEB147AC52A92F7C56594EA9F80A">Zettel, H.</mods:namePart>
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<mods:date id="BFF3636602CA324BCE8447234B30E2E9">2009</mods:date>
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<mods:number id="6CC612CDA9FDDFF1D2E484E17ECDF4B6">2009-03-20</mods:number>
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<mods:number id="A4277CD97DC64CEDFA69CED34416CF0E">2046</mods:number>
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<mods:url id="7F70EFABF9D2F3D5D17399DDCF94107B">https://biotaxa.org/Zootaxa/article/view/zootaxa.2046.1.2</mods:url>
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<mods:identifier id="CD6C2A2DD6113C50D23824DE3EF873D7" type="DOI">10.11646/zootaxa.2046.1.2</mods:identifier>
<mods:identifier id="6EAA73CC747627980A0AB3E84AFD5CC0" type="ISSN">1175-5326</mods:identifier>
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<treatment id="082CE471FFF4871175A930BC4BDE948B" ID-DOI="http://doi.org/10.5281/zenodo.5333966" ID-Zenodo-Dep="5333966" LSID="urn:lsid:plazi:treatment:082CE471FFF4871175A930BC4BDE948B" httpUri="http://treatment.plazi.org/id/082CE471FFF4871175A930BC4BDE948B" lastPageId="13" lastPageNumber="39" pageId="11" pageNumber="37">
<subSubSection id="C89F06ECFFF4871775A930BC4EA497BB" box="[151,459,1187,1214]" pageId="11" pageNumber="37" type="nomenclature">
<paragraph id="803A5567FFF4871775A930BC4EA497BB" blockId="11.[151,459,1187,1252]" box="[151,459,1187,1214]" pageId="11" pageNumber="37">
<heading id="DB72E20BFFF4871775A930BC4EA497BB" bold="true" box="[151,459,1187,1214]" fontSize="11" level="1" pageId="11" pageNumber="37" reason="1">
<emphasis id="B2F18975FFF4871775A930BC4EA497BB" bold="true" box="[151,459,1187,1214]" pageId="11" pageNumber="37">
<taxonomicName id="47852EE4FFF4871775A930BC4E1E97B8" ID-CoL="62D3S" authorityName="Zettel" authorityYear="2009" box="[151,369,1188,1214]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B2F18975FFF4871775A930BC4E1E97B8" bold="true" box="[151,369,1188,1214]" italics="true" pageId="11" pageNumber="37">Archaeoptilomera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9C2340EFFF48717744630BB4EA497BB" box="[376,459,1187,1213]" pageId="11" pageNumber="37" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C89F06ECFFF4871775A930D24E5B97E2" box="[151,308,1226,1252]" pageId="11" pageNumber="37" type="description">
<paragraph id="803A5567FFF4871775A930D24E5B97E2" blockId="11.[151,459,1187,1252]" box="[151,308,1226,1252]" pageId="11" pageNumber="37">
(
<figureCitation id="18BE49E2FFF4871775A130D24E4297E2" box="[159,301,1226,1252]" captionStart-0="FIGURES 25–26" captionStart-1="FIGURES 27–34" captionStartId-0="12.[151,269,1943,1967]" captionStartId-1="13.[151,270,841,865]" captionTargetBox-0="[169,1429,335,1929]" captionTargetBox-1="[288,1300,202,813]" captionTargetId-0="figure-129@12.[169,1429,331,1929]" captionTargetId-1="figure-428@13.[283,1304,194,817]" captionTargetPageId-0="12" captionTargetPageId-1="13" captionText-0="FIGURES 25–26. Archaeoptilomera gen. n. (25) Habitus of apterous female of A. derlethi sp. n. (holotype, body length 8.1 mm), dorsal aspect. (26) Habitus of apterous female of A. kodadai sp. n. (holotype, body length 8.5 mm), dorsal aspect. © NHMW Hemiptera Image Collection, published with permission." captionText-1="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi-0="http://doi.org/10.5281/zenodo.5321298" figureDoi-1="http://doi.org/10.5281/zenodo.5321300" httpUri-0="https://zenodo.org/record/5321298/files/figure.png" httpUri-1="https://zenodo.org/record/5321300/files/figure.png" pageId="11" pageNumber="37">Figs. 25–34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C89F06ECFFF4871775A9310B4C8D9628" box="[151,994,1299,1326]" pageId="11" pageNumber="37" type="type_taxon">
<paragraph id="803A5567FFF4871775A9310B4C8D9628" blockId="11.[151,1436,1299,2022]" box="[151,994,1299,1326]" pageId="11" pageNumber="37">
<emphasis id="B2F18975FFF4871775A9310B4E56962B" bold="true" box="[151,313,1299,1325]" pageId="11" pageNumber="37">
<typeStatus id="5F3EEBC5FFF4871775A9310B4FBB962B" box="[151,212,1299,1325]" pageId="11" pageNumber="37">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="47852EE4FFF48717747E310C4D18962B" ID-CoL="G7SW" authorityName="Zettel" authorityYear="2009" box="[320,631,1300,1325]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="37" phylum="Arthropoda" rank="species" species="derlethi" status="sp. nov.">
<emphasis id="B2F18975FFF48717747E310C4D18962B" box="[320,631,1300,1325]" italics="true" pageId="11" pageNumber="37">Archaeoptilomera derlethi</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF487177740310B4DAF962B" bold="true" box="[638,704,1299,1325]" pageId="11" pageNumber="37">
<taxonomicNameLabel id="A9C2340EFFF487177740310B4DAF962B" box="[638,704,1299,1325]" pageId="11" pageNumber="37" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(by present designation)
</paragraph>
</subSubSection>
<subSubSection id="C89F06ECFFF4871775FB31224BE4953A" pageId="11" pageNumber="37" type="diagnosis">
<paragraph id="803A5567FFF4871775FB31224BE4953A" blockId="11.[151,1436,1299,2022]" pageId="11" pageNumber="37">
<emphasis id="B2F18975FFF4871775FB31224E279652" bold="true" box="[197,328,1338,1364]" pageId="11" pageNumber="37">Diagnosis.</emphasis>
Small
<taxonomicName id="47852EE4FFF48717749F31224D549652" authorityName="Bianchi" authorityYear="1896" box="[417,571,1338,1364]" class="Insecta" family="Gerridae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="37" phylum="Arthropoda" rank="subFamily" subFamily="Ptilomerinae">Ptilomerinae</taxonomicName>
of predominately yellowish brown to pale yellow colour; thoracic and abdominal dorsum with smooth areas along midline (
<figureCitation id="18BE49E2FFF48717761431794CAC967D" box="[810,963,1377,1403]" captionStart="FIGURES 25–26" captionStartId="12.[151,269,1943,1967]" captionTargetBox="[169,1429,335,1929]" captionTargetId="figure-129@12.[169,1429,331,1929]" captionTargetPageId="12" captionText="FIGURES 25–26. Archaeoptilomera gen. n. (25) Habitus of apterous female of A. derlethi sp. n. (holotype, body length 8.1 mm), dorsal aspect. (26) Habitus of apterous female of A. kodadai sp. n. (holotype, body length 8.5 mm), dorsal aspect. © NHMW Hemiptera Image Collection, published with permission." figureDoi="http://doi.org/10.5281/zenodo.5321298" httpUri="https://zenodo.org/record/5321298/files/figure.png" pageId="11" pageNumber="37">Figs. 25, 26</figureCitation>
). Mesonotum in anterior two-thirds of length with pair of narrow black stripes (at medial margins of dull areas). Metanotum without median furrow. Protarsus short, about half length of protibia (
<figureCitation id="18BE49E2FFF48717778531B64C7A96CE" box="[699,789,1454,1480]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="11" pageNumber="37">Fig. 28</figureCitation>
), pale. Mesofemur and metafemur subequal in length, both with pale distal portions. Metacoxa without apical spine. Metatarsomeres fused. Abdomen of female moderately shortened, at dorsal midline slightly longer than thorax. Sternite 7 of female at connexival corner with spine (
<figureCitation id="18BE49E2FFF48717741E323A4EC0953A" box="[288,431,1570,1596]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="11" pageNumber="37">Figs. 29–32</figureCitation>
), ventrally with broad, short, round medial lobe (
<figureCitation id="18BE49E2FFF4871776D0323A4B12953A" box="[1006,1149,1570,1596]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="11" pageNumber="37">Figs. 33, 34</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C89F06ECFFF4871175FB32504D50941C" lastPageId="13" lastPageNumber="39" pageId="11" pageNumber="37" type="description">
<paragraph id="803A5567FFF4871075FB32504F8E924A" blockId="11.[151,1436,1299,2022]" lastBlockId="12.[151,1436,152,332]" lastPageId="12" lastPageNumber="38" pageId="11" pageNumber="37">
<emphasis id="B2F18975FFF4871775FB32504E0E9564" bold="true" box="[197,353,1608,1634]" pageId="11" pageNumber="37">Description.</emphasis>
Only apterous female known. Body length
<quantity id="477DF882FFF48717764832514B649565" box="[886,1035,1609,1635]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="8.5" metricValueMin="8.1" pageId="11" pageNumber="37" unit="mm" value="8.3" valueMax="8.5" valueMin="8.1">8.1–8.5 mm</quantity>
. Dorsum yellowish brown; black stripes distributed as in
<figureCitation id="18BE49E2FFF48717749032684D34958C" box="[430,603,1648,1674]" captionStart="FIGURES 25–26" captionStartId="12.[151,269,1943,1967]" captionTargetBox="[169,1429,335,1929]" captionTargetId="figure-129@12.[169,1429,331,1929]" captionTargetPageId="12" captionText="FIGURES 25–26. Archaeoptilomera gen. n. (25) Habitus of apterous female of A. derlethi sp. n. (holotype, body length 8.1 mm), dorsal aspect. (26) Habitus of apterous female of A. kodadai sp. n. (holotype, body length 8.5 mm), dorsal aspect. © NHMW Hemiptera Image Collection, published with permission." figureDoi="http://doi.org/10.5281/zenodo.5321298" httpUri="https://zenodo.org/record/5321298/files/figure.png" pageId="11" pageNumber="37">Figures 25, 26</figureCitation>
. Dorsum of head brownish yellow, anteclypeus and two small spots on vertex black. Pronotum with black stripe on anterior margin behind vertex. Mesonotum in anterior two-thirds of length with pair of narrow, black stripes lining smooth medial area; meso- and metanotum with distinct lateral stripes and black anterior margin interrupted in middle. All acetabula distally with black marks. Legs mainly yellowish brown to orange, profemur with two distinct stripes, protibia and apex of protarsus blackish, meso- and metafemora each with one thin black stripe at base and distal part pale; tibiae and tarsi of middle and hind legs orange to pale. Lateral areas of tergites black. Distal portions of meso- and metafemora pale. Venter pale. Meso- and metacetabula and lateral portions of tergites 2–5 (strongly reduced on tergite 1) with moderately developed, silvery hair layer (
<figureCitation id="18BE49E2FFF4871777B033BD4C4C94B9" box="[654,803,1957,1983]" captionStart="FIGURES 25–26" captionStartId="12.[151,269,1943,1967]" captionTargetBox="[169,1429,335,1929]" captionTargetId="figure-129@12.[169,1429,331,1929]" captionTargetPageId="12" captionText="FIGURES 25–26. Archaeoptilomera gen. n. (25) Habitus of apterous female of A. derlethi sp. n. (holotype, body length 8.1 mm), dorsal aspect. (26) Habitus of apterous female of A. kodadai sp. n. (holotype, body length 8.5 mm), dorsal aspect. © NHMW Hemiptera Image Collection, published with permission." figureDoi="http://doi.org/10.5281/zenodo.5321298" httpUri="https://zenodo.org/record/5321298/files/figure.png" pageId="11" pageNumber="37">Figs. 25, 26</figureCitation>
). Head (
<figureCitation id="18BE49E2FFF4871776B733BD4C8E94B9" box="[905,993,1957,1983]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="11" pageNumber="37">Fig. 27</figureCitation>
) with very large eyes and prominent antenniferous tubercles. Antenna slender, antennomere 1 subequal in length with antennomeres 2–4, antennomere 4 without sulcus. Pronotum robust. Metanotum without median furrow. Flexor side of profemur with numerous stout setae at base and one minute tooth subapically (
<figureCitation id="18BE49E2FFF38710768034A64B7993DE" box="[958,1046,190,216]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="12" pageNumber="38">Fig. 28</figureCitation>
); protarsus short, ca. 0.5 times as long as protibia; first protarsomere 1.7 times as long as second and slightly thicker. Claws or foreleg smooth. Metafemur and mesofemur subequal in length. Metacoxa without spine. Hind leg with two small but distinct claws.
</paragraph>
<caption id="D4FA05EFFFF3871075A9338F4CD294F5" ID-DOI="http://doi.org/10.5281/zenodo.5321298" ID-Zenodo-Dep="5321298" httpUri="https://zenodo.org/record/5321298/files/figure.png" pageId="12" pageNumber="38" startId="12.[151,269,1943,1967]" targetBox="[169,1429,335,1929]" targetPageId="12">
<paragraph id="803A5567FFF3871075A9338F4CD294F5" blockId="12.[151,1436,1943,2035]" pageId="12" pageNumber="38">
<emphasis id="B2F18975FFF3871075A9338F4E3294A9" bold="true" box="[151,349,1943,1967]" pageId="12" pageNumber="38">FIGURES 25–26.</emphasis>
<taxonomicName id="47852EE4FFF38710745D338F4D4B94A8" authorityName="Zettel" authorityYear="2009" box="[355,548,1943,1966]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="38" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B2F18975FFF38710745D338F4D4B94A8" box="[355,548,1943,1966]" italics="true" pageId="12" pageNumber="38">Archaeoptilomera</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF387107715338F4D1A94A9" bold="true" box="[555,629,1943,1967]" pageId="12" pageNumber="38">
<taxonomicNameLabel id="A9C2340EFFF387107715338F4D1A94A9" box="[555,629,1943,1967]" pageId="12" pageNumber="38" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
(25) Habitus of apterous female of
<taxonomicName id="47852EE4FFF3871076CF338F4B0E94A8" authorityName="Zettel" authorityYear="2009" box="[1009,1121,1943,1966]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="38" phylum="Arthropoda" rank="species" species="derlethi" status="sp. nov.">
<emphasis id="B2F18975FFF3871076CF338F4B0E94A8" box="[1009,1121,1943,1966]" italics="true" pageId="12" pageNumber="38">A. derlethi</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF387107156338F4BCC94A9" bold="true" box="[1128,1187,1943,1967]" pageId="12" pageNumber="38">
<taxonomicNameLabel id="A9C2340EFFF387107156338F4BCC94A9" box="[1128,1187,1943,1967]" pageId="12" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype, body length 8.1 mm), dorsal aspect. (26) Habitus of apterous female of
<taxonomicName id="47852EE4FFF38710761833A24CF194D7" authorityName="Zettel" authorityYear="2009" box="[806,926,1978,2001]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="38" phylum="Arthropoda" rank="species" species="kodadai" status="sp. nov.">
<emphasis id="B2F18975FFF38710761833A24CF194D7" box="[806,926,1978,2001]" italics="true" pageId="12" pageNumber="38">A. kodadai</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF38710769B33A14C8C94D7" bold="true" box="[933,995,1977,2001]" pageId="12" pageNumber="38">
<taxonomicNameLabel id="A9C2340EFFF38710769B33A14C8C94D7" box="[933,995,1977,2001]" pageId="12" pageNumber="38" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype, body length 8.5 mm), dorsal aspect. © NHMW
<taxonomicName id="47852EE4FFF38710745E33C44D7694F5" authority="Image" authorityName="Image" box="[352,537,2012,2035]" class="Insecta" kingdom="Animalia" order="Hemiptera" pageId="12" pageNumber="38" phylum="Arthropoda" rank="order">Hemiptera Image</taxonomicName>
Collection, published with permission.
</paragraph>
</caption>
<caption id="D4FA05EFFFF2871175A937514D1990CE" ID-DOI="http://doi.org/10.5281/zenodo.5321300" ID-Zenodo-Dep="5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="13" pageNumber="39" startId="13.[151,270,841,865]" targetBox="[288,1300,202,813]" targetPageId="13">
<paragraph id="803A5567FFF2871175A937514D1990CE" blockId="13.[151,1436,841,968]" pageId="13" pageNumber="39">
<emphasis id="B2F18975FFF2871175A937514E0F9067" bold="true" box="[151,352,841,865]" pageId="13" pageNumber="39">FIGURES 27–34.</emphasis>
<taxonomicName id="47852EE4FFF28711745937514D459066" authorityName="Zettel" authorityYear="2009" box="[359,554,841,864]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B2F18975FFF28711745937514D459066" box="[359,554,841,864]" italics="true" pageId="13" pageNumber="39">Archaeoptilomera</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711770C37514D119067" bold="true" box="[562,638,841,865]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711770C37514D119067" box="[562,638,841,865]" pageId="13" pageNumber="39" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
(holotypes, females). (27) Head of pronotum of
<taxonomicName id="47852EE4FFF2871171AA37514A699066" authorityName="Zettel" authorityYear="2009" box="[1172,1286,841,864]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="derlethi" status="sp. nov.">
<emphasis id="B2F18975FFF2871171AA37514A699066" box="[1172,1286,841,864]" italics="true" pageId="13" pageNumber="39">A. derlethi</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711703037514A249067" bold="true" box="[1294,1355,841,865]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711703037514A249067" box="[1294,1355,841,865]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, dorsal aspect. (28) Foreleg of
<taxonomicName id="47852EE4FFF2871174AE37744D6E9085" authorityName="Zettel" authorityYear="2009" box="[400,513,876,899]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="derlethi" status="sp. nov.">
<emphasis id="B2F18975FFF2871174AE37744D6E9085" box="[400,513,876,899]" italics="true" pageId="13" pageNumber="39">A. derlethi</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711773937734D2B9085" bold="true" box="[519,580,875,899]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711773937734D2B9085" box="[519,580,875,899]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33)
<taxonomicName id="47852EE4FFF28711779F37974C7D90A0" authorityName="Zettel" authorityYear="2009" box="[673,786,911,934]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="derlethi" status="sp. nov.">
<emphasis id="B2F18975FFF28711779F37974C7D90A0" box="[673,786,911,934]" italics="true" pageId="13" pageNumber="39">A. derlethi</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711762737964C3A90A0" bold="true" box="[793,853,910,934]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711762737964C3A90A0" box="[793,853,910,934]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(30, 32, 34)
<taxonomicName id="47852EE4FFF2871176DF37974B3A90A0" authorityName="Zettel" authorityYear="2009" box="[993,1109,911,934]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="kodadai" status="sp. nov.">
<emphasis id="B2F18975FFF2871176DF37974B3A90A0" box="[993,1109,911,934]" italics="true" pageId="13" pageNumber="39">A. kodadai</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711716337964BF690A0" bold="true" box="[1117,1177,910,934]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711716337964BF690A0" box="[1117,1177,910,934]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8.
</paragraph>
</caption>
<paragraph id="803A5567FFF2871175FB37E24EE49712" blockId="13.[151,1436,1018,1934]" box="[197,395,1018,1044]" pageId="13" pageNumber="39">Male. Unknown.</paragraph>
<paragraph id="803A5567FFF2871175FB30394C639625" blockId="13.[151,1436,1018,1934]" pageId="13" pageNumber="39">
Female. Metanotum, abdominal segments 1–6, and venter of thorax and abdomen without modifications. Abdomen moderately shortened, slightly longer than thorax (
<figureCitation id="18BE49E2FFF28711765130504B6D9764" box="[879,1026,1096,1122]" captionStart="FIGURES 25–26" captionStartId="12.[151,269,1943,1967]" captionTargetBox="[169,1429,335,1929]" captionTargetId="figure-129@12.[169,1429,331,1929]" captionTargetPageId="12" captionText="FIGURES 25–26. Archaeoptilomera gen. n. (25) Habitus of apterous female of A. derlethi sp. n. (holotype, body length 8.1 mm), dorsal aspect. (26) Habitus of apterous female of A. kodadai sp. n. (holotype, body length 8.5 mm), dorsal aspect. © NHMW Hemiptera Image Collection, published with permission." figureDoi="http://doi.org/10.5281/zenodo.5321298" httpUri="https://zenodo.org/record/5321298/files/figure.png" pageId="13" pageNumber="39">Figs. 25, 26</figureCitation>
). Medial areas of sternites slightly flattened. Connexiva not flapped mediodorsad over tergites 1–6, but slightly so over posterolateral parts of tergite 7 (
<figureCitation id="18BE49E2FFF287117437308D4EF597A9" box="[265,410,1173,1199]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="13" pageNumber="39">Figs. 31, 32</figureCitation>
). Tergites 7 and 8 simple, directed posteriad; proctiger simple, directed posteroventrad or almost ventrad (
<figureCitation id="18BE49E2FFF2871174BF30A44D7597D0" box="[385,538,1212,1238]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="13" pageNumber="39">Figs. 29, 30</figureCitation>
). Sternite 7 elongate, with short to moderately long spines at connexival corners (
<figureCitation id="18BE49E2FFF2871175C130FA4EFD97FA" box="[255,402,1250,1276]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="13" pageNumber="39">Figs. 29–32</figureCitation>
), with large, but relatively short, posteriorly rounded ventromedial lobe (
<figureCitation id="18BE49E2FFF2871171C530FA4AE197FA" box="[1275,1422,1250,1276]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="13" pageNumber="39">Figs. 33, 34</figureCitation>
). Gonocoxae 1 (
<figureCitation id="18BE49E2FFF28711747C31114EA09625" box="[322,463,1289,1315]" captionStart="FIGURES 27–34" captionStartId="13.[151,270,841,865]" captionTargetBox="[288,1300,202,813]" captionTargetId="figure-428@13.[283,1304,194,817]" captionTargetPageId="13" captionText="FIGURES 27–34. Archaeoptilomera gen. n. (holotypes, females). (27) Head of pronotum of A. derlethi sp. n., dorsal aspect. (28) Foreleg of A. derlethi sp. n., posterior aspect. (29–34) Posterior part of abdomen in (29, 30) lateral, (31, 32) dorsal, and (33, 34) ventral aspects. (29, 31, 33) A. derlethi sp. n. (30, 32, 34) A. kodadai sp. n. gc1 = gonocoxa 1; pr = proctiger; s7 = sternite 7; t7, t8 = tergite 7, 8." figureDoi="http://doi.org/10.5281/zenodo.5321300" httpUri="https://zenodo.org/record/5321300/files/figure.png" pageId="13" pageNumber="39">Figs. 29, 30</figureCitation>
) large, plate-like, exposed.
</paragraph>
<paragraph id="803A5567FFF2871175FB31374D50941C" blockId="13.[151,1436,1018,1934]" pageId="13" pageNumber="39">
<emphasis id="B2F18975FFF2871175FB31374D1B964F" bold="true" box="[197,628,1327,1353]" pageId="13" pageNumber="39">Comparative notes and discussion.</emphasis>
This new genus is so far only known from female specimens, and for that reason some aspects of its accurate position in a phylogenetic system remain uncertain. However,
<taxonomicName id="47852EE4FFF2871175A931654E109690" authorityName="Zettel" authorityYear="2009" box="[151,383,1405,1430]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B2F18975FFF2871175A931654E109690" box="[151,383,1405,1430]" italics="true" pageId="13" pageNumber="39">Archaeoptilomera</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF2871174B431654E8B9691" bold="true" box="[394,484,1405,1431]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF2871174B431654E8B9691" box="[394,484,1405,1431]" pageId="13" pageNumber="39" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
clearly belongs to the
<taxonomicName id="47852EE4FFF28711762A31654CFB9690" authorityName="Amyot and Serville" authorityYear="1843" box="[788,916,1405,1430]" class="Insecta" family="Gerridae" genus="Ptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B2F18975FFF28711762A31654CFB9690" box="[788,916,1405,1430]" italics="true" pageId="13" pageNumber="39">Ptilomera</emphasis>
</taxonomicName>
genus group (see Comparative notes of
<taxonomicName id="47852EE4FFF2871175A931BC4E4496BB" authorityName="Zettel" authorityYear="2009" box="[151,299,1444,1469]" class="Insecta" family="Gerridae" genus="Ptilomerella" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B2F18975FFF2871175A931BC4E4496BB" box="[151,299,1444,1469]" italics="true" pageId="13" pageNumber="39">Ptilomerella</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711740A31BB4EE796BB" bold="true" box="[308,392,1443,1469]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711740A31BB4EE796BB" box="[308,392,1443,1469]" pageId="13" pageNumber="39" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
). Within this group, it has strong similarities with the Himalayan monotypic subgenus
<taxonomicName id="47852EE4FFF2871175A931D34D1C96E2" authorityName="Hungerford &amp; Matsuda" authorityYear="1958" box="[151,627,1482,1508]" class="Insecta" family="Gerridae" genus="Ptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="himalayensis" subGenus="Proptilomera">
<emphasis id="B2F18975FFF2871175A931D34E6096E2" box="[151,271,1483,1508]" italics="true" pageId="13" pageNumber="39">Ptilomera</emphasis>
(
<emphasis id="B2F18975FFF28711741C31D34EAC96E2" box="[290,451,1483,1508]" italics="true" pageId="13" pageNumber="39">Proptilomera</emphasis>
)
<emphasis id="B2F18975FFF2871174E831D34D1C96E2" box="[470,627,1483,1508]" italics="true" pageId="13" pageNumber="39">himalayensis</emphasis>
</taxonomicName>
, which is morphologically distant from the species-rich
<taxonomicName id="47852EE4FFF28711701A31D34AF396E2" authorityName="Amyot and Serville" authorityYear="1843" box="[1316,1436,1483,1508]" class="Insecta" family="Gerridae" genus="Ptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B2F18975FFF28711701A31D34AF396E2" box="[1316,1436,1483,1508]" italics="true" pageId="13" pageNumber="39">Ptilomera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9C2340EFFF2871175A931E94FA3950D" box="[151,204,1521,1547]" pageId="13" pageNumber="39" sensu="stricto">s.str.</taxonomicNameLabel>
and may deserve genus rank, too.
<taxonomicName id="47852EE4FFF28711775931E94C54950C" authorityName="Zettel" authorityYear="2009" box="[615,827,1521,1546]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B2F18975FFF28711775931E94C54950C" box="[615,827,1521,1546]" italics="true" pageId="13" pageNumber="39">Archaeoptilomera</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711767D31E94CF9950D" bold="true" box="[835,918,1521,1547]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711767D31E94CF9950D" box="[835,918,1521,1547]" pageId="13" pageNumber="39" rank="genus">gen. n.</taxonomicNameLabel>
</emphasis>
can be distinguished from
<taxonomicName id="47852EE4FFF2871171E931E94A23950C" authorityName="Amyot and Serville" authorityYear="1843" box="[1239,1356,1521,1546]" class="Insecta" family="Gerridae" genus="Ptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B2F18975FFF2871171E931E94A23950C" box="[1239,1356,1521,1546]" italics="true" pageId="13" pageNumber="39">Ptilomera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9C2340EFFF28711706A31E94A18950D" box="[1364,1399,1521,1547]" pageId="13" pageNumber="39" sensu="lato">s.l.</taxonomicNameLabel>
by its small size, the absence of a medial furrow on the metanotum, the subequal length of meso- and metafemur, and by the black sublateral stripes on the mesonotum which line the smooth medial area. Moreover,
<taxonomicName id="47852EE4FFF28711701932274AF3955E" authorityName="Amyot and Serville" authorityYear="1843" box="[1319,1436,1599,1624]" class="Insecta" family="Gerridae" genus="Ptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B2F18975FFF28711701932274AF3955E" box="[1319,1436,1599,1624]" italics="true" pageId="13" pageNumber="39">Ptilomera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9C2340EFFF2871175A9327D4FA29579" box="[151,205,1637,1663]" pageId="13" pageNumber="39" sensu="stricto">s.str.</taxonomicNameLabel>
can be easily recognized by the prominent metacoxal spines. The sternite 7 of the female of
<taxonomicName id="47852EE4FFF28711701B327D4E8895A3" authorityName="Hungerford &amp; Matsuda" authorityYear="1958" class="Insecta" family="Gerridae" genus="Ptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="himalayensis" subGenus="Proptilomera">
<emphasis id="B2F18975FFF28711701B327D4AF49578" box="[1317,1435,1637,1662]" italics="true" pageId="13" pageNumber="39">Ptilomera</emphasis>
(
<emphasis id="B2F18975FFF28711759F32944E5195A3" box="[161,318,1676,1701]" italics="true" pageId="13" pageNumber="39">Proptilomera</emphasis>
)
<emphasis id="B2F18975FFF28711747032944E8895A3" box="[334,487,1676,1701]" italics="true" pageId="13" pageNumber="39">himalayensis</emphasis>
</taxonomicName>
differs strongly from both species of
<taxonomicName id="47852EE4FFF28711769E32944B1B95A3" authorityName="Zettel" authorityYear="2009" box="[928,1140,1676,1701]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B2F18975FFF28711769E32944B1B95A3" box="[928,1140,1676,1701]" italics="true" pageId="13" pageNumber="39">Archaeoptilomera</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711714532934BD295A3" bold="true" box="[1147,1213,1675,1701]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711714532934BD295A3" box="[1147,1213,1675,1701]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in the unique bifid medial lobe that embraces the proctiger, and by absence of connexival spines. The protarsus of
<taxonomicName id="47852EE4FFF2871175A932C14E1F95F4" authorityName="Zettel" authorityYear="2009" box="[151,368,1753,1778]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus" status="sp. nov.">
<emphasis id="B2F18975FFF2871175A932C14E1F95F4" box="[151,368,1753,1778]" italics="true" pageId="13" pageNumber="39">Archaeoptilomera</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711744732C14ED295F5" bold="true" box="[377,445,1753,1779]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711744732C14ED295F5" box="[377,445,1753,1779]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is pale as it is in
<taxonomicName id="47852EE4FFF2871177AA32C14CC695F4" authorityName="Zettel" authorityYear="2009" box="[660,937,1753,1778]" class="Insecta" family="Gerridae" genus="Ptilomerella" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="anderseni" status="sp. nov.">
<emphasis id="B2F18975FFF2871177AA32C14CC695F4" box="[660,937,1753,1778]" italics="true" pageId="13" pageNumber="39">Ptilomerella anderseni</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF28711768F32C14C9A95F5" bold="true" box="[945,1013,1753,1779]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF28711768F32C14C9A95F5" box="[945,1013,1753,1779]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, but the author does not know any other
<taxonomicName id="47852EE4FFF2871175E433184E1F941C" authorityName="Bianchi" authorityYear="1896" box="[218,368,1792,1818]" class="Insecta" family="Gerridae" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="subFamily" subFamily="Ptilomerinae">Ptilomerinae</taxonomicName>
with that feature.
</paragraph>
</subSubSection>
<subSubSection id="C89F06ECFFF2871175FB333E4EA59446" box="[197,458,1830,1856]" pageId="13" pageNumber="39" type="distribution">
<paragraph id="803A5567FFF2871175FB333E4EA59446" blockId="13.[151,1436,1018,1934]" box="[197,458,1830,1856]" pageId="13" pageNumber="39">
<emphasis id="B2F18975FFF2871175FB333E4E0A9446" bold="true" box="[197,357,1830,1856]" pageId="13" pageNumber="39">Distribution.</emphasis>
Borneo.
</paragraph>
</subSubSection>
<subSubSection id="C89F06ECFFF2871175FB33554BDE948B" pageId="13" pageNumber="39" type="etymology">
<paragraph id="803A5567FFF2871175FB33554A249461" blockId="13.[151,1436,1018,1934]" box="[197,1355,1869,1895]" pageId="13" pageNumber="39">
<emphasis id="B2F18975FFF2871175FB33554E3D9461" bold="true" box="[197,338,1869,1895]" pageId="13" pageNumber="39">Etymology.</emphasis>
The generic epithet refers to the close relationships with
<taxonomicName id="47852EE4FFF2871176CC33554B089460" authorityName="Amyot and Serville" authorityYear="1843" box="[1010,1127,1869,1894]" class="Insecta" family="Gerridae" genus="Ptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="genus">
<emphasis id="B2F18975FFF2871176CC33554B089460" box="[1010,1127,1869,1894]" italics="true" pageId="13" pageNumber="39">Ptilomera</emphasis>
</taxonomicName>
. Gender: feminine.
</paragraph>
<paragraph id="803A5567FFF2871175FB336B4BDE948B" blockId="13.[151,1436,1018,1934]" box="[197,1201,1907,1934]" pageId="13" pageNumber="39">
<emphasis id="B2F18975FFF2871175FB336B4EF7948B" bold="true" box="[197,408,1907,1933]" pageId="13" pageNumber="39">Included species.</emphasis>
<taxonomicName id="47852EE4FFF28711749E336C4DB9948B" authorityName="Zettel" authorityYear="2009" box="[416,726,1908,1933]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="derlethi" status="sp. nov.">
<emphasis id="B2F18975FFF28711749E336C4DB9948B" box="[416,726,1908,1933]" italics="true" pageId="13" pageNumber="39">Archaeoptilomera derlethi</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF2871177E3336B4C70948B" bold="true" box="[733,799,1907,1933]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF2871177E3336B4C70948B" box="[733,799,1907,1933]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="47852EE4FFF287117610336C4B07948B" authorityName="Zettel" authorityYear="2009" box="[814,1128,1908,1933]" class="Insecta" family="Gerridae" genus="Archaeoptilomera" kingdom="Animalia" order="Hemiptera" pageId="13" pageNumber="39" phylum="Arthropoda" rank="species" species="kodadai" status="sp. nov.">
<emphasis id="B2F18975FFF287117610336C4B07948B" box="[814,1128,1908,1933]" italics="true" pageId="13" pageNumber="39">Archaeoptilomera kodadai</emphasis>
</taxonomicName>
<emphasis id="B2F18975FFF287117151336B4BDE948B" bold="true" box="[1135,1201,1907,1933]" pageId="13" pageNumber="39">
<taxonomicNameLabel id="A9C2340EFFF287117151336B4BDE948B" box="[1135,1201,1907,1933]" pageId="13" pageNumber="39" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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