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<mods:title id="01668950DC8E9AFD2CF4B780E7C9D074">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
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<subSubSection id="C310651CFFCEFFA7FF70FE37FE42C7CF" box="[151,407,476,502]" pageId="40" pageNumber="41" type="nomenclature">
<paragraph id="8BB53697FFCEFFA7FF70FE37FE42C7CF" blockId="40.[151,407,476,502]" box="[151,407,476,502]" pageId="40" pageNumber="41">
<heading id="D0FD81FBFFCEFFA7FF70FE37FE42C7CF" bold="true" box="[151,407,476,502]" fontSize="11" level="1" pageId="40" pageNumber="41" reason="1">
<materialsCitation id="3B623CCAFFCEFFA7FF70FE37FE42C7CF" ID-GBIF-Occurrence="1502685008" box="[151,407,476,502]" location="Papua" pageId="40" pageNumber="41" specimenCount="1" stateProvince="Papua">
<emphasis id="B97EEA85FFCEFFA7FF70FE37FE42C7CF" bold="true" box="[151,407,476,502]" pageId="40" pageNumber="41">
<taxonomicName id="4C0A4D14FFCEFFA7FF70FE37FF32C7CF" box="[151,231,476,502]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="43" phylum="Arthropoda" rank="species" species="papua">
<collectingRegion id="49CEF875FFCEFFA7FF70FE37FF32C7CF" box="[151,231,476,502]" country="Indonesia" name="Papua Barat" pageId="40" pageNumber="41">Papua</collectingRegion>
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species group
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<emphasis id="B97EEA85FFCEFFA7FF70FDC9FE86C402" bold="true" box="[151,339,546,571]" pageId="40" pageNumber="41">Adult diagnosis</emphasis>
. The
<taxonomicName id="4C0A4D14FFCEFFA7FE7CFDC9FD03C402" box="[411,726,546,571]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFCEFFA7FE7CFDC9FDFFC402" box="[411,554,546,571]" italics="true" pageId="40" pageNumber="41">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFCEFFA7FDDAFDC9FD19C402" box="[573,716,546,571]" italics="true" pageId="40" pageNumber="41">Cycadophila</emphasis>
)
</taxonomicName>
<taxonomicName id="4C0A4D14FFCEFFA7FD07FDC8FCF6C403" box="[736,803,547,570]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="species" species="papua">papua</taxonomicName>
species group is distinguished from other members of
<taxonomicName id="4C0A4D14FFCEFFA7FF70FDADFEF3C466" box="[151,294,582,607]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFCEFFA7FF70FDADFEF3C466" box="[151,294,582,607]" italics="true" pageId="40" pageNumber="41">Cycadophila</emphasis>
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by the body ventrally reddish-brown, appendages reddish-brown, dorsally body mostly black with reddish-brown margins on head and pronotum and central stripe on each elytron; head dorsal surface flattened in profile; supraocular stria present, long, full length of eye; maxillary palpi long, almost 2× longer than labial palpi; ventral interocular distance approximately 1/31/2 head width; gular-submental suture primarily flattened, medially with shallow U-shaped depression deeper on sided, with setose punctures; pronotal anterior angles rounded, surface near anterior angle convex; lateral margins evenly arched most of length, basally appearing evenly sinuate; pronotal lateral carina thin in lateral view, with single row of setose punctuation on marginal surface; prosternal process narrow (&lt;1/2 width of profemur), appearing rounded at apex, apex depressed, in lateral view convex; protibia weakly dilated distally, lacking fringe of stout setae along apical and lateral margins; coxal line behind mesocoxae with weakly impressed large punctures; male genitalia with tegmen moderately twisted basally, penile strut about 4.0 0× length of median lobe; and sexual dimorphism present with males having more slightly dilated pro- and mesotibial apices.
</paragraph>
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<subSubSection id="C310651CFFCEFFA7FF20FC1DFF35C242" pageId="40" pageNumber="41" type="discussion">
<paragraph id="8BB53697FFCEFFA7FF20FC1DFF35C242" blockId="40.[151,1437,546,1147]" pageId="40" pageNumber="41">
<emphasis id="B97EEA85FFCEFFA7FF20FC1DFEEEC236" bold="true" box="[199,315,1014,1039]" pageId="40" pageNumber="41">Remarks.</emphasis>
The elongate maxillary palpi and broad flattened body readily distinguish the species of this group from all other. Their widespread distribution in the
<collectingCountry id="F31D7607FFCEFFA7FD3FFBF1FC83C20A" box="[728,854,1050,1075]" name="Philippines" pageId="40" pageNumber="41">Philippines</collectingCountry>
and New
<collectingCountry id="F31D7607FFCEFFA7FC2DFBF1FBCEC20A" box="[970,1051,1050,1075]" name="Guinea-Bissau" pageId="40" pageNumber="41">Guinea</collectingCountry>
would indicate there are probably additional species in
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occurring with the widespread and isolated members of
<taxonomicName id="4C0A4D14FFCEFFA7FB6CFBD4FF0EC242" authority="Lindstrom et al. 2009" authorityName="Lindstrom et al." authorityYear="2009" class="Cycadopsida" family="Cycadaceae" genus="Cycas" kingdom="Plantae" order="Cycadales" pageId="40" pageNumber="41" phylum="Tracheophyta" rank="genus">
<emphasis id="B97EEA85FFCEFFA7FB6CFBD4FB1AC26F" box="[1163,1231,1087,1110]" italics="true" pageId="40" pageNumber="41">Cycas</emphasis>
(
<bibRefCitation id="EF9B4B66FFCEFFA7FB06FBD5FF01C242" author="Lindstrom" pageId="40" pageNumber="41" refString="Lindstrom, A. J., Hill, K. D. &amp; Stanberg, L. C. (2009) The genus Cycas (Cycadaceae) in Indonesia. Telopea, 12 (3), 385 - 418. https: // doi. org / 10.7751 / telopea 20095826" type="journal article" year="2009">
Lindstrom
<emphasis id="B97EEA85FFCEFFA7FA86FBD4FA49C26E" box="[1377,1436,1086,1111]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
2009
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)
</taxonomicName>
.
</paragraph>
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