treatments-xml/data/03/A3/87/03A387B9FFBAFF97FDD61C95FBEBA103.xml
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<document id="3108AA62028D9126ED982838FD73D4FC" ID-DOI="10.5852/ejt.2024.927.2475" ID-ISSN="2118-9773" ID-Zenodo-Dep="10853688" ID-ZooBank="B6E43365-FACA-49BA-8CCD-77E4BF8F0016" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1711115020933" checkinUser="plazi" docAuthor="Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De &amp; Gittenberger, Edmund" docDate="2024" docId="03A387B9FFBAFF97FDD61C95FBEBA103" docLanguage="en" docName="EJT.2024.927.1-163.pdf" docOrigin="European Journal of Taxonomy 927" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cochlostoma (Turritus) macei Cochl." docType="treatment" docVersion="9" lastPageNumber="127" masterDocId="FF9AFFC1FFC4FFE9FFD91D5FFFB5A629" masterDocTitle="Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)" masterLastPageNumber="163" masterPageNumber="1" pageNumber="127" updateTime="1711726029180" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="350197C11E64EB04DB24193745D505AE">Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)</mods:title>
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<mods:namePart id="41C37EA554DFBD07BF1BF07F8B08AD24">Zallot, Enrico</mods:namePart>
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<mods:namePart id="299E157A09516B9D5762D47CF2C3A89D">Kamchev, Panche</mods:namePart>
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<mods:namePart id="EBF53CA7B54F8848330AD28870A81F70">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="B7A8F8105D6435719A8F476F9592037D">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. &amp; Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands.</mods:affiliation>
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<mods:namePart id="8FB3CEB6C802EDE80621606541C058C3">Fehér, Zoltán</mods:namePart>
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<mods:namePart id="C2DA5987586CAF8C2308EA7040B3EF78">Mattia, Willy De</mods:namePart>
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<mods:affiliation id="CE3E832BF2A0771C1F4E260D19223E4F">Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria.</mods:affiliation>
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<mods:namePart id="4070C319A95DE65E52475113521D19AE">Gittenberger, Edmund</mods:namePart>
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<mods:affiliation id="3C2690C67A68C9705A00C5E1A63DAF51">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.</mods:affiliation>
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<mods:title id="136F693CCD61F42B7A5D3E08D6129A6D">European Journal of Taxonomy</mods:title>
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<mods:date id="B0E6808978F0DA8F49B41F3800666C5A">2024</mods:date>
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<taxonomicName id="4C0A4D2CFFBAFF97FDD61C95FB91A7CC" ID-CoL="WNNQ" authority="(Bourguignat, 1870)" authorityName="Cochl." baseAuthorityName="Bourguignat" baseAuthorityYear="1870" box="[527,1060,458,485]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="126" pageNumber="127" phylum="Mollusca" rank="species" species="macei" subGenus="Turritus">
<emphasis id="B97EEABDFFBAFF97FDD61C95FD18A7CC" bold="true" box="[527,685,458,485]" italics="true" pageId="126" pageNumber="127">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFFBAFF97FD661C94FD60A7CC" bold="true" box="[703,725,459,485]" italics="true" pageId="126" pageNumber="127">T.</emphasis>
)
<emphasis id="B97EEABDFFBAFF97FD3F1C94FC9BA7CC" bold="true" box="[742,814,459,485]" italics="true" pageId="126" pageNumber="127">macei</emphasis>
(
<bibRefCitation id="EF9B4B5EFFBAFF97FCE41C94FBA9A7CC" author="Bourguignat" box="[829,1052,459,485]" firstAuthor="Bourguignat" pageId="126" pageNumber="127" refId="ref36496" refString="Bourguignat J. - R. 1870. Description d'especes nouvelles de Mollusques terrestres du departement des Alpes-Maritimes. L. Maccarry, Cannes." type="book" year="1870">Bourguignat, 1870</bibRefCitation>
)
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</paragraph>
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<figureCitation id="13312A2AFFBAFF97FD591CB1FD53A420" box="[640,742,494,521]" captionStart="Fig" captionStartId="118.[189,232,1856,1882]" captionTargetBox="[189,1398,1147,1827]" captionTargetId="figure-199@118.[189,1398,1147,1827]" captionTargetPageId="118" captionText="Fig. 105. Distribution of samples with undetermined position in the phylogenetic trees: white= Cochlostoma (T.) macei (Bourguignat, 1870); purple =C. (T.) sturanii (Wagner, 1897); orange=C. (T.) kleciaki (Braun, 1887); yellow=C. (T.) arnautorum (Wagner, 1906); red =C. (T.) atlanticum (Bourguignat, 1868); green =C. (T.) euboicum (Westerlund, 1885); blue =C. (T.) martensianum (Möllendorff, 1873); magenta= C. (T.) simrothi (Caziot, 1908); cyan =C. (T.) fontqueri Haas, 1924." figureDoi="http://doi.org/10.5281/zenodo.10853918" httpUri="https://zenodo.org/record/10853918/files/figure.png" pageId="126" pageNumber="127">Figs 105</figureCitation>
(white dots), 111
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<treatmentCitation id="0AAB10BEFFBAFF97FF641F6FFD04A463" author="Bourguignat J. - R." box="[189,689,559,586]" page="57" pageId="126" pageNumber="127" year="1870">
<taxonomicName id="4C0A4D2CFFBAFF97FF641F6FFD04A463" authority="Bourguignat 1870: 56 - 57" authorityName="Bourguignat" authorityPageNumber="56 - 57" authorityYear="1870" box="[189,689,559,586]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="126" pageNumber="127" phylum="Mollusca" rank="species" species="macei">
<emphasis id="B97EEABDFFBAFF97FF641F6FFECCA460" box="[189,377,560,585]" italics="true" pageId="126" pageNumber="127">Pomatias macei</emphasis>
<bibRefCitation id="EF9B4B5EFFBAFF97FE581F70FD04A463" author="Bourguignat" box="[385,689,559,586]" firstAuthor="Bourguignat" page="56 - 57" pageId="126" pageNumber="127" refId="ref36496" refString="Bourguignat J. - R. 1870. Description d'especes nouvelles de Mollusques terrestres du departement des Alpes-Maritimes. L. Maccarry, Cannes." type="book" year="1870">Bourguignat 1870: 5657</bibRefCitation>
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.
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<typeStatus id="54B1880DFFBAFF97FF641F2EFEB3A4A2" box="[189,262,625,651]" pageId="126" pageNumber="127">Types</typeStatus>
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<paragraph id="8BB536AFFFBAFF97FF641FFFFE99A493" blockId="126.[189,300,672,698]" box="[189,300,672,698]" pageId="126" pageNumber="127">Not seen.</paragraph>
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<emphasis id="B97EEABDFFBAFF97FF641FBFFE3BA4D3" bold="true" box="[189,398,736,762]" pageId="126" pageNumber="127">Other specimens</emphasis>
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<materialsCitation id="3B623CF2FFBAFF97FF641E50FBE6A500" box="[189,1107,783,810]" collectionCode="RMNH" collectorName="RMNH" country="France" latitude="43.7305" location="Gourdon" longLatPrecision="40" longitude="6.959" pageId="126" pageNumber="127" specimenCode="EZ0399" specimenCount="1">
<collectingCountry id="F31D763FFFBAFF97FF641E50FE84A500" box="[189,305,783,809]" name="France" pageId="126" pageNumber="127">FRANCE</collectingCountry>
• 1-
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;
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,
<geoCoordinate id="EE3E5068FFBAFF97FD521E50FD4DA503" box="[651,760,783,810]" degrees="6.959" direction="east" orientation="longitude" pageId="126" pageNumber="127" precision="55" value="6.959">6.959° E</geoCoordinate>
; 1976;
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leg.;
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2-
<location id="8ED56074FFBAFF97FB4F1E50FAC7A503" LSID="urn:lsid:plazi:treatment:03A387B9FFBAFF97FDD61C95FBEBA103:8ED56074FFBAFF97FB4F1E50FAC7A503" box="[1174,1394,783,810]" country="France" latitude="43.9471" longLatPrecision="6" longitude="6.2267" name="Gorge du Trevans" pageId="126" pageNumber="127">Gorge du Trevans</location>
;
<geoCoordinate id="EE3E5068FFBAFF97FF641E6CFEF2A564" box="[189,327,819,845]" degrees="43.9471" direction="north" orientation="latitude" pageId="126" pageNumber="127" precision="5" value="43.9471">43.9471° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FFBAFF97FE8A1E6DFE79A564" box="[339,460,818,845]" degrees="6.2267" direction="east" orientation="longitude" pageId="126" pageNumber="127" precision="5" value="6.2267">6.2267° E</geoCoordinate>
; 2014;
<collectorName id="26FF5379FFBAFF97FDFA1E6DFDDCA565" box="[547,617,818,844]" pageId="126" pageNumber="127">Zallot</collectorName>
leg.;
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.
</paragraph>
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locality
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<collectingCountry id="F31D763FFFBAFF97FF641EFCFE84A594" box="[189,305,931,957]" name="France" pageId="126" pageNumber="127">FRANCE</collectingCountry>
• Drôme, Grasse, Saint Vallier.
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<emphasis id="B97EEABDFFBAFF97FF641EBBFEFBA5D7" bold="true" box="[189,334,996,1022]" pageId="126" pageNumber="127">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFFBAFF97FF64194CFABCA279" blockId="126.[189,1399,1042,1104]" pageId="126" pageNumber="127">
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. Closely spaced riblets on last part of protoconh. Teleoconch spotless with moderately strong ribs. Relatively strong lip with quite abruptly inwardly curved columellar lobe covering umbilicus.
</paragraph>
<paragraph id="8BB536AFFFBAFF97FF641927FB3CA29C" blockId="126.[189,1398,1143,1205]" pageId="126" pageNumber="127">
<smallCapsWord id="8D53A073FFBAFF97FF641927FED9A2B9" baselines="1163,1163" box="[189,364,1144,1170]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="126" pageNumber="127">MEASUREMENTS</smallCapsWord>
.
<specimenCount id="9D0CFD26FFBAFF97FEA41928FE71A2BB" box="[381,452,1143,1170]" count="3" pageId="126" pageNumber="127" type="female">3 ♀♀</specimenCount>
: whorls= 8.39.1, H =
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, H/W=2.793.1, roundness =0.10.14, ribs incl.= 5759°, apert. incl.= 1719°, ribs/mm 1
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wh.= 513, ribs/mm 4
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wh.=715.
</paragraph>
<paragraph id="8BB536AFFFBAFF97FF641983FE2AA376" blockId="126.[189,1399,1244,1375]" pageId="126" pageNumber="127">
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<smallCapsWord id="8D53A073FFBAFF97FEC619BEFEC3A2DD" baselines="1263" box="[287,374,1249,1268]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genital" pageId="126" pageNumber="127">GENITAL</smallCapsWord>
<smallCapsWord id="8D53A073FFBAFF97FE5919BEFE61A2DD" baselines="1263" box="[384,468,1249,1268]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="organs" pageId="126" pageNumber="127">ORGANS</smallCapsWord>
.
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analysed. Slightly posterior connection of pedunculus to bursa copulatrix. Short, roughly oval seminal receptacle with undefined basis. 24 loops clustered beside apex of seminal receptacle. Junction of uterus gland far from connection of pedunculus of bursa copulatrix with distal oviduct.
</paragraph>
</subSubSection>
<subSubSection id="C3106524FFBAFF97FF6418D8FBEBA103" pageId="126" pageNumber="127" type="discussion">
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<emphasis id="B97EEABDFFBAFF97FF6418D8FE9AA388" bold="true" box="[189,303,1415,1441]" pageId="126" pageNumber="127">Remarks</emphasis>
</heading>
</paragraph>
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Our sample closest to the
<typeStatus id="54B1880DFFBAFF97FE3F18E9FDACA3F9" box="[486,537,1462,1488]" pageId="126" pageNumber="127">type</typeStatus>
locality originates from Gourdon, ca
<quantity id="4CF29B4AFFBAFF97FC1218EAFBA6A3E6" box="[971,1043,1461,1487]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="126" pageNumber="127" unit="km" value="10.0">10 km</quantity>
east of Saint Vallier. The other sample, from the Gorge du Trevans, is located
<quantity id="4CF29B4AFFBAFF97FD041887FC93A3DB" box="[733,806,1496,1523]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="126" pageNumber="127" unit="km" value="50.0">50 km</quantity>
northeast of the
<typeStatus id="54B1880DFFBAFF97FC301886FBA9A3DA" box="[1001,1052,1497,1523]" pageId="126" pageNumber="127">type</typeStatus>
locality. Only of the latter we were able to analyse the genitals.
</paragraph>
<paragraph id="8BB536AFFFBAFF97FF641B62FBEBA103" blockId="126.[189,1399,1597,1835]" pageId="126" pageNumber="127">
The shell morphology and the female genitalia suggest that this species belongs to
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<emphasis id="B97EEABDFFBAFF97FB791B61FB4AA07E" box="[1184,1279,1598,1623]" italics="true" pageId="126" pageNumber="127">Turritus</emphasis>
</taxonomicName>
. Only H3 was sequenced and the results support this conclusion. Based on some similarity of the female genital morphology with that of
<taxonomicName id="4C0A4D2CFFBAFF97FE341BDBFD1EA0B4" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1890" box="[493,683,1667,1694]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="126" pageNumber="127" phylum="Mollusca" rank="species" species="sardoum" subGenus="Turritus">
<emphasis id="B97EEABDFFBAFF97FE341BDBFDBDA0B7" box="[493,520,1668,1694]" italics="true" pageId="126" pageNumber="127">C.</emphasis>
(
<emphasis id="B97EEABDFFBAFF97FDC51BDBFD87A0B4" box="[540,562,1668,1693]" italics="true" pageId="126" pageNumber="127">T.</emphasis>
)
<emphasis id="B97EEABDFFBAFF97FD9C1BDCFD1EA0B4" box="[581,683,1667,1693]" italics="true" pageId="126" pageNumber="127">sardoum</emphasis>
</taxonomicName>
, it could belong to the same subclade within clade B. This would be interesting in a paleogographic perspective because it would be the remnant on the European plate of the closest relative to the
<taxonomicName id="4C0A4D2CFFBAFF97FD9F1B95FD19A0CD" authorityName="Cochl." baseAuthorityName="Westerlund" baseAuthorityYear="1890" box="[582,684,1738,1764]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="126" pageNumber="127" phylum="Mollusca" rank="species" species="sardoum">
<emphasis id="B97EEABDFFBAFF97FD9F1B95FD19A0CD" box="[582,684,1738,1764]" italics="true" pageId="126" pageNumber="127">sardoum</emphasis>
</taxonomicName>
,
<emphasis id="B97EEABDFFBAFF97FD631B95FCD1A0CD" box="[698,868,1738,1764]" italics="true" pageId="126" pageNumber="127">
<taxonomicName id="4C0A4D2CFFBAFF97FD631B95FCA5A0CD" baseAuthorityName="Westerlund" baseAuthorityYear="1879" box="[698,784,1738,1764]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="126" pageNumber="127" phylum="Mollusca" rank="species" species="agriotes">agriotes</taxonomicName>
/
<taxonomicName id="4C0A4D2CFFBAFF97FCC41B95FCD1A0CD" authorityName="Cochl." baseAuthorityName="Benoit" baseAuthorityYear="1876" box="[797,868,1738,1764]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="126" pageNumber="127" phylum="Mollusca" rank="species" species="affine">affine</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C0A4D2CFFBAFF97FC7A1B95FB95A0CD" baseAuthorityName="Bourguignat" baseAuthorityYear="1868" box="[931,1056,1738,1764]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="126" pageNumber="127" phylum="Mollusca" rank="species" species="atlanticum">
<emphasis id="B97EEABDFFBAFF97FC7A1B95FB95A0CD" box="[931,1056,1738,1764]" italics="true" pageId="126" pageNumber="127">atlanticum</emphasis>
</taxonomicName>
taxa (see Discussion), which originated on the southern border of the European plates before the geological eras which led to the current shape of the western Mediterranean Sea and the lands that sourround it.
</paragraph>
</subSubSection>
</treatment>
</document>