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<document id="BB6EE64F7B8958542801019FF3EFC919" ID-CLB-Dataset="94528" ID-DOI="10.11646/zootaxa.5102.1.1" ID-GBIF-Dataset="ad30faa6-eab1-4dae-b686-9e5fee56f55c" ID-ISSN="1175-5326" ID-Zenodo-Dep="6245602" ID-ZooBank="BEA963B7-F7B9-458A-92EE-A4740AC390AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1645601333553" checkinUser="plazi" docAuthor="Ward, Philip S." docDate="2022" docId="03A33567FFA154201CD087C5FCEE3F59" docLanguage="en" docName="zootaxa.5102.1.1.pdf" docOrigin="Zootaxa 5102 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Tetraponera dispar Ward 2022, sp. nov." docType="treatment" docVersion="10" lastPageNumber="30" masterDocId="FF9A4D1FFFBD543D1C478336FFC13E45" masterDocTitle="The ant genus Tetraponera (Hymenoptera: Formicidae) in the Afrotropical region: taxonomic review and key to species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="29" updateTime="1699316131195" updateUser="ExternalLinkService">
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<mods:title id="7027945F46512E274BB0086F3649ACB3">The ant genus Tetraponera (Hymenoptera: Formicidae) in the Afrotropical region: taxonomic review and key to species</mods:title>
</mods:titleInfo>
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<mods:namePart id="A509C9BF652C31ED2A54840B3FECEE12">Ward, Philip S.</mods:namePart>
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<mods:typeOfResource id="6EAE36360E9752C3F2964E466D5F199A">text</mods:typeOfResource>
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<mods:title id="F7B98EF11AD501E703DBF7A11E955144">Zootaxa</mods:title>
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<mods:part id="F21E5D3BA9150DF3B2DE99873D40A56A">
<mods:date id="01F768BBAE1321A30E8430C535A89992">2022</mods:date>
<mods:detail id="98AF5DEECD47DA0825197569260203F4" type="pubDate">
<mods:number id="B07240FC789CD713138FC951703B1FFA">2022-02-23</mods:number>
</mods:detail>
<mods:detail id="8EDADFEA455D31EA65C2BB950AA2D63E" type="volume">
<mods:number id="1DE1339B8AA7C0E48C4EDBF882896988">5102</mods:number>
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<mods:number id="8491B0D43541150526312FAB947CF5A3">1</mods:number>
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<mods:classification id="80175BAE873FEAD3599855A49A1244F4">journal article</mods:classification>
<mods:identifier id="9985D247083C3822FC5873697EB2F148" type="CLB-Dataset">94528</mods:identifier>
<mods:identifier id="A98911BAAB556174247A0A8BF049180E" type="DOI">10.11646/zootaxa.5102.1.1</mods:identifier>
<mods:identifier id="C0EDEC43390A106F2EC5BA40F03FE69F" type="GBIF-Dataset">ad30faa6-eab1-4dae-b686-9e5fee56f55c</mods:identifier>
<mods:identifier id="D82DD85E9023B0FA55560399CD436A75" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="07F119A8DCCC362D10AEB206890010AF" type="Zenodo-Dep">6245602</mods:identifier>
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<treatment id="03A33567FFA154201CD087C5FCEE3F59" ID-DOI="http://doi.org/10.5281/zenodo.6822876" ID-GBIF-Taxon="197227894" ID-Zenodo-Dep="6822876" LSID="urn:lsid:plazi:treatment:03A33567FFA154201CD087C5FCEE3F59" httpUri="http://treatment.plazi.org/id/03A33567FFA154201CD087C5FCEE3F59" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<subSubSection id="C310D7FAFFA154211CD087C5FE233B48" box="[151,482,1266,1293]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BB58471FFA154211CD087C5FE233B48" blockId="28.[151,482,1266,1330]" box="[151,482,1266,1293]" pageId="28" pageNumber="29">
<heading id="D0FD331DFFA154211CD087C5FE233B48" bold="true" box="[151,482,1266,1293]" fontSize="11" level="2" pageId="28" pageNumber="29" reason="4">
<emphasis id="B97E5863FFA154211CD087C5FE233B48" bold="true" box="[151,482,1266,1293]" pageId="28" pageNumber="29">
<taxonomicName id="4C0AFFF2FFA154211CD087C5FEBC3B48" authority="Ward, 2022" authorityName="Ward" authorityYear="2022" box="[151,381,1266,1293]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="B97E5863FFA154211CD087C5FEBC3B48" bold="true" box="[151,381,1266,1293]" italics="true" pageId="28" pageNumber="29">Tetraponera dispar</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24DE518FFA154211DC387C5FE233B48" box="[388,482,1267,1293]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA154211CD08621FED83B77" box="[151,281,1303,1330]" pageId="28" pageNumber="29" type="description">
<paragraph id="8BB58471FFA154211CD08621FED83B77" blockId="28.[151,482,1266,1330]" box="[151,281,1303,1330]" pageId="28" pageNumber="29">
(
<figureCitation id="133198F4FFA154211CD88621FF293B74" box="[159,232,1303,1329]" captionStart="FIGURES 7–9" captionStartId="14.[151,264,1692,1716]" captionTargetBox="[152,1437,181,1666]" captionTargetId="figure-29@14.[151,1437,181,1667]" captionTargetPageId="14" captionText="FIGURES 7–9. Tetraponera allaborans group, full-face (dorsal) view of head (a) and lateral profile of body (b). 7, T. dispar, worker, holotype, Republic of Congo (CASENT0888518); 8, T. emeryi, worker, South Africa (CASENT0220743); 9, T. exactor, alate queen, holotype, South Africa (CASENT0794350). Images from AntWeb (www.antweb.org); photographers Peter Hawkes (7), Matthew Prebus (8, 9)." figureDoi="http://doi.org/10.5281/zenodo.6245620" httpUri="https://zenodo.org/record/6245620/files/figure.png" pageId="28" pageNumber="29">Figs 7</figureCitation>
,
<figureCitation id="133198F4FFA154211CB38621FED03B77" box="[244,273,1303,1330]" captionStart="FIGURES 35–40" captionStartId="23.[151,264,1943,1967]" captionTargetBox="[156,1433,182,1919]" captionTargetId="figure-17@23.[155,1433,181,1919]" captionTargetPageId="23" captionText="FIGURES 35–40. Distribution maps for Afrotropical Tetraponera. 35, T. clypeata (circles), T. tessmanni (triangles); 36, T. emeryi (circles), T. dispar (triangle); 37, T. exactor (circle), T. furtiva (triangle), T. liengmei (squares); 38, T. continua (circles), T. cortina (triangles), T. gerdae (squares); 39, T. pedana (circles), T. penzigi (triangles); 40, T. pumila (circles)." figureDoi="http://doi.org/10.5281/zenodo.6245653" httpUri="https://zenodo.org/record/6245653/files/figure.png" pageId="28" pageNumber="29">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA154211CD08669FBE3396C" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BB58471FFA154211CD08669FC063B84" blockId="28.[151,1437,1374,2013]" pageId="28" pageNumber="29">
<emphasis id="B97E5863FFA154211CD08669FE843B3D" bold="true" box="[151,325,1374,1401]" pageId="28" pageNumber="29">Type material.</emphasis>
<materialsCitation id="3B628E2CFFA154211D0B8669FA583BD8" ID-GBIF-Occurrence="3861810359" collectingDate="2013-11-16" collectionCode="SAMC" collectorName="L. Niemand &amp; C. Ngoulou " country="Republic of the Congo" county="Afog" elevation="27803" latitude="-2.47106" location="Niari" longLatPrecision="1" longitude="12.96037" municipality="Tsingidi" pageId="28" pageNumber="29" specimenCode="CASENT0888518" specimenCount="1" typeStatus="holotype">
<emphasis id="B97E5863FFA154211D0B8669FDC83B3C" box="[332,521,1375,1401]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="54B13AD3FFA154211D0B8669FE723B3C" box="[332,435,1375,1401]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
worker
</emphasis>
:
<collectingCountry id="F31DC4E1FFA154211E5E8669FD303B3C" box="[537,753,1375,1401]" name="Republic of the Congo" pageId="28" pageNumber="29">Republic of Congo</collectingCountry>
,
<location id="8ED5D2AAFFA154211EBA8669FCF83B3C" LSID="urn:lsid:plazi:treatment:03A33567FFA154201CD087C5FCEE3F59:8ED5D2AAFFA154211EBA8669FCF83B3C" box="[765,825,1375,1401]" country="Republic of the Congo" county="Afog" latitude="-2.47106" longLatPrecision="1" longitude="12.96037" municipality="Tsingidi" name="Niari" pageId="28" pageNumber="29">Niari</location>
,
<collectingMunicipality id="6BD11E0BFFA154211F038669FC633B3C" box="[836,930,1375,1401]" pageId="28" pageNumber="29">Tsingidi</collectingMunicipality>
,
<collectingCounty id="62D4FCFDFFA154211FEB8669FC283B3C" box="[940,1001,1375,1401]" pageId="28" pageNumber="29">Afog</collectingCounty>
<quantity id="4CF22994FFA154211FAE8669FB9B3B3C" box="[1001,1114,1375,1401]" metricMagnitude="4" metricUnit="m" metricValue="2.7803" pageId="28" pageNumber="29" unit="m" value="27803.0">
<elevation id="00276342FFA154211FAE8669FB9B3B3C" box="[1001,1114,1375,1401]" metricMagnitude="4" metricUnit="m" metricValue="2.7803" pageId="28" pageNumber="29" unit="m" value="27803.0">27, 803 m</elevation>
</quantity>
,
<geoCoordinate id="EE3EE2B6FFA1542118218669FB0B3B3C" box="[1126,1226,1375,1401]" degrees="2.47106" direction="south" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="-2.47106">-2.47106</geoCoordinate>
<geoCoordinate id="EE3EE2B6FFA1542118968669FAFC3B3C" box="[1233,1341,1375,1401]" degrees="12.96037" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="12.96037">12.96037</geoCoordinate>
,
<date id="FFB4A2B1FFA15421190E8669FF133BD8" pageId="28" pageNumber="29" value="2013-11-16">
<collectingDate id="EFF05B59FFA15421190E8669FF133BD8" pageId="28" pageNumber="29" value="2013-11-16">16 Nov 2013</collectingDate>
</date>
, tall mature forest, canopy fogging,
<collectorName id="26FFE1A7FFA154211E1B86B5FD203BD8" box="[604,737,1411,1437]" pageId="28" pageNumber="29">L. Niemand</collectorName>
&amp;
<collectorName id="26FFE1A7FFA154211F4786B5FC403BD8" box="[768,897,1411,1437]" pageId="28" pageNumber="29">C. Ngoulou</collectorName>
LN-RC1-
<collectorName id="26FFE1A7FFA154211FA886B5FBED3BD8" box="[1007,1068,1411,1437]" pageId="28" pageNumber="29">Afog</collectorName>
27 (
<specimenCode id="DBAC2C0AFFA15421181386B4FAE63BD8" box="[1108,1319,1410,1437]" collectionCode="CASENT" pageId="28" pageNumber="29">CASENT0888518</specimenCode>
) (
<collectionCode id="ED1B1CB4FFA15421197C86B5FA503BD8" box="[1339,1425,1411,1437]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="28" pageNumber="29" type="Museum">SAMC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B628E2CFFA154211CD08691FC023B84" ID-GBIF-Occurrence="3861810328" box="[151,963,1446,1473]" collectingDate="2013-11-16" collectionCode="CASC" collectorName="L. Niemand &amp; C. Ngoulou" country="Republic of the Congo" county="Afog" elevation="27803" latitude="-2.47106" location="Niari" longLatPrecision="1" longitude="12.96037" municipality="Tsingidi" pageId="28" pageNumber="29" specimenCode="CASENT0888519" specimenCount="1" specimenCount-worker="1" typeStatus="paratype">
<emphasis id="B97E5863FFA154211CD08691FF3F3B85" box="[151,254,1447,1472]" italics="true" pageId="28" pageNumber="29">
<typeStatus id="54B13AD3FFA154211CD08691FF3F3B85" box="[151,254,1447,1472]" pageId="28" pageNumber="29" type="paratype">Paratype</typeStatus>
</emphasis>
:
<specimenCount id="9D0C4FF8FFA154211D4A8691FEB23B84" box="[269,371,1447,1473]" pageId="28" pageNumber="29" type="worker">1 worker</specimenCount>
, same data as holotype (
<specimenCode id="DBAC2C0AFFA154211EC48690FC973B84" box="[643,854,1446,1473]" collectionCode="CASENT" pageId="28" pageNumber="29">CASENT0888519</specimenCode>
) (
<collectionCode id="ED1B1CB4FFA154211F2B8691FC7A3B84" box="[876,955,1447,1473]" pageId="28" pageNumber="29">CASC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BB58471FFA154211C8086FDFEA33868" blockId="28.[151,1437,1374,2013]" pageId="28" pageNumber="29">
<emphasis id="B97E5863FFA154211C8086FDFE163BA0" bold="true" box="[199,471,1483,1509]" pageId="28" pageNumber="29">Worker measurements</emphasis>
(n = 2). HW 0.51–0.52, HL 0.66–0.67, LHT 0.43–0.44, CI 0.77–0.78, FCI 0.10, REL 0.35, REL2 0.45, SI 0.62–0.63, SI3 1.39–1.40, FI 0.38–0.39, PLI 0.59–0.60, PWI 0.44–0.48, LHT/HW 0.83–0.86, CSC 2, MSC 1–2.
</paragraph>
<paragraph id="8BB58471FFA154211C808501FBE3396C" blockId="28.[151,1437,1374,2013]" pageId="28" pageNumber="29">
<emphasis id="B97E5863FFA154211C808501FE5D3815" bold="true" box="[199,412,1590,1617]" pageId="28" pageNumber="29">Worker diagnosis</emphasis>
. A member of the
<taxonomicName id="4C0AFFF2FFA154211E3D8501FD363814" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[634,759,1591,1617]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA154211E3D8501FD363814" box="[634,759,1591,1617]" italics="true" pageId="28" pageNumber="29">T. clypeata</emphasis>
</taxonomicName>
complex (see characterization under
<taxonomicName id="4C0AFFF2FFA1542118EF8501FAE43814" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[1192,1317,1591,1617]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA1542118EF8501FAE43814" box="[1192,1317,1591,1617]" italics="true" pageId="28" pageNumber="29">T. clypeata</emphasis>
</taxonomicName>
), with the following more specific features: very small (HL &lt;0.70), with relatively broad head (CI 0.77–0.78) and large eyes (
<collectionCode id="ED1B1CB4FFA154211CE78549FF1438DC" box="[160,213,1663,1689]" pageId="28" pageNumber="29">REL</collectionCode>
0.35, EL/LHT 0.52–0.54); mesopropodeal impression simple, lacking longitudinal rugulae; profemur rather slender (FI 0.38–0.39); petiole relatively short (for this complex), such that PLI 0.59–0.60, PL/HW 0.71, and PL/SL 1.13–1.14. Integument predominantly smooth and shiny, with scattered fine punctures. Standing pilosity sparse, present on frontal carinae, vertex (one supraocular pair), and pronotum (one anterolateral pair), absent from remainder of mesosoma and from the petiole and postpetiole. Body light orange-brown.
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA154211C808404FA453998" pageId="28" pageNumber="29" type="discussion">
<paragraph id="8BB58471FFA154211C808404FA453998" blockId="28.[151,1437,1374,2013]" pageId="28" pageNumber="29">
<emphasis id="B97E5863FFA154211C808404FE893909" bold="true" box="[199,328,1842,1868]" pageId="28" pageNumber="29">Comments</emphasis>
.
<taxonomicName id="4C0AFFF2FFA154211D1F8405FDF23908" authorityName="Ward" authorityYear="2022" box="[344,563,1843,1869]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E5863FFA154211D1F8405FDF23908" box="[344,563,1843,1869]" italics="true" pageId="28" pageNumber="29">Tetraponera dispar</emphasis>
</taxonomicName>
is a diminutive, orange-brown species in the
<taxonomicName id="4C0AFFF2FFA1542118068405FB7F3908" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[1089,1214,1843,1869]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA1542118068405FB7F3908" box="[1089,1214,1843,1869]" italics="true" pageId="28" pageNumber="29">T. clypeata</emphasis>
</taxonomicName>
complex. It can be distinguished from other species in that complex by its small size, broad head, and large eyes. In addition, the petiole is relatively short: PL/HW 0.71 and PL/SL 1.13–1.14, compared with PL/HW 0.72–0.94 and PL/SL
<quantity id="4CF22994FFA154211949844DFA5D39D0" box="[1294,1436,1915,1941]" metricMagnitude="-2" metricUnit="m" metricValue="3.3528" metricValueMax="3.6322" metricValueMin="3.0734" pageId="28" pageNumber="29" unit="in" value="1.3199999999999998" valueMax="1.43" valueMin="1.21">1.21–1.43 in</quantity>
other species (these measurements refer to workers only). Finally, the worker of
<taxonomicName id="4C0AFFF2FFA15421186484A9FB4639FC" authorityName="Ward" authorityYear="2022" box="[1059,1159,1951,1977]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E5863FFA15421186484A9FB4639FC" box="[1059,1159,1951,1977]" italics="true" pageId="28" pageNumber="29">T. dispar</emphasis>
</taxonomicName>
has an anterolateral pair of setae on the pronotum, whereas these setae are lacking in workers of other species in the
<taxonomicName id="4C0AFFF2FFA1542118DC84F5FAD73998" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[1179,1302,1987,2013]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA1542118DC84F5FAD73998" box="[1179,1302,1987,2013]" italics="true" pageId="28" pageNumber="29">T. clypeata</emphasis>
</taxonomicName>
complex.
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA054201C8083A0FA423EBD" pageId="29" pageNumber="30" type="distribution">
<paragraph id="8BB58471FFA054201C8083A0FA423EBD" blockId="29.[151,1437,150,285]" pageId="29" pageNumber="30">
<emphasis id="B97E5863FFA054201C8083A0FE2E3EF5" bold="true" box="[199,495,150,176]" pageId="29" pageNumber="30">Distribution and biology</emphasis>
.
<taxonomicName id="4C0AFFF2FFA054201DB883A1FD1B3EF5" authorityName="Ward" authorityYear="2022" box="[511,730,150,176]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E5863FFA054201DB883A1FD1B3EF5" box="[511,730,150,176]" italics="true" pageId="29" pageNumber="30">Tetraponera dispar</emphasis>
</taxonomicName>
is known only from
<specimenCount id="9D0C4FF8FFA054201F8B83A1FB983EF5" box="[972,1113,150,177]" pageId="29" pageNumber="30" type="worker">two workers</specimenCount>
taken by canopy fogging in primary rainforest in the
<collectingCountry id="F31DC4E1FFA054201DEC838CFD403E91" box="[427,641,186,212]" name="Republic of the Congo" pageId="29" pageNumber="30">Republic of Congo</collectingCountry>
. The habitat is illustrated in
<bibRefCitation id="EF9BF980FFA054201FFF838CFB323E91" author="Janak, J. &amp; Bordoni, A." box="[952,1267,186,213]" pageId="29" pageNumber="30" pagination="1 - 94" refId="ref66081" refString="Janak, J. &amp; Bordoni, A. (2015) Revision of the genus Thyreocephalus and description of Afrus gen. nov. of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae). Zootaxa, 4038 (1), 1 - 94. https: // doi. org / 10.11646 / zootaxa. 4038.1.1" type="journal article" year="2015">Janák &amp; Bordoni (2015: 91)</bibRefCitation>
. The four other species in the
<taxonomicName id="4C0AFFF2FFA054201D7283E9FE713EBD" baseAuthorityName="Emery" baseAuthorityYear="1886" box="[309,432,222,248]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="clypeata">
<emphasis id="B97E5863FFA054201D7283E9FE713EBD" box="[309,432,222,248]" italics="true" pageId="29" pageNumber="30">T. clypeata</emphasis>
</taxonomicName>
complex are confined to southern Africa, so
<taxonomicName id="4C0AFFF2FFA054201FEF83E9FBCA3EBD" authorityName="Ward" authorityYear="2022" box="[936,1035,222,248]" class="Insecta" family="Formicidae" genus="Tetraponera" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B97E5863FFA054201FEF83E9FBCA3EBD" box="[936,1035,222,248]" italics="true" pageId="29" pageNumber="30">T. dispar</emphasis>
</taxonomicName>
is a striking geographical outlier.
</paragraph>
</subSubSection>
<subSubSection id="C310D7FAFFA054201C808234FCEE3F59" box="[199,815,258,285]" pageId="29" pageNumber="30" type="materials_examined">
<paragraph id="8BB58471FFA054201C808234FCEE3F59" blockId="29.[151,1437,150,285]" box="[199,815,258,285]" pageId="29" pageNumber="30">
<emphasis id="B97E5863FFA054201C808234FE703F59" bold="true" box="[199,433,258,284]" pageId="29" pageNumber="30">Material examined.</emphasis>
Known only from the
<typeStatus id="54B13AD3FFA054201EF78235FD213F58" box="[688,736,259,285]" pageId="29" pageNumber="30">type</typeStatus>
series.
</paragraph>
</subSubSection>
</treatment>
</document>
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