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<document id="B1EDC6A0C0DAFFEEF9B6C1A79EBEE587" ID-DOI="10.5281/zenodo.177736" ID-GBIF-Dataset="0b4f2b31-604a-464e-a680-96aa0d2e170b" ID-ISSN="1175-5326" ID-Zenodo-Dep="177736" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459871882067" checkinUser="plazi" docAuthor="Shear, William A. &amp; Krejca, Jean K." docDate="2007" docId="03E0612EFF9EFFEFFF02CFD22822F820" docLanguage="en" docName="zt01532p039.pdf" docOrigin="Zootaxa 1532" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Amplaria eldora Chamberlin" docType="treatment" docVersion="11" lastPageNumber="27" masterDocId="FFD91956FF9AFFEBFF95CD47292FFFD4" masterDocTitle="Revalidation of the milliped genus Amplaria Chamberlin 1941 (Diplopoda, Chordeumatida, Striariidae), and description of two new species from caves in Sequoia and Kings Canyon National Parks, California" masterLastPageNumber="39" masterPageNumber="23" pageNumber="27" updateTime="1698226697101" updateUser="plazi">
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<mods:title id="1B55DF9D4F5CFA151C030BBFB854A814">Revalidation of the milliped genus Amplaria Chamberlin 1941 (Diplopoda, Chordeumatida, Striariidae), and description of two new species from caves in Sequoia and Kings Canyon National Parks, California</mods:title>
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<mods:namePart id="A125C62F50DED2E2504DBA777C3FAA00">Shear, William A.</mods:namePart>
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<treatment id="03E0612EFF9EFFEFFF02CFD22822F820" ID-DOI="http://doi.org/10.5281/zenodo.5628955" ID-GBIF-Taxon="119369694" ID-Zenodo-Dep="5628955" LSID="urn:lsid:plazi:treatment:03E0612EFF9EFFEFFF02CFD22822F820" httpUri="http://treatment.plazi.org/id/03E0612EFF9EFFEFFF02CFD22822F820" lastPageNumber="27" pageId="4" pageNumber="27">
<subSubSection id="C35383B3FF9EFFEFFF02CFD2282BFD06" pageId="4" pageNumber="27" type="nomenclature">
<paragraph id="8BF6D038FF9EFFEFFF02CFD22B20FD7B" blockId="4.[151,527,661,722]" box="[151,527,661,687]" pageId="4" pageNumber="27">
<heading id="D0BE6754FF9EFFEFFF02CFD22B20FD7B" bold="true" box="[151,527,661,687]" fontSize="11" level="1" pageId="4" pageNumber="27" reason="1">
<taxonomicName id="4C49ABBBFF9EFFEFFF02CFD22B20FD7B" ID-CoL="846LB" authority="Chamberlin" authorityName="Chamberlin" box="[151,527,661,687]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eldora">
<emphasis id="B93D0C2AFF9EFFEFFF02CFD22B20FD7B" bold="true" box="[151,527,661,687]" pageId="4" pageNumber="27">
<emphasis id="B93D0C2AFF9EFFEFFF02CFD22872FD7B" bold="true" box="[151,349,661,687]" italics="true" pageId="4" pageNumber="27">Amplaria eldora</emphasis>
(Chamberlin)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF6D038FF9EFFEFFF02CFFF282BFD06" blockId="4.[151,527,661,722]" box="[151,260,696,722]" pageId="4" pageNumber="27">
<figureCitation id="1372CCBDFF9EFFEFFF02CFFF282BFD06" box="[151,260,696,722]" captionStart="FIGURES 1 3" captionStartId="5.[151,269,929,953]" captionTargetBox="[235,1318,210,907]" captionTargetId="figure@5.[226,1357,194,908]" captionTargetPageId="5" captionText="FIGURES 1 3. Amplaria eldora, Grapevine Cave specimen, male. 1. Right gonopod, lateral view; aac, anterior angiocoxite; pac, posterior angiocoxite; llcc, lateral lobe of colpocoxite; f, flagellocoxite. 2. Posterior angiocoxite of right gonopod, mesal view. 3. Tip of sheathing branch of posterior angiocoxite, mesal view." httpUri="https://zenodo.org/record/177737/files/figure.png" pageId="4" pageNumber="27">Figs. 1, 2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35383B3FF9EFFEFFF02CE452822F820" pageId="4" pageNumber="27" type="description">
<paragraph id="8BF6D038FF9EFFEFFF02CE452A0EFC90" blockId="4.[151,1437,770,2036]" pageId="4" pageNumber="27">
<emphasis id="B93D0C2AFF9EFFEFFF02CE4529CEFCCF" box="[151,225,770,795]" italics="true" pageId="4" pageNumber="27">
<typeStatus id="54F26E9AFF9EFFEFFF02CE4529F8FCCF" box="[151,215,770,795]" pageId="4" pageNumber="27">Types</typeStatus>
:
</emphasis>
Three female
<typeStatus id="54F26E9AFF9EFFEFFE05CE4528D8FCC8" box="[400,503,770,796]" pageId="4" pageNumber="27" type="syntype">syntypes</typeStatus>
from Crystal Cosumnes Cave, Eldorado Co., California, collected by A. Lang and G. Lange,
<date id="FFF7F6F8FF9EFFEFFED4CE6D2B2AFC90" box="[321,517,810,836]" pageId="4" pageNumber="27" value="1952-02-02">2 February 1952</date>
, whereabouts unknown.
</paragraph>
<paragraph id="8BF6D038FF9EFFEFFF53CE152DD3FB88" blockId="4.[151,1437,770,2036]" pageId="4" pageNumber="27">
<emphasis id="B93D0C2AFF9EFFEFFF53CE15283CFCBF" box="[198,275,850,875]" italics="true" pageId="4" pageNumber="27">Notes:</emphasis>
<bibRefCitation id="EFD8ADC9FF9EFFEFFE8FCE152B28FCB8" author="Chamberlin" box="[282,519,850,876]" pageId="4" pageNumber="38" refString="Chamberlin, R. V. (1953 a) Two new millipeds taken in California Caves. Entomological News, 64, 93 - 95." type="journal article" year="1953" yearSuffix="a">Chamberlin (1953a)</bibRefCitation>
did not formally designate a
<typeStatus id="54F26E9AFF9EFFEFFCCBCE152AEBFCB8" box="[862,964,850,876]" pageId="4" pageNumber="27" type="holotype">holotype</typeStatus>
from among the three females available to him, and these specimens cannot be located; they are not in the USNM, where
<typeStatus id="54F26E9AFF9EFFEFFBF8CE3D2D8FFC40" box="[1133,1184,890,916]" pageId="4" pageNumber="27">type</typeStatus>
material Chamberlin retained in his personal collection is now deposited. The specimens were part of a collection made by Edward Danehy and others in
<date id="FFF7F6F8FF9EFFEFFE0BCE8D2B7DFC30" box="[414,594,970,996]" pageId="4" pageNumber="27" value="1950" valueMax="1951">1950 and 1951</date>
, and sent to Chamberlin for identification. The USNM does contain a vial labeled “
<taxonomicName id="4C49ABBBFF9EFFEFFEA1CEB52B57FBD8" authority="Chamberlin" authorityName="Chamberlin" box="[308,632,1010,1036]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eldora">
<emphasis id="B93D0C2AFF9EFFEFFEA1CEB528CCFBDF" box="[308,483,1010,1035]" italics="true" pageId="4" pageNumber="27">Striaria eldora</emphasis>
Chamberlin
</taxonomicName>
,
<typeStatus id="54F26E9AFF9EFFEFFD11CEB52A32FBD8" box="[644,797,1010,1036]" pageId="4" pageNumber="27" type="paratype">PARATYPE</typeStatus>
.” This single male specimen is from Grapevine Cave, Calaveras Co., California, and was collected
<date id="FFF7F6F8FF9EFFEFFD33C95D2A65FBE0" box="[678,842,1050,1076]" pageId="4" pageNumber="27" value="1951-04-17">17 April 1951</date>
by E. Jacobus. Morley Hardaker, of Davis, California, kindly provided us with information about Crystal Cosumnes Cave and Grapevine Cave:
</paragraph>
<paragraph id="8BF6D038FF9EFFEFFF53C92D2D7CFAA0" blockId="4.[151,1437,770,2036]" pageId="4" pageNumber="27">
“Crystal Cosumnes cave is near the little town of Fairplay in El Dorado County, and is located above the Middle Fork of the Cosumnes River…at about
<quantity id="4CB17DDDFF9EFFEFFD56C9D52A1DFB78" box="[707,818,1170,1196]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="4" pageNumber="27" unit="ft" value="2000.0">2000 feet</quantity>
elevation….Grapevine Cave is about 36 air miles in a southerly direction and located about
<quantity id="4CB17DDDFF9EFFEFFDF3C9FD2B93FB00" box="[614,700,1210,1236]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="4" pageNumber="27" unit="mi" value="3.0">3 miles</quantity>
east of the little town of Vallecito in Calaveras County. It is in a karst area high above and on the north side of the Stanislaus….Unlike Crystal Cosumnes its passages are maybe
<quantity id="4CB17DDDFF9EFFEFFF78C84D28AFFAF0" box="[237,384,1290,1316]" metricMagnitude="0" metricUnit="m" metricValue="7.62" metricValueMax="9.144" metricValueMin="6.096" pageId="4" pageNumber="27" unit="ft" value="25.0" valueMax="30.0" valueMin="20.0">20 to 30 feet</quantity>
below the surface, and many of its ceilings contain tree roots, some up to a couple feet in length. Grapevine is one of our richer caves in containing cave fauna. Millipedes and salamanders are commonly found here. I have personally seen millipedes in both caves on most trips.”
</paragraph>
<paragraph id="8BF6D038FF9EFFEFFF53C8C52D9DF8A8" blockId="4.[151,1437,770,2036]" pageId="4" pageNumber="27">
In a later paper,
<bibRefCitation id="EFD8ADC9FF9EFFEFFE17C8C52B5FFA48" author="Chamberlin" box="[386,624,1410,1436]" pageId="4" pageNumber="38" refString="Chamberlin, R. V. (1953 b) Six new American millipeds, with notes on several cave-dwelling species. Proceedings of the Biological Society of Washington, 66, 67 - 72." type="journal article" year="1953" yearSuffix="b">Chamberlin (1953b)</bibRefCitation>
mentioned this Grapevine Cave collection without specifying the sex or number of specimens, or designating the specimen as a
<typeStatus id="54F26E9AFF9EFFEFFCD9C8ED2A51FA10" box="[844,894,1450,1476]" pageId="4" pageNumber="27">type</typeStatus>
of any sort, so the museum labeling of it as a
<typeStatus id="54F26E9AFF9EFFEFFF02C89529D3FA38" box="[151,252,1490,1516]" pageId="4" pageNumber="27" type="paratype">paratype</typeStatus>
has no formal standing. It would be a particularly maddening aspect of Chamberlins often capricious work if he had a male available but chose to describe the species from females. And why did he decide to describe two of the species from the Danehy collection separately (
<bibRefCitation id="EFD8ADC9FF9EFFEFFC7ECB652DE1F9E8" author="Chamberlin" box="[1003,1230,1570,1596]" pageId="4" pageNumber="38" refString="Chamberlin, R. V. (1953 a) Two new millipeds taken in California Caves. Entomological News, 64, 93 - 95." type="journal article" year="1953" yearSuffix="a">Chamberlin 1953a</bibRefCitation>
), then just a few months later mix the descriptions of two more, plus more records of the first two, into a paper that was fundamentally about material from
<collectingCountry id="F35E90A8FF9EFFEFFE62CB352B46F958" box="[503,617,1650,1676]" name="Honduras" pageId="4" pageNumber="27">Honduras</collectingCountry>
(
<bibRefCitation id="EFD8ADC9FF9EFFEFFDECCB352A77F958" author="Chamberlin" box="[633,856,1650,1676]" pageId="4" pageNumber="38" refString="Chamberlin, R. V. (1953 b) Six new American millipeds, with notes on several cave-dwelling species. Proceedings of the Biological Society of Washington, 66, 67 - 72." type="journal article" year="1953" yearSuffix="b">Chamberlin 1953b</bibRefCitation>
)? Furthermore, in a checklist published in 1958, only five years after reporting the male
<taxonomicName id="4C49ABBBFF9EFFEFFDFDCBDC2B9BF960" box="[616,692,1691,1716]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eldora">
<emphasis id="B93D0C2AFF9EFFEFFDFDCBDC2B9BF960" box="[616,692,1691,1716]" italics="true" pageId="4" pageNumber="27">eldora</emphasis>
</taxonomicName>
specimen from Grapevine Cave, Chamberlin stated that
<taxonomicName id="4C49ABBBFF9EFFEFFADACBDC2CB4F960" box="[1359,1435,1691,1716]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eldora">
<emphasis id="B93D0C2AFF9EFFEFFADACBDC2CB4F960" box="[1359,1435,1691,1716]" italics="true" pageId="4" pageNumber="27">eldora</emphasis>
</taxonomicName>
was “known only from the
<typeStatus id="54F26E9AFF9EFFEFFE7CCB852B34F908" box="[489,539,1730,1756]" pageId="4" pageNumber="27">type</typeStatus>
locality (
<bibRefCitation id="EFD8ADC9FF9EFFEFFD04CB852D3DF908" author="Chamberlin" box="[657,1042,1730,1756]" pageId="4" pageNumber="38" refString="Chamberlin, R. V., and Hoffman, R. L. 1958. Checklist of the millipeds of North America. United States National Museum Bulletin, 212, 1 - 236." type="journal article" year="1958">Chamberlin and Hoffman 1958</bibRefCitation>
).” Chamberlins (1953) sketchy description of female
<taxonomicName id="4C49ABBBFF9EFFEFFE0DCBAC28CBF8D0" box="[408,484,1771,1796]" class="Diplopoda" family="Striariidae" genus="Striaria" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eldora">
<emphasis id="B93D0C2AFF9EFFEFFE0DCBAC28CBF8D0" box="[408,484,1771,1796]" italics="true" pageId="4" pageNumber="27">eldora</emphasis>
</taxonomicName>
could apply to almost any species of medium-sized striariid, and so is of no help in determining the identity of this specimen. Until males from Crystal Cosumnes Cave appear, no harm is done by considering the Grapevine Cave male to be
<taxonomicName id="4C49ABBBFF9EFFEFFD6ECA7C2A93F880" box="[763,956,1851,1876]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eldora">
<emphasis id="B93D0C2AFF9EFFEFFD6ECA7C2A93F880" box="[763,956,1851,1876]" italics="true" pageId="4" pageNumber="27">Amplaria eldora</emphasis>
</taxonomicName>
. However, as stated above, it remains to be proven that
<taxonomicName id="4C49ABBBFF9EFFEFFED3CA2428BDF8A8" box="[326,402,1891,1916]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eldora">
<emphasis id="B93D0C2AFF9EFFEFFED3CA2428BDF8A8" box="[326,402,1891,1916]" italics="true" pageId="4" pageNumber="27">eldora</emphasis>
</taxonomicName>
is a junior synonym of
<emphasis id="B93D0C2AFF9EFFEFFD3CCA242BD1F8A8" box="[681,766,1891,1916]" italics="true" pageId="4" pageNumber="27">
<taxonomicName id="4C49ABBBFF9EFFEFFD3CCA242BD5F8A8" box="[681,762,1891,1916]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="eutypa">eutypa</taxonomicName>
,
</emphasis>
though we strongly suspect that it is.
</paragraph>
<paragraph id="8BF6D038FF9EFFEFFF53CACD2822F820" blockId="4.[151,1437,770,2036]" pageId="4" pageNumber="27">
The gonopods of the Grapevine Cave male are illustrated in figs. 1 and 2; unfortunately the specimen was in poor condition, but the distal parts of the gonopods are clearly distinct from other species now placed in
<emphasis id="B93D0C2AFF9EFFEFFF02CA9C2822F820" box="[151,269,2011,2036]" italics="true" pageId="4" pageNumber="27">
<taxonomicName id="4C49ABBBFF9EFFEFFF02CA9C2825F820" box="[151,266,2011,2036]" class="Diplopoda" family="Striariidae" genus="Amplaria" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">Amplaria</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
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