140 lines
18 KiB
XML
140 lines
18 KiB
XML
<document id="A35FEF6857F2D6F2EBD306617B3D728B" ID-ISSN="11755334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFFA3AF02D1DBD83EFC53163A" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Feeridium Vaz-De-Mello 2008, new genus" docType="treatment" docVersion="2" lastPageNumber="25" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="24" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="570B733C4CF4CD9C97BC00EE6F99552B">
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<mods:title id="83AF1BE29650CA48913AC292F3A2F2DE">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
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<mods:name id="3A07045836C3C56C00D507FE5D30C3E5" type="personal">
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<mods:roleTerm id="44B6EFA5E7A19F72CDE860A13B41ED69">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="76B45750149A05ED86792192429E2F5C">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="AE1AFF986BD92CCB5D8B66BCCAAFB2F6">text</mods:typeOfResource>
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<mods:titleInfo id="3995D471BFC2538C175065A3DA7AA0D1">
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<mods:title id="0B3E45A7D71C9D3BD2FD3F13BCB18A94">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="15E2FC8AFD47D8596629A8570C1DD7C7">
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<mods:date id="F2B069196E9608FE2D758FC504B673D6">2008</mods:date>
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<mods:detail id="487813C9DDA955FF2F3D2B93A113CED0" type="pubDate">
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<mods:number id="4E3858F457A64FED7E35B33BF203F4FE">2008-12-05</mods:number>
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</mods:detail>
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<mods:detail id="089E7DC6F1B68166E58B4859C8AC1948" type="volume">
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<mods:number id="283632EB7DED222349E7C6BA35ED6B22">1955</mods:number>
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<mods:extent id="E23DCF51CF93CBF7C804A98E752D17DD" unit="page">
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<mods:start id="88A7AD3636DCE0364883629A45FBDD23">1</mods:start>
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<mods:end id="0C94DF26FF1304D1F46E0944300E53FE">75</mods:end>
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</mods:part>
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<mods:classification id="018706F0D59B35838CCAC82E9FA403B9">journal article</mods:classification>
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<mods:identifier id="C30FE1E85676DBCAD89FB369E2A607F6" type="ISSN">11755334</mods:identifier>
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</mods:mods>
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<treatment id="03D287CEFFA3AF02D1DBD83EFC53163A" LSID="urn:lsid:plazi:treatment:03D287CEFFA3AF02D1DBD83EFC53163A" httpUri="http://treatment.plazi.org/id/03D287CEFFA3AF02D1DBD83EFC53163A" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<subSubSection id="C3616553FFA3AF0DD1DBD83EFE1D143C" box="[151,443,1866,1892]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BC436D8FFA3AF0DD1DBD83EFE1D143C" blockId="23.[151,443,1866,1892]" box="[151,443,1866,1892]" pageId="23" pageNumber="24">
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<heading id="D08C81B4FFA3AF0DD1DBD83EFE1D143C" bold="true" box="[151,443,1866,1892]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B90FEACAFFA3AF0DD1DBD83EFE1D143C" bold="true" box="[151,443,1866,1892]" pageId="23" pageNumber="24">
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5.
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<taxonomicName id="4C7B4D5BFFA3AF0DD1F8D83EFE95143C" ID-CoL="4JP4" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[180,307,1866,1892]" class="Insecta" family="Scarabaeidae" genus="Feeridium" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90FEACAFFA3AF0DD1F8D83EFE95143C" bold="true" box="[180,307,1866,1892]" italics="true" pageId="23" pageNumber="24">Feeridium</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23C57B1FFA3AF0DD076D83EFE1D143C" box="[314,443,1866,1892]" pageId="23" pageNumber="24" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFA3AF0DD1DBD8FEFB9A14AC" pageId="23" pageNumber="24" type="diagnosis">
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<paragraph id="8BC436D8FFA3AF0DD1DBD8FEFB9A14AC" blockId="23.[151,1436,1930,2036]" pageId="23" pageNumber="24">
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<emphasis id="B90FEACAFFA3AF0DD1DBD8FEFEBF14FC" bold="true" box="[151,281,1930,1956]" pageId="23" pageNumber="24">Diagnosis:</emphasis>
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Size large (
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<quantity id="4C839B3DFFA3AF0DD0EBD8FEFD9D14FC" box="[423,571,1930,1956]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="23" pageNumber="24" unit="mm" value="4.85" valueMax="5.2" valueMin="4.5">4.5–5.2 mm</quantity>
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), very elongated (
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<figureCitation id="13402A5DFFA3AF0DD25CD8FEFCCF14FC" box="[784,873,1930,1956]" captionStart="FIGURES 15–22" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 15–22. Dorsal views. 15–16: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 21–22: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="23" pageNumber="24">Fig. 20</figureCitation>
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). Mesofemora and metafemora very strong and rounded (
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<figureCitation id="13402A5DFFA3AF0DD045D8C6FEC71494" box="[265,353,1970,1996]" captionStart="FIGURES 47–54" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 47–54. 47–48: Degallieridium lilliputanum: 47. head; 48. parameres; 49–51: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 52–54. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="23" pageNumber="24">Fig. 54</figureCitation>
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), eyes extremely large dorsally, interocular width narrower than twice eye width (
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<figureCitation id="13402A5DFFA3AF0DD479D8C6FA281494" box="[1333,1422,1970,1996]" captionStart="FIGURES 47–54" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 47–54. 47–48: Degallieridium lilliputanum: 47. head; 48. parameres; 49–51: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 52–54. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="23" pageNumber="24">Fig. 52</figureCitation>
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). Eytral interstriae sparsely punctate, with uniseriate setae only at apex and sides.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFACAF02D18ADFE3FC4A10F2" pageId="24" pageNumber="25" type="description">
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<paragraph id="8BC436D8FFACAF02D18ADFE3FD4B1002" blockId="24.[151,1436,151,1378]" pageId="24" pageNumber="25">
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<emphasis id="B90FEACAFFACAF02D18ADFE3FEC613E9" bold="true" box="[198,352,151,177]" pageId="24" pageNumber="25">Description:</emphasis>
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Length
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<quantity id="4C839B3DFFACAF02D086DFECFDF813EA" box="[458,606,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="4.85" metricValueMax="5.2" metricValueMin="4.5" pageId="24" pageNumber="25" unit="mm" value="4.85" valueMax="5.2" valueMin="4.5">4.5–5.2 mm</quantity>
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, body very elongated, with subparallel sides (
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<figureCitation id="13402A5DFFACAF02D5C2DFECFB4C13EA" box="[1166,1258,152,178]" captionStart="FIGURES 15–22" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 15–22. Dorsal views. 15–16: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 21–22: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="24" pageNumber="25">Fig. 20</figureCitation>
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); color brown, opaque, without metallic sheen. Clypeus with two strong teeth, separated by narrow U-shaped emargination, clypeus laterally and gena continuously rounded. Clypeofrontal and clypeogenal sutures not discernible. Eyes, in dorsal view, enormous, as long as wide; interocular width less than 1.5 times eye width (
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<figureCitation id="13402A5DFFACAF02D5F0DE64FAB21272" box="[1212,1300,272,298]" captionStart="FIGURES 47–54" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 47–54. 47–48: Degallieridium lilliputanum: 47. head; 48. parameres; 49–51: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 52–54. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="24" pageNumber="25">Fig. 52</figureCitation>
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). Pronotum lacking anterior and posterior beads, lateral pronotal callosity rounded and darkened; disc covered by simple strong punctures, glabrous; separated of hypomeron by row of punctures, without carina. Hypomeron without lateral longitudinal carina; mesoepimeron anterior transverse carina strong and distinct; metasternal disc with minute impressed punctures. Elytral interstriae flat, with unorganized punctures on disc, sparse uniseriate setae present only apically and laterally; striae slightly deeper posteriorly, not widened. Pseudoepipleura gradually narrowed posteriorly, with slight longitudinal fold near metacoxa, completely glabrous. Protibia with three very strong teeth, distributed along apical three-fifths, basally not denticulate, lacking ventral scale-like setae. Mesofemora wider than third of their length, metafemora about as wide as half of their length (
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<figureCitation id="13402A5DFFACAF02D47ADD24FA2B1132" box="[1334,1421,592,618]" captionStart="FIGURES 47–54" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 47–54. 47–48: Degallieridium lilliputanum: 47. head; 48. parameres; 49–51: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 52–54. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="24" pageNumber="25">Fig. 54</figureCitation>
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); mesotibiae apically wider than one of their length, metatibiae apically quite as wide as half of their length (
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<figureCitation id="13402A5DFFACAF02D1D3DDD4FF5F11E2" box="[159,249,672,698]" captionStart="FIGURES 47–54" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 47–54. 47–48: Degallieridium lilliputanum: 47. head; 48. parameres; 49–51: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 52–54. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="24" pageNumber="25">Fig. 54</figureCitation>
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). Metatarsi with basal tarsomere as long as 1.5 times the following tarsomere. Parameres as long as two-thirds of phallobase, flattened and truncate, apically divergent (
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<figureCitation id="13402A5DFFACAF02D2F2DDBCFBB111BA" box="[958,1047,712,738]" captionStart="FIGURES 47–54" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 47–54. 47–48: Degallieridium lilliputanum: 47. head; 48. parameres; 49–51: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 52–54. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="24" pageNumber="25">Fig. 53</figureCitation>
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). Internal sac with pseudoflagellum relativelly short and bent at mid-length, rounded by flat large accessory lamella. Coxites triangular, very sclerotized, and pointed. Spermateca C-shaped, bulbous basally (narrower at duct insertion), drastically narrowed apically, with duct short and not sclerotized.
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</paragraph>
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<paragraph id="8BC436D8FFACAF02D18ADC13FC4A10F2" blockId="24.[151,1436,151,1378]" pageId="24" pageNumber="25">
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<emphasis id="B90FEACAFFACAF02D18ADC13FE6510D9" bold="true" box="[198,451,871,897]" pageId="24" pageNumber="25">Sexual dimorphism:</emphasis>
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Males have central portion of last abdominal sternite flat to slightly concave, pygidium more elongated, and a protibial apicomesal tooth, ventrally directed.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFACAF02D18ADCC3FC30108A" box="[198,918,951,978]" pageId="24" pageNumber="25" type="type_taxon">
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<paragraph id="8BC436D8FFACAF02D18ADCC3FC30108A" blockId="24.[151,1436,151,1378]" box="[198,918,951,978]" pageId="24" pageNumber="25">
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<emphasis id="B90FEACAFFACAF02D18ADCC3FECC1089" bold="true" box="[198,362,951,977]" pageId="24" pageNumber="25">
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<typeStatus id="54C0887AFFACAF02D18ADCC3FEA51089" box="[198,259,951,977]" pageId="24" pageNumber="25">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C7B4D5BFFACAF02D03EDCCCFDC01089" ID-CoL="3DXC7" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[370,614,952,977]" class="Insecta" family="Scarabaeidae" genus="Feeridium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="woodruffi" status="sp. nov.">
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<emphasis id="B90FEACAFFACAF02D03EDCCCFDC01089" box="[370,614,952,977]" italics="true" pageId="24" pageNumber="25">Feeridium woodruffi</emphasis>
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</taxonomicName>
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<emphasis id="B90FEACAFFACAF02D321DCC3FD5B1089" bold="true" box="[621,765,951,977]" pageId="24" pageNumber="25">
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<taxonomicNameLabel id="A23C57B1FFACAF02D321DCC3FD5B1089" box="[621,765,951,977]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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(monotypy).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFACAF02D18ADCABFBB01712" pageId="24" pageNumber="25" type="etymology">
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<paragraph id="8BC436D8FFACAF02D18ADCABFEBB177A" blockId="24.[151,1436,151,1378]" pageId="24" pageNumber="25">
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<emphasis id="B90FEACAFFACAF02D18ADCABFEF110A1" bold="true" box="[198,343,991,1017]" pageId="24" pageNumber="25">Etymology:</emphasis>
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After François Feer, who collected and sent me the first specimens I saw of this genus. Gen- der neutral.
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</paragraph>
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<paragraph id="8BC436D8FFACAF02D18ADB5BFBB01712" blockId="24.[151,1436,151,1378]" box="[198,1046,1071,1098]" pageId="24" pageNumber="25">
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<emphasis id="B90FEACAFFACAF02D18ADB5BFEC91711" bold="true" box="[198,367,1071,1097]" pageId="24" pageNumber="25">Composition:</emphasis>
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The new genus currently includes solely the new species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFACAF02D18ADB23FDC317C2" pageId="24" pageNumber="25" type="distribution">
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<paragraph id="8BC436D8FFACAF02D18ADB23FDC317C2" blockId="24.[151,1436,151,1378]" pageId="24" pageNumber="25">
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<emphasis id="B90FEACAFFACAF02D18ADB23FECF1729" bold="true" box="[198,361,1111,1137]" pageId="24" pageNumber="25">Distribution:</emphasis>
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French
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<collectingCountry id="F36C7648FFACAF02D080DB2CFD81172A" box="[460,551,1112,1138]" name="Guyana" pageId="24" pageNumber="25">Guyana</collectingCountry>
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and Amazonian
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<collectingCountry id="F36C7648FFACAF02D3BFDB2CFC98172A" box="[755,830,1112,1138]" name="Brazil" pageId="24" pageNumber="25">Brazil</collectingCountry>
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. Known from Humid Guiana,
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<collectingRegion id="49BFF83AFFACAF02D5E7DB2CFAB6172A" box="[1195,1296,1112,1138]" country="Brazil" name="Roraima" pageId="24" pageNumber="25">Roraima</collectingRegion>
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and Varzea provinces of the Amazonian subregion.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616553FFACAF02D18ADBD3FC53163A" pageId="24" pageNumber="25" type="discussion">
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<paragraph id="8BC436D8FFACAF02D18ADBD3FC53163A" blockId="24.[151,1436,151,1378]" pageId="24" pageNumber="25">
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<emphasis id="B90FEACAFFACAF02D18ADBD3FEE71799" bold="true" box="[198,321,1191,1217]" pageId="24" pageNumber="25">Remarks:</emphasis>
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The condition of the parameres and pseudepipleura relates this genus to
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<taxonomicName id="4C7B4D5BFFACAF02D5DBDBDCFAEC1799" authorityName="Vaz-De-Mello" authorityYear="2008" box="[1175,1354,1192,1217]" class="Insecta" family="Scarabaeidae" genus="Degallieridium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFFACAF02D5DBDBDCFAEC1799" box="[1175,1354,1192,1217]" italics="true" pageId="24" pageNumber="25">Degallieridium</emphasis>
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</taxonomicName>
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. However, it superficially resembles both
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<taxonomicName id="4C7B4D5BFFACAF02D307DBA4FD7917B1" authorityName="Vaz-De-Mello" authorityYear="2008" box="[587,735,1232,1257]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFFACAF02D307DBA4FD7917B1" box="[587,735,1232,1257]" italics="true" pageId="24" pageNumber="25">Genieridium</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7B4D5BFFACAF02D251DBA4FC3917B1" authorityName="Vaz-De-Mello" authorityYear="2008" box="[797,927,1232,1257]" class="Insecta" family="Scarabaeidae" genus="Nunoidium" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEACAFFACAF02D251DBA4FC3917B1" box="[797,927,1232,1257]" italics="true" pageId="24" pageNumber="25">Nunoidium</emphasis>
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</taxonomicName>
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, from which it can be immediately distinguished by the condition of pseudepipleuron, head, and elytra. Synapomorphies defining this genus include lack of row-organized elytral disc punctation, absence of pronotal-hypomeral carina, lack of sexual differences in protarsi, size of eyes and psammophiliform development of legs.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |