367 lines
52 KiB
XML
367 lines
52 KiB
XML
<document id="4F7726637484810B9E59E064D92A21B5" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E31E8C23A2EDFC43FAE2FA61" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum spinatum Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="224" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="221" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:titleInfo id="E216D7DAEED38FC8A4F41949F3818153">
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<mods:title id="5F638E39404E4FD27EC1280C02438ECB">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="2AEB44F5787CF0D948BDC32B3284069B" type="personal">
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<mods:roleTerm id="E5543A03B14CF7D46299C23DFA0F38D2">Author</mods:roleTerm>
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<mods:namePart id="42426DC884680E4A89DAEC1A7F5577AE">Wesener, Thomas</mods:namePart>
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<mods:title id="C05316516FA6AD44A93D369F55BDC4C6">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F8F2137575327D046E46FFD3800BF261">
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<mods:date id="29FF6C9C1A22D1E09ED13F9758104A4E">2019</mods:date>
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<mods:number id="1EE9EF086A32B46021C4433509DE2FCF">2019-03-01</mods:number>
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<mods:number id="CFB8700379D4AD6983BFDF4197027174">4563</mods:number>
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<mods:detail id="E7F8BB4A9B89A0A2030D96A724ED05C0" type="issue">
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<mods:identifier id="8B03A32FDAE8AF95747BF016E03EDFC2" type="CLB-Dataset">28149</mods:identifier>
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<mods:identifier id="A9DA5A2BF64B2EB9825EC9BF602A42CA" type="DOI">10.11646/zootaxa.4563.2.1</mods:identifier>
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<mods:identifier id="8287D2CDCDC4C7C129E15E98722CBA61" type="GBIF-Dataset">e6037c5d-a96f-41e5-8e70-71f2659659e2</mods:identifier>
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<mods:identifier id="5C2EDA49E1E6A7699042B3A54B70D883" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="78AB017B5E3089974FCEC285108EAA8D" type="Zenodo-Dep">2601072</mods:identifier>
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<mods:identifier id="150BC52302757001929D92FF1ED5999B" type="ZooBank">CF79B01B-8B5F-4B3A-B642-2CADE4B339AF</mods:identifier>
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</mods:mods>
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<treatment id="03EB5506E31E8C23A2EDFC43FAE2FA61" ID-DOI="http://doi.org/10.5281/zenodo.5934528" ID-GBIF-Taxon="154474548" ID-Zenodo-Dep="5934528" LSID="urn:lsid:plazi:treatment:03EB5506E31E8C23A2EDFC43FAE2FA61" httpUri="http://treatment.plazi.org/id/03EB5506E31E8C23A2EDFC43FAE2FA61" lastPageId="23" lastPageNumber="224" pageId="20" pageNumber="221">
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<subSubSection id="C358B79BE31E8C20A2EDFC43FDBCFC5A" box="[151,597,908,934]" pageId="20" pageNumber="221" type="nomenclature">
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<paragraph id="8BFDE410E31E8C20A2EDFC43FDBCFC5A" blockId="20.[151,597,908,967]" box="[151,597,908,934]" pageId="20" pageNumber="221">
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<heading id="D0B5537CE31E8C20A2EDFC43FDBCFC5A" bold="true" box="[151,597,908,934]" fontSize="11" level="1" pageId="20" pageNumber="221" reason="1">
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<emphasis id="B9363802E31E8C20A2EDFC43FDBCFC5A" bold="true" box="[151,597,908,934]" pageId="20" pageNumber="221">
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<taxonomicName id="4C429F93E31E8C20A2EDFC43FE56FC5A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,447,908,934]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
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<emphasis id="B9363802E31E8C20A2EDFC43FE56FC5A" bold="true" box="[151,447,908,934]" italics="true" pageId="20" pageNumber="221">Sphaerobelum spinatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E31E8C20A3BFFC43FDBCFC5A" box="[453,597,908,934]" pageId="20" pageNumber="221" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE31E8C20A2EDFC61FE28FC3B" box="[151,449,942,967]" pageId="20" pageNumber="221" type="description">
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<paragraph id="8BFDE410E31E8C20A2EDFC61FE28FC3B" blockId="20.[151,597,908,967]" box="[151,449,942,967]" pageId="20" pageNumber="221">
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<figureCitation id="1379F895E31E8C20A2EDFC61FEFFFC3B" box="[151,278,942,967]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="20" pageNumber="221">Figures 6D</figureCitation>
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,
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<figureCitation id="1379F895E31E8C20A358FC60FEADFC3B" box="[290,324,943,967]" captionStart="FIGURE 9" captionStartId="13.[151,250,1423,1445]" captionTargetBox="[152,1434,1021,1400]" captionTargetId="figure@13.[152,1435,1021,1402]" captionTargetPageId="13" captionText="FIGURE 9. Endoterga of the body ring 9 of different Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum spinatum n. sp.; D: Sphaerobelum splendidum n. sp. Abbreviations: cp = cuticular patterns; iA = inner area." httpUri="https://zenodo.org/record/2601090/files/figure.png" pageId="20" pageNumber="221">9C</figureCitation>
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,
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<figureCitation id="1379F895E31E8C20A32BFC60FE86FC3B" box="[337,367,943,967]" captionStart="FIGURE 14" captionStartId="20.[151,250,1947,1969]" captionTargetBox="[152,1434,1454,1923]" captionTargetId="figure@20.[151,1435,1454,1925]" captionTargetPageId="20" captionText="FIGURE 14. Sphaerobelum spinatum n. sp., holotype (ZMUC), SEM, A: left antenna, lateral view; B: apical disc with sensory cones; C: gnathochilarium, ventral view. Abbreviations: cP = central pads; d = apical disc; LL = lamellae linguales; LP = lateral palpus; sb = sensilla basiconica; sc = sensory cones; St = stipes; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601102/files/figure.png" pageId="20" pageNumber="221">14</figureCitation>
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,
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<figureCitation id="1379F895E31E8C20A301FC61FE70FC3B" box="[379,409,942,967]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="20" pageNumber="221">15</figureCitation>
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,
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<figureCitation id="1379F895E31E8C20A3DFFC61FE28FC3B" box="[421,449,942,967]" captionStart="FIGURE 16" captionStartId="22.[151,250,1927,1949]" captionTargetBox="[151,1435,373,1906]" captionTargetId="figure@22.[151,1435,373,1906]" captionTargetPageId="22" captionText="FIGURE 16. Sphaerobelum spinatum n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Not to scale." httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="20" pageNumber="221">16</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE31E8C20A2EDFC39FC6CFB87" pageId="20" pageNumber="221" type="materials_examined">
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<paragraph id="8BFDE410E31E8C20A2EDFC39FC6CFB87" blockId="20.[151,1437,1014,1399]" pageId="20" pageNumber="221">
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<emphasis id="B9363802E31E8C20A2EDFC39FE6EFBF3" bold="true" box="[151,391,1014,1039]" pageId="20" pageNumber="221">Material examined:</emphasis>
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Type specimens. 1 M
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<typeStatus id="54F95AB2E31E8C20A0E6FC39FD17FBF3" box="[668,766,1014,1039]" pageId="20" pageNumber="221" type="holotype">holotype</typeStatus>
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(ZMUC00040258) from
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<collectingCountry id="F355A480E31E8C20A656FC38FB8CFBF2" box="[1068,1125,1015,1038]" name="Laos" pageId="20" pageNumber="221">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E31E8C20A60FFC39FB09FBF3" box="[1141,1248,1014,1039]" country="Laos" name="Viangchan" pageId="20" pageNumber="221">Vientiane</collectingRegion>
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Province, Phou Khao Khouay (
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<geoCoordinate id="EE7682D7E31E8C20A332FBD5FE30FBCF" box="[328,473,1050,1075]" direction="north" orientation="latitude" pageId="20" pageNumber="221" precision="1" value="18.339483">N18°20.369'</geoCoordinate>
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N,
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<geoCoordinate id="EE7682D7E31E8C20A386FBD5FD71FBCF" box="[508,664,1050,1075]" direction="east" orientation="longitude" pageId="20" pageNumber="221" precision="1" value="102.808716">102°48.523'E</geoCoordinate>
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),
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<quantity id="4CBA49F5E31E8C20A0D4FBD4FCDBFBCE" box="[686,818,1051,1075]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="20" pageNumber="221" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700–800 m</quantity>
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, strongly disturbed primary evergreen forest, leg.
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<date id="FFFCC2D0E31E8C20A70DFBD5FEE5FBAB" pageId="20" pageNumber="221" value="2008-05-26" valueMax="2008-05-31" valueMin="2008-05-26">26- 31.V.2008</date>
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, A. Solodovnikov & J. Pedersen; 1 F
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<typeStatus id="54F95AB2E31E8C20A0AEFBF0FCD1FBAB" box="[724,824,1087,1111]" pageId="20" pageNumber="221" type="paratype">paratype</typeStatus>
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; ZMUC00040259; same data as
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<typeStatus id="54F95AB2E31E8C20A6C6FBF0FAC9FBAB" box="[1212,1312,1087,1111]" pageId="20" pageNumber="221" type="paratype">paratype</typeStatus>
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;
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<materialsCitation id="3B2AEE4DE31E8C20A755FBF0FC6BFB87" ID-GBIF-Occurrence="2013952577" ID-GBIF-Specimen="3774779754" collectionCode="ZFMK" httpUri="https://www.gbif.org/occurrence/3774779754" pageId="20" pageNumber="221" specimenCode="MYR8101" specimenCount="1" typeStatus="paratype">
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1 M, 1 F
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<typeStatus id="54F95AB2E31E8C20A2EDFBACFF1EFB87" box="[151,247,1123,1147]" pageId="20" pageNumber="221" type="paratype">paratype</typeStatus>
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(
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<collectionCode id="ED537CD5E31E8C20A37CFBACFEBFFB87" box="[262,342,1123,1147]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="221" type="Museum">ZFMK</collectionCode>
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<specimenCode id="DBE44C6BE31E8C20A326FBACFE3DFB87" box="[348,468,1123,1147]" collectionCode="ZFMK" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="20" pageNumber="221" type="Museum">MYR8101</specimenCode>
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& MYR8102), same data as holotype
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE31E8C20A2BDFB49FE59FA8B" pageId="20" pageNumber="221" type="diagnosis">
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<paragraph id="8BFDE410E31E8C20A2BDFB49FE59FA8B" blockId="20.[151,1437,1014,1399]" pageId="20" pageNumber="221">
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<emphasis id="B9363802E31E8C20A2BDFB49FED3FB63" bold="true" box="[199,314,1158,1183]" pageId="20" pageNumber="221">Diagnosis</emphasis>
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:
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<taxonomicName id="4C429F93E31E8C20A330FB48FE26FB62" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[330,463,1159,1182]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
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<emphasis id="B9363802E31E8C20A330FB48FE26FB62" box="[330,463,1159,1182]" italics="true" pageId="20" pageNumber="221">S. spinatum</emphasis>
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</taxonomicName>
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<emphasis id="B9363802E31E8C20A3ADFB49FDF1FB63" bold="true" box="[471,536,1158,1183]" pageId="20" pageNumber="221">
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<taxonomicNameLabel id="A2058579E31E8C20A3ADFB49FDF1FB63" box="[471,536,1158,1183]" pageId="20" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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belongs to a group of
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<taxonomicName id="4C429F93E31E8C20A164FB49FC56FB62" authorityName="Verhoeff" authorityYear="1924" box="[798,959,1158,1182]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="genus">
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<emphasis id="B9363802E31E8C20A164FB49FC56FB62" box="[798,959,1158,1182]" italics="true" pageId="20" pageNumber="221">Sphaerobelum</emphasis>
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</taxonomicName>
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species in which the mesal margin of the femur is extended with several teeth (
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<figureCitation id="1379F895E31E8C20A010FB65FD31FB3F" box="[618,728,1194,1219]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="20" pageNumber="221">Fig. 15B</figureCitation>
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).
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<taxonomicName id="4C429F93E31E8C20A08EFB64FC96FB3E" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[756,895,1195,1218]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
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<emphasis id="B9363802E31E8C20A08EFB64FC96FB3E" box="[756,895,1195,1218]" italics="true" pageId="20" pageNumber="221">S. spinatum</emphasis>
|
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</taxonomicName>
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<emphasis id="B9363802E31E8C20A1F7FB65FC3DFB3F" bold="true" box="[909,980,1194,1219]" pageId="20" pageNumber="221">
|
||
<taxonomicNameLabel id="A2058579E31E8C20A1F7FB65FC3DFB3F" box="[909,980,1194,1219]" pageId="20" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
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shares only with
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<taxonomicName id="4C429F93E31E8C20A6C3FB64FAA8FB3E" box="[1209,1345,1195,1218]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="cattiense">
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||
<emphasis id="B9363802E31E8C20A6C3FB64FAA8FB3E" box="[1209,1345,1195,1218]" italics="true" pageId="20" pageNumber="221">S. cattiense</emphasis>
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||
</taxonomicName>
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and
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<taxonomicName id="4C429F93E31E8C20A7FDFB64FED5FB1A" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="konkakinhense">
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<emphasis id="B9363802E31E8C20A7FDFB64FED5FB1A" italics="true" pageId="20" pageNumber="221">S. konkakinhense</emphasis>
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</taxonomicName>
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a straight telopoditomere 4 (
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<figureCitation id="1379F895E31E8C20A0FBFB01FCE0FB1B" box="[641,777,1230,1255]" captionStart="FIGURE 16" captionStartId="22.[151,250,1927,1949]" captionTargetBox="[151,1435,373,1906]" captionTargetId="figure@22.[151,1435,373,1906]" captionTargetPageId="22" captionText="FIGURE 16. Sphaerobelum spinatum n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Not to scale." httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="20" pageNumber="221">Figs 16E, F</figureCitation>
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).
|
||
<taxonomicName id="4C429F93E31E8C20A167FB00FC48FB1A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[797,929,1231,1254]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
|
||
<emphasis id="B9363802E31E8C20A167FB00FC48FB1A" box="[797,929,1231,1254]" italics="true" pageId="20" pageNumber="221">S. spinatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E31E8C20A1D2FB01FC0EFB1B" bold="true" box="[936,999,1230,1255]" pageId="20" pageNumber="221">
|
||
<taxonomicNameLabel id="A2058579E31E8C20A1D2FB01FC0EFB1B" box="[936,999,1230,1255]" pageId="20" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs in several characters from both other species: mesal margin of prefemur with indentations like the femur (
|
||
<figureCitation id="1379F895E31E8C20A1ADFB3DFBD4FAF7" box="[983,1085,1266,1291]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="20" pageNumber="221">Fig. 15B</figureCitation>
|
||
), male antenna with <60 apical cones (>
|
||
<quantity id="4CBA49F5E31E8C20A280FAD8FED1FAD3" box="[250,312,1302,1327]" metricMagnitude="0" metricUnit="m" metricValue="1.778" pageId="20" pageNumber="221" unit="in" value="70.0">70 in</quantity>
|
||
<taxonomicName id="4C429F93E31E8C20A339FAD8FE1FFAD2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[323,502,1302,1326]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
|
||
<emphasis id="B9363802E31E8C20A339FAD8FE1FFAD2" box="[323,502,1302,1326]" italics="true" pageId="20" pageNumber="221">S. denticulatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E31E8C20A07BFAD9FDADFAD3" bold="true" box="[513,580,1302,1327]" pageId="20" pageNumber="221">
|
||
<taxonomicNameLabel id="A2058579E31E8C20A07BFAD9FDADFAD3" box="[513,580,1302,1327]" pageId="20" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
), tergites shiny, but setose (tergites covered with setose pits in the other species), first laterotergite strongly projecting, extended into a sharp tip (first laterotergite only weakly projecting in the other two species).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF3DB498E31E8C20A2EDF854FB1CF812" httpUri="https://zenodo.org/record/2601102/files/figure.png" pageId="20" pageNumber="221" startId="20.[151,250,1947,1969]" targetBox="[152,1434,1454,1923]" targetPageId="20">
|
||
<paragraph id="8BFDE410E31E8C20A2EDF854FB1CF812" blockId="20.[151,1436,1947,2030]" pageId="20" pageNumber="221">
|
||
<emphasis id="B9363802E31E8C20A2EDF854FEF6F84D" bold="true" box="[151,287,1947,1969]" pageId="20" pageNumber="221">FIGURE 14.</emphasis>
|
||
<taxonomicName id="4C429F93E31E8C20A35EF854FDF0F84D" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[292,537,1947,1969]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="20" pageNumber="221" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
|
||
<emphasis id="B9363802E31E8C20A35EF854FDF0F84D" box="[292,537,1947,1969]" italics="true" pageId="20" pageNumber="221">Sphaerobelum spinatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E31E8C20A064F854FDBEF84D" bold="true" box="[542,599,1947,1969]" pageId="20" pageNumber="221">
|
||
<taxonomicNameLabel id="A2058579E31E8C20A064F854FDBEF84D" box="[542,599,1947,1969]" pageId="20" pageNumber="221" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (ZMUC), SEM,
|
||
<emphasis id="B9363802E31E8C20A112F854FC93F84D" bold="true" box="[872,890,1947,1969]" pageId="20" pageNumber="221">A</emphasis>
|
||
: left antenna, lateral view;
|
||
<emphasis id="B9363802E31E8C20A6F7F854FB74F84D" bold="true" box="[1165,1181,1947,1969]" pageId="20" pageNumber="221">B</emphasis>
|
||
: apical disc with sensory cones;
|
||
<emphasis id="B9363802E31E8C20A29AF876FF12F833" bold="true" box="[224,251,1977,1999]" pageId="20" pageNumber="221">C:</emphasis>
|
||
gnathochilarium, ventral view. Abbreviations:
|
||
<emphasis id="B9363802E31E8C20A09CF876FCE8F833" bold="true" box="[742,769,1977,1999]" pageId="20" pageNumber="221">cP</emphasis>
|
||
= central pads;
|
||
<emphasis id="B9363802E31E8C20A1D7F876FC52F833" bold="true" box="[941,955,1977,1999]" pageId="20" pageNumber="221">d</emphasis>
|
||
= apical disc;
|
||
<emphasis id="B9363802E31E8C20A623F876FB92F833" bold="true" box="[1113,1147,1977,1999]" pageId="20" pageNumber="221">LL</emphasis>
|
||
= lamellae linguales;
|
||
<emphasis id="B9363802E31E8C20A71EF876FA6DF833" bold="true" box="[1380,1412,1977,1999]" pageId="20" pageNumber="221">LP</emphasis>
|
||
= lateral palpus;
|
||
<emphasis id="B9363802E31E8C20A356F817FEADF812" bold="true" box="[300,324,2008,2030]" pageId="20" pageNumber="221">sb</emphasis>
|
||
= sensilla basiconica;
|
||
<emphasis id="B9363802E31E8C20A053F817FDD7F812" bold="true" box="[553,574,2008,2030]" pageId="20" pageNumber="221">sc</emphasis>
|
||
= sensory cones;
|
||
<emphasis id="B9363802E31E8C20A08EF817FCE1F812" bold="true" box="[756,776,2008,2030]" pageId="20" pageNumber="221">St</emphasis>
|
||
= stipes; numbers refer to antennomere number.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C358B79BE31F8C23A2BDFF58FB03FA84" lastPageId="23" lastPageNumber="224" pageId="21" pageNumber="222" type="description">
|
||
<paragraph id="8BFDE410E31F8C21A2BDFF58FCFBFF4C" blockId="21.[151,1437,151,357]" box="[199,786,151,176]" pageId="21" pageNumber="222">
|
||
<emphasis id="B9363802E31F8C21A2BDFF58FEB0FF4C" bold="true" box="[199,345,151,176]" pageId="21" pageNumber="222">Description.</emphasis>
|
||
Based on
|
||
<typeStatus id="54F95AB2E31F8C21A3B4FF58FDD9FF4C" box="[462,560,151,176]" pageId="21" pageNumber="222" type="holotype">holotype</typeStatus>
|
||
and both
|
||
<typeStatus id="54F95AB2E31F8C21A0E5FF57FCE7FF4C" box="[671,782,152,176]" pageId="21" pageNumber="222" type="paratype">paratypes</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E31F8C21A2BDFF73FA74FEE1" blockId="21.[151,1437,151,357]" pageId="21" pageNumber="222">
|
||
<emphasis id="B9363802E31F8C21A2BDFF73FE96FF29" bold="true" box="[199,383,188,213]" pageId="21" pageNumber="222">Measurements:</emphasis>
|
||
Body length:
|
||
<typeStatus id="54F95AB2E31F8C21A050FF73FD65FF29" box="[554,652,188,213]" pageId="21" pageNumber="222" type="holotype">holotype</typeStatus>
|
||
male: length ca
|
||
<quantity id="4CBA49F5E31F8C21A129FF73FC57FF28" box="[851,958,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.69" pageId="21" pageNumber="222" unit="mm" value="26.9">26.9 mm</quantity>
|
||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E31F8C21A76BFF73FA90FF28" box="[1297,1401,188,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.3599999999999999" pageId="21" pageNumber="222" unit="mm" value="13.6">13.6 mm</quantity>
|
||
(= broadest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E31F8C21A033FF10FD76FF0B" box="[585,671,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="21" pageNumber="222" unit="mm" value="7.5">7.5 mm</quantity>
|
||
(= highest). Female: length ca
|
||
<quantity id="4CBA49F5E31F8C21A187FF10FB8DFF0B" box="[1021,1124,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="21" pageNumber="222" unit="mm" value="32.5">32.5 mm</quantity>
|
||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E31F8C21A2EDFECBFF15FEE0" box="[151,252,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="21" pageNumber="222" unit="mm" value="14.5">14.5 mm</quantity>
|
||
, of tergite 7 =
|
||
<quantity id="4CBA49F5E31F8C21A3E7FECBFDE8FEE0" box="[413,513,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="21" pageNumber="222" unit="mm" value="15.7">15.7 mm</quantity>
|
||
(=broadest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E31F8C21A1B4FECAFBCCFEE0" box="[974,1061,261,284]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="21" pageNumber="222" unit="mm" value="7.7">7.7 mm</quantity>
|
||
, of tergite 9 =
|
||
<quantity id="4CBA49F5E31F8C21A6BCFECBFAFDFEE0" box="[1222,1300,260,284]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="21" pageNumber="222" unit="mm" value="9.1">9.1mm</quantity>
|
||
(= highest).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E31F8C21A2EDFEE8FBE0FE99" blockId="21.[151,1437,151,357]" pageId="21" pageNumber="222">
|
||
<emphasis id="B9363802E31F8C21A2EDFEE8FEF1FEBC" bold="true" box="[151,280,295,320]" pageId="21" pageNumber="222">Coloration</emphasis>
|
||
: in preserved specimens black, paratergite depressions and groove of thoracic shield dark green (
|
||
<figureCitation id="1379F895E31F8C21A714FEE8FF54FE99" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="21" pageNumber="222">Fig. 6D</figureCitation>
|
||
). Antennae brown-orange, legs light brown, but prefemur and femur green.
|
||
</paragraph>
|
||
<caption id="DF3DB498E31F8C21A2EDF8D8FB21F896" httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="21" pageNumber="222" startId="21.[151,250,1815,1837]" targetBox="[151,1435,409,1793]" targetPageId="21">
|
||
<paragraph id="8BFDE410E31F8C21A2EDF8D8FB21F896" blockId="21.[151,1437,1815,1898]" pageId="21" pageNumber="222">
|
||
<emphasis id="B9363802E31F8C21A2EDF8D8FEF0F8D1" bold="true" box="[151,281,1815,1837]" pageId="21" pageNumber="222">FIGURE 15</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E31F8C21A35CF8D8FDF5F8D1" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[294,540,1815,1837]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
|
||
<emphasis id="B9363802E31F8C21A35CF8D8FDF5F8D1" box="[294,540,1815,1837]" italics="true" pageId="21" pageNumber="222">Sphaerobelum spinatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E31F8C21A058F8D8FDB5F8D1" bold="true" box="[546,604,1815,1837]" pageId="21" pageNumber="222">
|
||
<taxonomicNameLabel id="A2058579E31F8C21A058F8D8FDB5F8D1" box="[546,604,1815,1837]" pageId="21" pageNumber="222" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B9363802E31F8C21A013F8D8FD70F8D1" bold="true" box="[617,665,1815,1837]" pageId="21" pageNumber="222">A, B</emphasis>
|
||
: holotype (ZMUC);
|
||
<emphasis id="B9363802E31F8C21A114F8D8FC49F8D1" bold="true" box="[878,928,1815,1837]" pageId="21" pageNumber="222">C, D</emphasis>
|
||
: female paratype (ZFMK MYR8102);
|
||
<emphasis id="B9363802E31F8C21A74AF8D8FAABF8D1" bold="true" box="[1328,1346,1815,1837]" pageId="21" pageNumber="222">A</emphasis>
|
||
: coxa of first left leg with first stigmatic plate;
|
||
<emphasis id="B9363802E31F8C21A048F8FAFDAAF8B7" bold="true" box="[562,579,1845,1867]" pageId="21" pageNumber="222">B</emphasis>
|
||
: 9th left leg;
|
||
<emphasis id="B9363802E31F8C21A0A1F8FAFD1FF8B7" bold="true" box="[731,758,1845,1867]" pageId="21" pageNumber="222">C:</emphasis>
|
||
coxa and prefemur of second leg with vulva;
|
||
<emphasis id="B9363802E31F8C21A693F8FAFB12F8B7" bold="true" box="[1257,1275,1845,1867]" pageId="21" pageNumber="222">D</emphasis>
|
||
: subanal plate. Abbreviations:
|
||
<emphasis id="B9363802E31F8C21A34EF89BFEBAF896" bold="true" box="[308,339,1876,1898]" pageId="21" pageNumber="222">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E31F8C21A3D1F89BFE56F896" bold="true" box="[427,447,1876,1898]" pageId="21" pageNumber="222">O</emphasis>
|
||
= operculum of vulva;
|
||
<emphasis id="B9363802E31F8C21A0D7F89BFD3BF896" bold="true" box="[685,722,1876,1898]" pageId="21" pageNumber="222">Pre</emphasis>
|
||
= prefemur;
|
||
<emphasis id="B9363802E31F8C21A12DF89BFC63F896" bold="true" box="[855,906,1876,1898]" pageId="21" pageNumber="222">St-Pl</emphasis>
|
||
= stigmatic plate. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFDE410E31F8C22A2BDF858FD94FEBC" blockId="21.[151,1436,1943,2004]" lastBlockId="22.[151,1437,151,320]" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">
|
||
<emphasis id="B9363802E31F8C21A2BDF858FEEEF84C" bold="true" box="[199,263,1943,1968]" pageId="21" pageNumber="222">Head</emphasis>
|
||
: Eyes with>60 ocelli. Aberrant ocellus located inside antennal groove. Antennae short, with rounded joints, extending posteriorly to leg-pair 5. Lengths of antennomeres: 1>2=3=4=5<<6 (
|
||
<figureCitation id="1379F895E31F8C21A620F874FB2BF828" box="[1114,1218,1979,2004]" captionStart="FIGURE 14" captionStartId="20.[151,250,1947,1969]" captionTargetBox="[152,1434,1454,1923]" captionTargetId="figure@20.[151,1435,1454,1925]" captionTargetPageId="20" captionText="FIGURE 14. Sphaerobelum spinatum n. sp., holotype (ZMUC), SEM, A: left antenna, lateral view; B: apical disc with sensory cones; C: gnathochilarium, ventral view. Abbreviations: cP = central pads; d = apical disc; LL = lamellae linguales; LP = lateral palpus; sb = sensilla basiconica; sc = sensory cones; St = stipes; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601102/files/figure.png" pageId="21" pageNumber="222">Fig. 14A</figureCitation>
|
||
). All antennomeres densely pubescent, sensilla basiconica surrounding apical disc. Shape of antennae sexually dimorphic, cylindrical in female, thickened, apically widened and slightly flattened in male (
|
||
<figureCitation id="1379F895E31C8C22A1CBFF73FBF0FF28" box="[945,1049,188,213]" captionStart="FIGURE 14" captionStartId="20.[151,250,1947,1969]" captionTargetBox="[152,1434,1454,1923]" captionTargetId="figure@20.[151,1435,1454,1925]" captionTargetPageId="20" captionText="FIGURE 14. Sphaerobelum spinatum n. sp., holotype (ZMUC), SEM, A: left antenna, lateral view; B: apical disc with sensory cones; C: gnathochilarium, ventral view. Abbreviations: cP = central pads; d = apical disc; LL = lamellae linguales; LP = lateral palpus; sb = sensilla basiconica; sc = sensory cones; St = stipes; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601102/files/figure.png" pageId="22" pageNumber="223">Fig. 14B</figureCitation>
|
||
). Apical disc with ca 52/52 apical cones (male) (
|
||
<figureCitation id="1379F895E31C8C22A32FFF10FE2FFF04" box="[341,454,223,248]" captionStart="FIGURE 14" captionStartId="20.[151,250,1947,1969]" captionTargetBox="[152,1434,1454,1923]" captionTargetId="figure@20.[151,1435,1454,1925]" captionTargetPageId="20" captionText="FIGURE 14. Sphaerobelum spinatum n. sp., holotype (ZMUC), SEM, A: left antenna, lateral view; B: apical disc with sensory cones; C: gnathochilarium, ventral view. Abbreviations: cP = central pads; d = apical disc; LL = lamellae linguales; LP = lateral palpus; sb = sensilla basiconica; sc = sensory cones; St = stipes; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601102/files/figure.png" pageId="22" pageNumber="223">Fig. 14b</figureCitation>
|
||
), or 30–36 (female). Organ of Tömösváry located inside antennal groove.
|
||
<emphasis id="B9363802E31C8C22A2EDFECBFE8FFEE1" bold="true" box="[151,358,260,285]" pageId="22" pageNumber="223">Gnathochilarium</emphasis>
|
||
: Damaged. Structure typical of the order (
|
||
<figureCitation id="1379F895E31C8C22A144FECBFC4CFEE0" box="[830,933,260,285]" captionStart="FIGURE 14" captionStartId="20.[151,250,1947,1969]" captionTargetBox="[152,1434,1454,1923]" captionTargetId="figure@20.[151,1435,1454,1925]" captionTargetPageId="20" captionText="FIGURE 14. Sphaerobelum spinatum n. sp., holotype (ZMUC), SEM, A: left antenna, lateral view; B: apical disc with sensory cones; C: gnathochilarium, ventral view. Abbreviations: cP = central pads; d = apical disc; LL = lamellae linguales; LP = lateral palpus; sb = sensilla basiconica; sc = sensory cones; St = stipes; numbers refer to antennomere number." httpUri="https://zenodo.org/record/2601102/files/figure.png" pageId="22" pageNumber="223">Fig. 14C</figureCitation>
|
||
). Palpi with sensory cones arranged in single field. Rudimentary lateral palps not visible.
|
||
</paragraph>
|
||
<caption id="DF3DB498E31C8C22A2EDF848FAA9F826" httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="22" pageNumber="223" startId="22.[151,250,1927,1949]" targetBox="[151,1435,373,1906]" targetPageId="22">
|
||
<paragraph id="8BFDE410E31C8C22A2EDF848FAA9F826" blockId="22.[151,1436,1927,2010]" pageId="22" pageNumber="223">
|
||
<emphasis id="B9363802E31C8C22A2EDF848FEF2F861" bold="true" box="[151,283,1927,1949]" pageId="22" pageNumber="223">FIGURE 16</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E31C8C22A351F848FDCBF861" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[299,546,1927,1949]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="spinatum" status="sp. nov.">
|
||
<emphasis id="B9363802E31C8C22A351F848FDCBF861" box="[299,546,1927,1949]" italics="true" pageId="22" pageNumber="223">Sphaerobelum spinatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E31C8C22A051F848FD8EF861" bold="true" box="[555,615,1927,1949]" pageId="22" pageNumber="223">
|
||
<taxonomicNameLabel id="A2058579E31C8C22A051F848FD8EF861" box="[555,615,1927,1949]" pageId="22" pageNumber="223" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (ZMUC);
|
||
<emphasis id="B9363802E31C8C22A13BF848FCBAF861" bold="true" box="[833,851,1927,1949]" pageId="22" pageNumber="223">A</emphasis>
|
||
: coxa of second left leg with gonopore;
|
||
<emphasis id="B9363802E31C8C22A778F848FAFAF861" bold="true" box="[1282,1299,1927,1949]" pageId="22" pageNumber="223">B</emphasis>
|
||
: left anterior telopod, anterior view;
|
||
<emphasis id="B9363802E31C8C22A3FEF869FE77F840" bold="true" box="[388,414,1958,1980]" pageId="22" pageNumber="223">C:</emphasis>
|
||
left anterior telopod, posterior view;
|
||
<emphasis id="B9363802E31C8C22A160F869FCC5F840" bold="true" box="[794,812,1958,1980]" pageId="22" pageNumber="223">D</emphasis>
|
||
: left anterior telopod, lateral view;
|
||
<emphasis id="B9363802E31C8C22A6EEF869FB4CF840" bold="true" box="[1172,1189,1958,1980]" pageId="22" pageNumber="223">E</emphasis>
|
||
: right posterior telopod, anterior view;
|
||
<emphasis id="B9363802E31C8C22A350F80BFED0F826" bold="true" box="[298,313,1988,2010]" pageId="22" pageNumber="223">F</emphasis>
|
||
: right posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E31C8C22A115F80BFC67F826" bold="true" box="[879,910,1988,2010]" pageId="22" pageNumber="223">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E31C8C22A19CF80BFBDFF826" bold="true" box="[998,1078,1988,2010]" pageId="22" pageNumber="223">Syn-Cx</emphasis>
|
||
= syncoxite. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFDE410E31D8C23A2BDFF58FE04FF4C" blockId="23.[151,1437,151,1437]" box="[199,493,151,176]" pageId="23" pageNumber="224">
|
||
<emphasis id="B9363802E31D8C23A2BDFF58FEACFF4C" bold="true" box="[199,325,151,176]" pageId="23" pageNumber="224">Mandibles</emphasis>
|
||
: not dissected.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E31D8C23A2BDFF73FC17FCE9" blockId="23.[151,1437,151,1437]" pageId="23" pageNumber="224">
|
||
<emphasis id="B9363802E31D8C23A2BDFF73FE62FF29" bold="true" box="[199,395,188,213]" pageId="23" pageNumber="224">Stigmatic plates</emphasis>
|
||
: first stigmatic plate rounded, apex well-rounded, straight towards coxa 1 (
|
||
<figureCitation id="1379F895E31D8C23A75AFF73FA79FF29" box="[1312,1424,188,213]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="23" pageNumber="224">Fig. 15A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E31D8C23A2EDFF10FEA9FF04" bold="true" box="[151,320,223,248]" pageId="23" pageNumber="224">Laterotergites</emphasis>
|
||
: laterotergite 1 strongly projecting into a sharp tip. Laterotergite 2 with a broad, stout, much shorter projection.
|
||
<emphasis id="B9363802E31D8C23A36DFECBFE86FEE1" bold="true" box="[279,367,260,285]" pageId="23" pageNumber="224">Collum</emphasis>
|
||
: with glabrous surface, margins with few isolated setae.
|
||
<emphasis id="B9363802E31D8C23A18AFECBFB40FEE1" bold="true" box="[1008,1193,260,285]" pageId="23" pageNumber="224">Thoracic shield</emphasis>
|
||
: surface glabrous but covered with tiny pits, setae only in grooves. Slope towards groove without anterior keels but with 4 or 5 posterior keels.
|
||
<emphasis id="B9363802E31D8C23A29EFE83FEAAFE99" bold="true" box="[228,323,332,357]" pageId="23" pageNumber="224">Tergites</emphasis>
|
||
: surface anterior half densely setose with short setae, posterior half glabrous, shiny. Tips of paratergites of midbody tergites projecting posteriorly (
|
||
<figureCitation id="1379F895E31D8C23A17EFEA0FCB7FE74" box="[772,862,367,392]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="23" pageNumber="224">Fig. 6D</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E31D8C23A108FEA0FBE0FE74" bold="true" box="[882,1033,367,392]" pageId="23" pageNumber="224">Endotergum</emphasis>
|
||
: inner section lacking any spines or setae. Middle area with a single row of large, sparse, elliptical, cuticular impressions. Distance between impressions twice as wide as their diameter. Apically, 2-3 dense rows of long marginal bristles, the tips of the longest setae slightly protruding beyond tergal margin (
|
||
<figureCitation id="1379F895E31D8C23A163FE13FC9CFE08" box="[793,885,476,501]" captionStart="FIGURE 9" captionStartId="13.[151,250,1423,1445]" captionTargetBox="[152,1434,1021,1400]" captionTargetId="figure@13.[152,1435,1021,1402]" captionTargetPageId="13" captionText="FIGURE 9. Endoterga of the body ring 9 of different Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum spinatum n. sp.; D: Sphaerobelum splendidum n. sp. Abbreviations: cp = cuticular patterns; iA = inner area." httpUri="https://zenodo.org/record/2601090/files/figure.png" pageId="23" pageNumber="224">Fig. 9C</figureCitation>
|
||
). Bristles not smooth, but with numerous small spinicles.
|
||
<emphasis id="B9363802E31D8C23A372FE30FE79FDE4" bold="true" box="[264,400,511,536]" pageId="23" pageNumber="224">Anal shield</emphasis>
|
||
: large, in both sexes well-rounded. Surface in both sexes completely covered by tiny setae located in small pits. Underside with a single, short, black, locking carina, located close to last laterotergite.
|
||
<emphasis id="B9363802E31D8C23A726FDEBFA7DFDC1" bold="true" box="[1372,1428,548,573]" pageId="23" pageNumber="224">Legs</emphasis>
|
||
: leg-pair 1 with 2 or 3 ventral spines, leg-pair 2 with 6, leg-pair 3 with 7 or 8. First two leg-pairs without an apical spine. Leg-pairs 4–21 with 10 ventral spines and one dorso-apical spine. In leg 9, femur 1.6 times, tarsus 3.7 times longer than wide (
|
||
<figureCitation id="1379F895E31D8C23A316FD40FE3CFD54" box="[364,469,655,680]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="23" pageNumber="224">Fig. 15B</figureCitation>
|
||
). All podomeres densely setose. Coxa with a large and marginally toothed process. Coxa process absent at first leg (
|
||
<figureCitation id="1379F895E31D8C23A074FD7BFD9CFD31" box="[526,629,692,717]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="23" pageNumber="224">Fig. 15A</figureCitation>
|
||
) and sharply projecting at second (
|
||
<figureCitation id="1379F895E31D8C23A673FD7BFB9CFD31" box="[1033,1141,692,717]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="23" pageNumber="224">Figs 15C</figureCitation>
|
||
,
|
||
<figureCitation id="1379F895E31D8C23A6F8FD7BFB5FFD31" box="[1154,1206,692,717]" captionStart="FIGURE 16" captionStartId="22.[151,250,1927,1949]" captionTargetBox="[151,1435,373,1906]" captionTargetId="figure@22.[151,1435,373,1906]" captionTargetPageId="22" captionText="FIGURE 16. Sphaerobelum spinatum n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Not to scale." httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="23" pageNumber="224">16A</figureCitation>
|
||
). Prefemur at apical margin with a projection laterally and mesally. Lateral projection triangular and sharply edged, juxtaposed to coxal process. Femur extended mesally into a dentate margin featuring 12–14 teeth.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E31D8C23A2BDFCD0FD14FC59" blockId="23.[151,1437,151,1437]" pageId="23" pageNumber="224">
|
||
<emphasis id="B9363802E31D8C23A2BDFCD0FE13FCC4" bold="true" box="[199,506,799,824]" pageId="23" pageNumber="224">Female sexual characters</emphasis>
|
||
: female gravid. Vulva large, covering 2/3 of coxa, extending mesally to anterior third of prefemur (
|
||
<figureCitation id="1379F895E31D8C23A30DFC8BFE0AFCA1" box="[375,483,836,861]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="23" pageNumber="224">Fig. 15C</figureCitation>
|
||
). Operculum rounded, very slightly invaginated medially, mesal margin strongly projecting into a well-rounded lobe twice as high as remaining operculum.
|
||
<emphasis id="B9363802E31D8C23A198FCA8FB6FFC7C" bold="true" box="[994,1158,871,896]" pageId="23" pageNumber="224">Subanal plate</emphasis>
|
||
: large and wide, divided by a suture into two halves. Densely setose (
|
||
<figureCitation id="1379F895E31D8C23A0F3FC43FD19FC59" box="[649,752,908,933]" captionStart="FIGURE 15" captionStartId="21.[151,250,1815,1837]" captionTargetBox="[151,1435,409,1793]" captionTargetId="figure@21.[150,1436,409,1794]" captionTargetPageId="21" captionText="FIGURE 15. Sphaerobelum spinatum n. sp., A, B: holotype (ZMUC); C, D: female paratype (ZFMK MYR8102); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa and prefemur of second leg with vulva; D: subanal plate. Abbreviations: Cx = coxa; O = operculum of vulva; Pre = prefemur; St-Pl = stigmatic plate. Not to scale." httpUri="https://zenodo.org/record/2601104/files/figure.png" pageId="23" pageNumber="224">Fig. 15D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E31D8C23A2BDFC60FB03FA84" blockId="23.[151,1437,151,1437]" pageId="23" pageNumber="224">
|
||
<emphasis id="B9363802E31D8C23A2BDFC60FE08FC34" bold="true" box="[199,481,943,968]" pageId="23" pageNumber="224">Male sexual characters</emphasis>
|
||
: gonopore covered with a single, undivided, circular, sclerotized plate (
|
||
<figureCitation id="1379F895E31D8C23A75EFC60FA66FC34" box="[1316,1423,943,968]" captionStart="FIGURE 16" captionStartId="22.[151,250,1927,1949]" captionTargetBox="[151,1435,373,1906]" captionTargetId="figure@22.[151,1435,373,1906]" captionTargetPageId="22" captionText="FIGURE 16. Sphaerobelum spinatum n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Not to scale." httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="23" pageNumber="224">Fig. 16A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E31D8C23A2EDFC1BFE83FC11" bold="true" box="[151,362,980,1005]" pageId="23" pageNumber="224">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E31D8C23A303FC1BFE26FC11" box="[377,463,980,1005]" captionStart="FIGURE 16" captionStartId="22.[151,250,1927,1949]" captionTargetBox="[151,1435,373,1906]" captionTargetId="figure@22.[151,1435,373,1906]" captionTargetPageId="22" captionText="FIGURE 16. Sphaerobelum spinatum n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Not to scale." httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="23" pageNumber="224">Figs 16</figureCitation>
|
||
B–D): consisting of 4 telopoditomeres above syncoxite. Telopoditomere 1 rectangular, as long as wide. Telopoditomere 2 large, as long as telopoditomere 1. Process of telopoditomere 2 located posteriorly, not visible in anterior view. Process of telopoditomere 2 wide, well-rounded, projecting to half of telopoditomere 3. Telopoditomere 3 small, slightly shorter than telopoditomere 4. telopoditomere 4 cylindrical, well-rounded, posterior side with two small spines. All telopoditomeres covered with long setae.
|
||
<emphasis id="B9363802E31D8C23A757FBABFF13FB5C" bold="true" pageId="23" pageNumber="224">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E31D8C23A373FB48FE79FB5C" box="[265,400,1159,1184]" captionStart="FIGURE 16" captionStartId="22.[151,250,1927,1949]" captionTargetBox="[151,1435,373,1906]" captionTargetId="figure@22.[151,1435,373,1906]" captionTargetPageId="22" captionText="FIGURE 16. Sphaerobelum spinatum n. sp., holotype (ZMUC); A: coxa of second left leg with gonopore; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left anterior telopod, lateral view; E: right posterior telopod, anterior view; F: right posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Not to scale." httpUri="https://zenodo.org/record/2601106/files/figure.png" pageId="23" pageNumber="224">Figs 16E, F</figureCitation>
|
||
): telopoditomere 1 short, half as long as wide. Immovable finger (process of telopoditomere 2) as long as movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex, well rounded apically, apex only slightly wider than base. Telopoditomere 3 rectangular, well rounded, apically with a well-rounded extension carrying a small spine. Telopoditomere 4 only slightly shorter and slightly more slender than telopoditomere 3, 2.1 times longer than wide, apically weakly tapering, straight, not curved, with 3 small spines at margin towards immovable finger. In anterior view telopoditomere 1–3 covered by setae, in posterior view telopoditomeres 2–4 mostly glabrous except for a few setae at the margins.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE31D8C23A2BDFA4BFAE2FA61" box="[199,1291,1412,1437]" pageId="23" pageNumber="224" type="etymology">
|
||
<paragraph id="8BFDE410E31D8C23A2BDFA4BFAE2FA61" blockId="23.[151,1437,151,1437]" box="[199,1291,1412,1437]" pageId="23" pageNumber="224">
|
||
<emphasis id="B9363802E31D8C23A2BDFA4BFE74FA61" bold="true" box="[199,413,1412,1437]" pageId="23" pageNumber="224">Derivatio nominis</emphasis>
|
||
: spinatum, noun in apposition, after the sharp process of the first laterotergite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |