290 lines
39 KiB
XML
290 lines
39 KiB
XML
<document id="28ED77C33CC6F0612722E406268E9EA8" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E3108C28A2EDFA4BFDA7FA3C" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum peterjaegeri Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="229" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="227" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:title id="AEE7E06E3F1C21176573A9A462659A3F">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="846860A34D9C743BE2F6C77BE7298FEE" type="personal">
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<mods:namePart id="8BFCEAB241FF9DCE201EB7FF333533CA">Wesener, Thomas</mods:namePart>
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<mods:title id="7EFD90B816837C6627B3F249007C63C7">Zootaxa</mods:title>
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<mods:part id="4758B5335B33986C11C63BF8B7C13DDF">
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<mods:date id="EE66F51D40340311E1D59D0B42B4DC89">2019</mods:date>
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<mods:number id="873FF91570DBFBA32DDE77A20ACCE0E9">2019-03-01</mods:number>
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<mods:number id="81D90D57DB362ABAB82E8CE3FBD20E3A">4563</mods:number>
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<mods:identifier id="2D6F05D16FFEB70419F1A62C0DC3E51E" type="DOI">10.11646/zootaxa.4563.2.1</mods:identifier>
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<treatment id="03EB5506E3108C28A2EDFA4BFDA7FA3C" ID-DOI="http://doi.org/10.5281/zenodo.5934532" ID-GBIF-Taxon="154474552" ID-Zenodo-Dep="5934532" LSID="urn:lsid:plazi:treatment:03EB5506E3108C28A2EDFA4BFDA7FA3C" httpUri="http://treatment.plazi.org/id/03EB5506E3108C28A2EDFA4BFDA7FA3C" lastPageId="28" lastPageNumber="229" pageId="26" pageNumber="227">
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<subSubSection id="C358B79BE3108C2EA2EDFA4BFD9DFA62" box="[151,628,1412,1438]" pageId="26" pageNumber="227" type="nomenclature">
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<paragraph id="8BFDE410E3108C2EA2EDFA4BFD9DFA62" blockId="26.[151,628,1412,1471]" box="[151,628,1412,1438]" pageId="26" pageNumber="227">
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<heading id="D0B5537CE3108C2EA2EDFA4BFD9DFA62" bold="true" box="[151,628,1412,1438]" fontSize="11" level="1" pageId="26" pageNumber="227" reason="1">
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<emphasis id="B9363802E3108C2EA2EDFA4BFD9DFA62" bold="true" box="[151,628,1412,1438]" pageId="26" pageNumber="227">
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<taxonomicName id="4C429F93E3108C2EA2EDFA4BFE34FA62" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,477,1412,1438]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
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<emphasis id="B9363802E3108C2EA2EDFA4BFE34FA62" bold="true" box="[151,477,1412,1438]" italics="true" pageId="26" pageNumber="227">Sphaerobelum peterjaegeri</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E3108C2EA39EFA4BFD9DFA62" box="[484,628,1412,1438]" pageId="26" pageNumber="227" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE3108C2EA2EDFA69FEDEFA43" box="[151,311,1446,1471]" pageId="26" pageNumber="227" type="description">
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<paragraph id="8BFDE410E3108C2EA2EDFA69FEDEFA43" blockId="26.[151,628,1412,1471]" box="[151,311,1446,1471]" pageId="26" pageNumber="227">
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<figureCitation id="1379F895E3108C2EA2EDFA69FEF9FA43" box="[151,272,1446,1471]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="26" pageNumber="227">Figures 6F</figureCitation>
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,
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<figureCitation id="1379F895E3108C2EA361FA68FEDEFA43" box="[283,311,1447,1471]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="26" pageNumber="227">19</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE3108C2EA2EDFA21FDC0F9B3" pageId="26" pageNumber="227" type="materials_examined">
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<paragraph id="8BFDE410E3108C2EA2EDFA21FDC0F9B3" blockId="26.[151,1436,1518,2011]" pageId="26" pageNumber="227">
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<emphasis id="B9363802E3108C2EA2EDFA21FE6AF9FB" bold="true" box="[151,387,1518,1543]" pageId="26" pageNumber="227">Material examined:</emphasis>
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<materialsCitation id="3B2AEE4DE3108C2EA3F1FA20FDCDF9B3" ID-GBIF-Occurrence="2013952573" collectingDate="2008-02-28" collectionCode="SMF" collectorName="P. Jager" country="Laos" county="Nam Khan" elevation="328" latitude="19.73586" location="Houay Kho" longLatPrecision="1" longitude="102.277084" municipality="Ban Pak Bak" pageId="26" pageNumber="227" specimenCode="SD553" specimenCount="1" stateProvince="Louangphabang" typeStatus="holotype">
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1 M,
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<typeStatus id="54F95AB2E3108C2EA3B3FA21FDC2F9FB" box="[457,555,1518,1543]" pageId="26" pageNumber="227" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED537CD5E3108C2EA040FA20FD9DF9FB" box="[570,628,1519,1543]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="227">SMF</collectionCode>
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<specimenCode id="DBE44C6BE3108C2EA006FA21FD22F9FB" box="[636,715,1518,1543]" collectionCode="SMF" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="26" pageNumber="227">SD553</specimenCode>
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) from
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<collectingCountry id="F355A480E3108C2EA163FA20FCBBF9FA" box="[793,850,1519,1542]" name="Laos" pageId="26" pageNumber="227">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E3108C2EA124FA20FB9EF9FB" box="[862,1143,1518,1543]" country="Laos" name="Louangphabang" pageId="26" pageNumber="227">Luang Prabang Province</collectingRegion>
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, SE
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<collectingRegion id="49862AF2E3108C2EA6D6FA20FAB3F9FB" box="[1196,1370,1518,1543]" country="Laos" name="Louangphabang" pageId="26" pageNumber="227">Luang Prabang</collectingRegion>
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,
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<collectingCounty id="629C9C9CE3108C2EA71CFA20FF3EF9D7" pageId="26" pageNumber="227">Nam Khan</collectingCounty>
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,
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<collectingMunicipality id="6B997E6AE3108C2EA298F9DCFE90F9D7" box="[226,377,1554,1579]" pageId="26" pageNumber="227">Ban Pak Bak</collectingMunicipality>
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,
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<location id="8E9DB2CBE3108C2EA3FFF9DCFDE3F9D7" LSID="urn:lsid:plazi:treatment:03EB5506E3108C28A2EDFA4BFDA7FA3C:8E9DB2CBE3108C2EA3FFF9DCFDE3F9D7" box="[389,522,1554,1579]" country="Laos" county="Nam Khan" latitude="19.73586" longLatPrecision="1" longitude="102.277084" municipality="Ban Pak Bak" name="Houay Kho" pageId="26" pageNumber="227" stateProvince="Louangphabang">Houay Kho</location>
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,
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<quantity id="4CBA49F5E3108C2EA06DF9DCFD89F9D6" box="[535,608,1555,1579]" metricMagnitude="2" metricUnit="m" metricValue="3.28" pageId="26" pageNumber="227" unit="m" value="328.0">
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<elevation id="006F0323E3108C2EA06DF9DCFD89F9D6" box="[535,608,1555,1579]" metricMagnitude="2" metricUnit="m" metricValue="3.28" pageId="26" pageNumber="227" unit="m" value="328.0">328 m</elevation>
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</quantity>
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, (
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<geoCoordinate id="EE7682D7E3108C2EA00CF9DCFCF8F9D7" box="[630,785,1555,1579]" direction="north" orientation="latitude" pageId="26" pageNumber="227" precision="1" value="19.73586">N19°44'09.1"</geoCoordinate>
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,
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<geoCoordinate id="EE7682D7E3108C2EA164F9DDFC2CF9D7" box="[798,965,1554,1579]" direction="east" orientation="longitude" pageId="26" pageNumber="227" precision="1" value="102.277084">102°16'37.5"E</geoCoordinate>
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), bank of stream, under stone, moist sand and gravel, leg.
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<date id="FFFCC2D0E3108C2EA330F9F8FE28F9B3" box="[330,449,1591,1615]" pageId="26" pageNumber="227" value="2008-02-28">
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<collectingDate id="EFB83B38E3108C2EA330F9F8FE28F9B3" box="[330,449,1591,1615]" pageId="26" pageNumber="227" value="2008-02-28">28.II.2008</collectingDate>
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</date>
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,
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<collectorName id="26B781C6E3108C2EA3B7F9F8FDCDF9B3" box="[461,548,1591,1615]" pageId="26" pageNumber="227">P. Jäger</collectorName>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE3108C2EA2BDF995FEF3F893" pageId="26" pageNumber="227" type="diagnosis">
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<paragraph id="8BFDE410E3108C2EA2BDF995FEF3F893" blockId="26.[151,1436,1518,2011]" pageId="26" pageNumber="227">
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<emphasis id="B9363802E3108C2EA2BDF995FED3F98F" bold="true" box="[199,314,1626,1651]" pageId="26" pageNumber="227">Diagnosis</emphasis>
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:
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<taxonomicName id="4C429F93E3108C2EA332F994FE05F98E" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[328,492,1627,1650]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
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<emphasis id="B9363802E3108C2EA332F994FE05F98E" box="[328,492,1627,1650]" italics="true" pageId="26" pageNumber="227">S. peterjaegeri</emphasis>
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</taxonomicName>
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<emphasis id="B9363802E3108C2EA38EF995FDDAF98F" bold="true" box="[500,563,1626,1651]" pageId="26" pageNumber="227">
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<taxonomicNameLabel id="A2058579E3108C2EA38EF995FDDAF98F" box="[500,563,1626,1651]" pageId="26" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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belongs to a group of
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<taxonomicName id="4C429F93E3108C2EA154F994FC26F98F" authorityName="Verhoeff" authorityYear="1924" box="[814,975,1627,1651]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="genus">
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<emphasis id="B9363802E3108C2EA154F994FC26F98F" box="[814,975,1627,1651]" italics="true" pageId="26" pageNumber="227">Sphaerobelum</emphasis>
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</taxonomicName>
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species in which the mesal margin of the femur is unmodified (
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<figureCitation id="1379F895E3108C2EA3F0F9B1FE19F96B" box="[394,496,1662,1687]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="26" pageNumber="227">Fig. 19B</figureCitation>
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).
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<taxonomicName id="4C429F93E3108C2EA07EF9B0FD41F96A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[516,680,1663,1686]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
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<emphasis id="B9363802E3108C2EA07EF9B0FD41F96A" box="[516,680,1663,1686]" italics="true" pageId="26" pageNumber="227">S. peterjaegeri</emphasis>
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</taxonomicName>
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<emphasis id="B9363802E3108C2EA0D5F9B1FD04F96B" bold="true" box="[687,749,1662,1687]" pageId="26" pageNumber="227">
|
||
<taxonomicNameLabel id="A2058579E3108C2EA0D5F9B1FD04F96B" box="[687,749,1662,1687]" pageId="26" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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shares only with
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<taxonomicName id="4C429F93E3108C2EA1C8F9B0FBAAF96A" authorityName="Attems" authorityYear="1953" box="[946,1091,1663,1686]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="separatum">
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<emphasis id="B9363802E3108C2EA1C8F9B0FBAAF96A" box="[946,1091,1663,1686]" italics="true" pageId="26" pageNumber="227">S. separatum</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C429F93E3108C2EA628F9B0FB09F96A" box="[1106,1248,1663,1686]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="truncatum">
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<emphasis id="B9363802E3108C2EA628F9B0FB09F96A" box="[1106,1248,1663,1686]" italics="true" pageId="26" pageNumber="227">S. truncatum</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C429F93E3108C2EA76CF9B0FA75F96A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1302,1436,1662,1686]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="lachneeis" status="sp. nov.">
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<emphasis id="B9363802E3108C2EA76CF9B0FA75F96A" box="[1302,1436,1662,1686]" italics="true" pageId="26" pageNumber="227">S. lachneeis</emphasis>
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</taxonomicName>
|
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<emphasis id="B9363802E3108C2EA2EDF96DFF31F947" bold="true" box="[151,216,1698,1723]" pageId="26" pageNumber="227">
|
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<taxonomicNameLabel id="A2058579E3108C2EA2EDF96DFF31F947" box="[151,216,1698,1723]" pageId="26" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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the absence of a "hook" at telopoditomere 4 of the posterior telopods (
|
||
<figureCitation id="1379F895E3108C2EA67DF96DFB7EF947" box="[1031,1175,1698,1723]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="26" pageNumber="227">Figs 19G, H</figureCitation>
|
||
).
|
||
<taxonomicName id="4C429F93E3108C2EA6D6F96CFABBF946" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1196,1362,1699,1722]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
|
||
<emphasis id="B9363802E3108C2EA6D6F96CFABBF946" box="[1196,1362,1699,1722]" italics="true" pageId="26" pageNumber="227">S. peterjaegeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3108C2EA721F96DFA72F947" bold="true" box="[1371,1435,1698,1723]" pageId="26" pageNumber="227">
|
||
<taxonomicNameLabel id="A2058579E3108C2EA721F96DFA72F947" box="[1371,1435,1698,1723]" pageId="26" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C429F93E3108C2EA359F908FE5CF922" authorityName="Attems" authorityYear="1953" box="[291,437,1735,1758]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="separatum">
|
||
<emphasis id="B9363802E3108C2EA359F908FE5CF922" box="[291,437,1735,1758]" italics="true" pageId="26" pageNumber="227">S. separatum</emphasis>
|
||
</taxonomicName>
|
||
in the absence of a lateral extension on podomere 3 of the anterior telopod, as well as the absence of an incurved process on telopoditomere 4 of the posterior telopods.
|
||
<taxonomicName id="4C429F93E3108C2EA67DF924FB44F8FE" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1031,1197,1771,1794]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
|
||
<emphasis id="B9363802E3108C2EA67DF924FB44F8FE" box="[1031,1197,1771,1794]" italics="true" pageId="26" pageNumber="227">S. peterjaegeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3108C2EA6CEF925FB1AF8FF" bold="true" box="[1204,1267,1770,1795]" pageId="26" pageNumber="227">
|
||
<taxonomicNameLabel id="A2058579E3108C2EA6CEF925FB1AF8FF" box="[1204,1267,1770,1795]" pageId="26" pageNumber="227" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is very similar to
|
||
<taxonomicName id="4C429F93E3108C2EA2CEF8C0FEADF8DA" box="[180,324,1807,1830]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="truncatum">
|
||
<emphasis id="B9363802E3108C2EA2CEF8C0FEADF8DA" box="[180,324,1807,1830]" italics="true" pageId="26" pageNumber="227">S. truncatum</emphasis>
|
||
</taxonomicName>
|
||
, but differs in several characters: <70 apical cones on the antenna (>
|
||
<quantity id="4CBA49F5E3108C2EA634F8C0FB60F8DB" box="[1102,1161,1806,1831]" metricMagnitude="0" metricUnit="m" metricValue="2.032" pageId="26" pageNumber="227" unit="in" value="80.0">80 in</quantity>
|
||
<taxonomicName id="4C429F93E3108C2EA6EAF8C0FAC9F8DA" box="[1168,1312,1807,1830]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="truncatum">
|
||
<emphasis id="B9363802E3108C2EA6EAF8C0FAC9F8DA" box="[1168,1312,1807,1830]" italics="true" pageId="26" pageNumber="227">S. truncatum</emphasis>
|
||
</taxonomicName>
|
||
), antennae and legs orange-red (yellow in
|
||
<taxonomicName id="4C429F93E3108C2EA381F8FCFD62F8B6" box="[507,651,1843,1866]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="truncatum">
|
||
<emphasis id="B9363802E3108C2EA381F8FCFD62F8B6" box="[507,651,1843,1866]" italics="true" pageId="26" pageNumber="227">S. truncatum</emphasis>
|
||
</taxonomicName>
|
||
), femur 1.7 times longer than wide (2.0 times longer than wide in
|
||
<taxonomicName id="4C429F93E3108C2EA7FDF8FCFEE3F892" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="26" pageNumber="227" phylum="Arthropoda" rank="species" species="truncatum">
|
||
<emphasis id="B9363802E3108C2EA7FDF8FCFEE3F892" italics="true" pageId="26" pageNumber="227">S. truncatum</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE3108C28A2BDF8B5FCD3FA84" lastPageId="28" lastPageNumber="229" pageId="26" pageNumber="227" type="description">
|
||
<paragraph id="8BFDE410E3108C2EA2BDF8B5FBA6F827" blockId="26.[151,1436,1518,2011]" pageId="26" pageNumber="227">
|
||
<emphasis id="B9363802E3108C2EA2BDF8B5FDF1F86F" bold="true" box="[199,536,1914,1939]" pageId="26" pageNumber="227">Description. Measurements:</emphasis>
|
||
Body length:
|
||
<typeStatus id="54F95AB2E3108C2EA0C2F8B5FCF3F86F" box="[696,794,1914,1939]" pageId="26" pageNumber="227" type="holotype">holotype</typeStatus>
|
||
male: length ca
|
||
<quantity id="4CBA49F5E3108C2EA1AEF8B4FBD2F86E" box="[980,1083,1915,1939]" metricMagnitude="-2" metricUnit="m" metricValue="2.38" pageId="26" pageNumber="227" unit="mm" value="23.8">23.8 mm</quantity>
|
||
. Width of thoracic shield =
|
||
<quantity id="4CBA49F5E3108C2EA716F8B5FF2AF84A" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="26" pageNumber="227" unit="mm" value="11.6">11.6 mm</quantity>
|
||
(= broadest). Height of thoracic shield =
|
||
<quantity id="4CBA49F5E3108C2EA0DEF851FD15F84A" box="[676,764,1950,1975]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="26" pageNumber="227" unit="mm" value="6.9">6.9 mm</quantity>
|
||
(= highest).
|
||
<emphasis id="B9363802E3108C2EA1EBF851FBFBF84B" bold="true" box="[913,1042,1950,1975]" pageId="26" pageNumber="227">Coloration</emphasis>
|
||
: in preserved specimens brown to orange (
|
||
<figureCitation id="1379F895E3108C2EA288F80DFEA1F827" box="[242,328,1986,2011]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="26" pageNumber="227">Fig. 6F</figureCitation>
|
||
). Collum dark olive brown, same as head. Antennae and legs orange.
|
||
</paragraph>
|
||
<caption id="DF3DB498E3118C2FA2EDF8B4FBD1F810" httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="27" pageNumber="228" startId="27.[151,250,1915,1937]" targetBox="[188,1408,195,1899]" targetPageId="27">
|
||
<paragraph id="8BFDE410E3118C2FA2EDF8B4FBD1F810" blockId="27.[151,1437,1915,2028]" pageId="27" pageNumber="228">
|
||
<emphasis id="B9363802E3118C2FA2EDF8B4FEF0F86D" bold="true" box="[151,281,1915,1937]" pageId="27" pageNumber="228">FIGURE 19</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E3118C2FA35DF8B4FDD2F86D" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[295,571,1915,1937]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="peterjaegeri" status="sp. nov.">
|
||
<emphasis id="B9363802E3118C2FA35DF8B4FDD2F86D" box="[295,571,1915,1937]" italics="true" pageId="27" pageNumber="228">Sphaerobelum peterjaegeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3118C2FA038F8B4FD95F86D" bold="true" box="[578,636,1915,1937]" pageId="27" pageNumber="228">
|
||
<taxonomicNameLabel id="A2058579E3118C2FA038F8B4FD95F86D" box="[578,636,1915,1937]" pageId="27" pageNumber="228" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (SMF);
|
||
<emphasis id="B9363802E3118C2FA141F8B4FCA4F86D" bold="true" box="[827,845,1915,1937]" pageId="27" pageNumber="228">A</emphasis>
|
||
: coxa of first left leg with first stigmatic plate;
|
||
<emphasis id="B9363802E3118C2FA74EF8B4FAACF86D" bold="true" box="[1332,1349,1915,1937]" pageId="27" pageNumber="228">B</emphasis>
|
||
: 9th left leg;
|
||
<emphasis id="B9363802E3118C2FA2B9F856FF3CF853" bold="true" box="[195,213,1945,1967]" pageId="27" pageNumber="228">C</emphasis>
|
||
: coxa of second left leg with gonopore;
|
||
<emphasis id="B9363802E3118C2FA016F856FD97F853" bold="true" box="[620,638,1945,1967]" pageId="27" pageNumber="228">D</emphasis>
|
||
: left anterior telopod, anterior view;
|
||
<emphasis id="B9363802E3118C2FA189F856FBE5F853" bold="true" box="[1011,1036,1945,1967]" pageId="27" pageNumber="228">E:</emphasis>
|
||
left anterior telopod, posterior view;
|
||
<emphasis id="B9363802E3118C2FA7FFF856FA7DF853" bold="true" box="[1413,1428,1945,1967]" pageId="27" pageNumber="228">F</emphasis>
|
||
: left anterior telopod, lateral view;
|
||
<emphasis id="B9363802E3118C2FA075F877FDCAF832" bold="true" box="[527,547,1976,1998]" pageId="27" pageNumber="228">G</emphasis>
|
||
: right posterior telopod, anterior view;
|
||
<emphasis id="B9363802E3118C2FA1ADF877FC02F832" bold="true" box="[983,1003,1976,1998]" pageId="27" pageNumber="228">H</emphasis>
|
||
: right posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E3118C2FA34EF819FEBAF810" bold="true" box="[308,339,2006,2028]" pageId="27" pageNumber="228">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E3118C2FA3D1F819FE37F810" bold="true" box="[427,478,2006,2028]" pageId="27" pageNumber="228">St-Pl</emphasis>
|
||
= stigmatic plate;
|
||
<emphasis id="B9363802E3118C2FA0E6F819FD05F810" bold="true" box="[668,748,2006,2028]" pageId="27" pageNumber="228">Syn-Cx</emphasis>
|
||
= syncoxite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFDE410E3168C28A2BDFF58FCCEFEBC" blockId="28.[151,1437,151,1472]" pageId="28" pageNumber="229">
|
||
<emphasis id="B9363802E3168C28A2BDFF58FEEEFF4C" bold="true" box="[199,263,151,176]" pageId="28" pageNumber="229">Head</emphasis>
|
||
: Eyes with ca. 60 ocelli. Aberrant ocellus located inside antennal groove. Antennae short, with rounded joints, extending posteriorly to leg-pair 3. First antennomere with cuticular scales. Lengths of antennomeres: 1=2=3=4=5<<6. Shape of apical joint thickened, apically widened. Apical disc with ca 66/68 apical cones. Organ of Tömösváry located inside antennal groove.
|
||
<emphasis id="B9363802E3168C28A0DBFECBFC86FEE1" bold="true" box="[673,879,260,285]" pageId="28" pageNumber="229">Gnathochilarium</emphasis>
|
||
: structure typical of the order. Palpi with sensory cones arranged in a single field.
|
||
<emphasis id="B9363802E3168C28A078FEE8FD69FEBC" bold="true" box="[514,640,295,320]" pageId="28" pageNumber="229">Mandibles</emphasis>
|
||
: not dissected.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3168C28A2BDFE83FC21FCC4" blockId="28.[151,1437,151,1472]" pageId="28" pageNumber="229">
|
||
<emphasis id="B9363802E3168C28A2BDFE83FE65FE99" bold="true" box="[199,396,332,357]" pageId="28" pageNumber="229">Stigmatic plates</emphasis>
|
||
: first stigmatic plate well-rounded (
|
||
<figureCitation id="1379F895E3168C28A132FE83FC50FE98" box="[840,953,332,357]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="28" pageNumber="229">Fig. 19A</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3168C28A1AEFE83FB94FE99" bold="true" box="[980,1149,332,357]" pageId="28" pageNumber="229">Laterotergites</emphasis>
|
||
: laterotergite 1 slightly projecting into a sharp but short tip. Laterotergite 2 with a broad, stout, much shorter projection.
|
||
<emphasis id="B9363802E3168C28A6A8FEA0FAC2FE74" bold="true" box="[1234,1323,367,392]" pageId="28" pageNumber="229">Collum</emphasis>
|
||
: with few isolated setae on entire surface.
|
||
<emphasis id="B9363802E3168C28A380FE5BFD58FE51" bold="true" box="[506,689,404,429]" pageId="28" pageNumber="229">Thoracic shield</emphasis>
|
||
: surface hirsute like tergites, longer setae only in grooves. Shallow grooves beset with numerous long setae, slope towards groove with 4 or 5 posterior keels, anterior keels absent.
|
||
<emphasis id="B9363802E3168C28A2EDFE13FF1FFE09" bold="true" box="[151,246,476,501]" pageId="28" pageNumber="229">Tergites</emphasis>
|
||
: surface densely setose with short setae. Tips of paratergites of midbody tergites projecting posteriorly (
|
||
<figureCitation id="1379F895E3168C28A2E5FE30FF10FDE4" box="[159,249,511,536]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="28" pageNumber="229">Fig. 6F</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3168C28A368FE30FE40FDE4" bold="true" box="[274,425,511,536]" pageId="28" pageNumber="229">Endotergum</emphasis>
|
||
: posterior margin conspicuously tesselated. Inner section without any setae or spines. Middle area with a single row of dense elliptical, cuticular impressions, distance between impressions slightly smaller than their diameter. Apically, 2-3 dense rows of short marginal bristles, the longest just protruding up to 2/ 3 of the distance to tergal margin. Bristles not smooth, but with numerous small spicules.
|
||
<emphasis id="B9363802E3168C28A6BEFDA3FAA6FD79" bold="true" box="[1220,1359,620,645]" pageId="28" pageNumber="229">Anal shield</emphasis>
|
||
: wellrounded. Surface completely covered by tiny setae. Underside with a single, very long black, locking carina, located close to last laterotergite.
|
||
<emphasis id="B9363802E3168C28A06FFD7BFDA4FD31" bold="true" box="[533,589,692,717]" pageId="28" pageNumber="229">Legs</emphasis>
|
||
: leg-pair 1 with 2 ventral spines, leg-pair 2 with 4, leg-pair 3 with 6. First two leg-pairs without an apical spine. Leg-pairs 4–21 with 7–10 ventral spines and one dorso-apical spine. In leg 9, femur 1.7 times, tarsus 4.6 times longer than wide (
|
||
<figureCitation id="1379F895E3168C28A0AEFD33FCD3FCE8" box="[724,826,764,789]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="28" pageNumber="229">Fig. 19B</figureCitation>
|
||
). All podomeres densely setose. Coxa with a large and well-rounded process. Coxal process absent at leg 1 and 2 (
|
||
<figureCitation id="1379F895E3168C28A154FCD0FC52FCC4" box="[814,955,799,824]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="28" pageNumber="229">Figs 19A, C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3168C28A2BDFC8BFE61FCA1" blockId="28.[151,1437,151,1472]" box="[199,392,836,861]" pageId="28" pageNumber="229">Female unknown</paragraph>
|
||
<paragraph id="8BFDE410E3168C28A2BDFCA8FCD3FA84" blockId="28.[151,1437,151,1472]" pageId="28" pageNumber="229">
|
||
<emphasis id="B9363802E3168C28A2BDFCA8FE08FC7C" bold="true" box="[199,481,871,896]" pageId="28" pageNumber="229">Male sexual characters</emphasis>
|
||
: gonopore covered with a single, undivided, circular, sclerotized plate (
|
||
<figureCitation id="1379F895E3168C28A75FFCA8FA66FC7C" box="[1317,1423,871,896]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="28" pageNumber="229">Fig. 19C</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3168C28A2EDFC43FE82FC59" bold="true" box="[151,363,908,933]" pageId="28" pageNumber="229">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3168C28A300FC43FE26FC58" box="[378,463,908,933]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="28" pageNumber="229">Figs 19</figureCitation>
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D–F): consisting of 4 telopoditomeres above syncoxite. Telopoditomere 1 rectangular, as long as wide. Telopoditomere 2 large, as long as telopoditomere 1. Process of telopoditomere 2 located posteriorly, visible in anterior view. Process of telopoditomere 2 wide, projecting to half of telopoditomere 4, apically with a well-rounded tip. Telopoditomere 3 cylindrical, straight, apically slightly tapering, as long as telopoditomere 4. telopoditomere 4 cylindrical, apex slightly bent posteriorly. Posterior side with a black sclerotized spot and two small, triangular spines. Telopoditomeres 1–4 with only few, mainly marginal long setae.
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||
<emphasis id="B9363802E3168C28A2EDFBABFE99FB81" bold="true" box="[151,368,1124,1149]" pageId="28" pageNumber="229">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E3168C28A3FBFBABFDE6FB80" box="[385,527,1124,1149]" captionStart="FIGURE 19" captionStartId="27.[151,250,1915,1937]" captionTargetBox="[188,1408,195,1899]" captionTargetId="figure@27.[188,1408,193,1899]" captionTargetPageId="27" captionText="FIGURE 19. Sphaerobelum peterjaegeri n. sp., holotype (SMF); A: coxa of first left leg with first stigmatic plate; B: 9th left leg; C: coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view; G: right posterior telopod, anterior view; H: right posterior telopod, posterior view. Abbreviations: Cx = coxa; St-Pl = stigmatic plate; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601113/files/figure.png" pageId="28" pageNumber="229">Figs 19G, H</figureCitation>
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): telopoditomere 1 elongated, twice as long as wide. Immovable finger (process of telopoditomere 2) shorter than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex, well rounded, apex not expanded, only slightly wider than base. Telopoditomere 3 elongated, towards immovable finger close to apex with a slight extension carrying a small spine. Telopoditomere 4 shorter and more slender than telopoditomere 3, 2.1 times longer than wide, apically weakly tapering, very slightly curved towards immovable finger, at inner margin with two small spines. Telopoditomere 1 and 2 at both sides covered by setae. Telopoditomere 3 at base at inner margin with a few setae, remaining parts of telopoditomeres 3 and 4 almost glabrous.
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</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="C358B79BE3168C28A2BDFA4BFDA7FA3C" pageId="28" pageNumber="229" type="etymology">
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||
<paragraph id="8BFDE410E3168C28A2BDFA4BFDA7FA3C" blockId="28.[151,1437,151,1472]" pageId="28" pageNumber="229">
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||
<emphasis id="B9363802E3168C28A2BDFA4BFE49FA61" bold="true" box="[199,416,1412,1437]" pageId="28" pageNumber="229">Derivatio nominis</emphasis>
|
||
: peterjaegeri, noun in apposition, after the Arachnologist Peter Jäger, great collector of arthropods in
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<collectingCountry id="F355A480E3168C28A348FA66FE81FA3C" box="[306,360,1449,1472]" name="Laos" pageId="28" pageNumber="229">Laos</collectingCountry>
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and other countries.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
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||
</document> |