312 lines
38 KiB
XML
312 lines
38 KiB
XML
<document id="B968D2DB91C0E96B90852402E0575704" ID-CLB-Dataset="28149" ID-DOI="10.11646/zootaxa.4563.2.1" ID-GBIF-Dataset="e6037c5d-a96f-41e5-8e70-71f2659659e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="2601072" ID-ZooBank="CF79B01B-8B5F-4B3A-B642-2CADE4B339AF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171433896" checkinUser="plazi" docAuthor="Wesener, Thomas" docDate="2019" docId="03EB5506E3058C20A2EDFD33FEEDFCC4" docLanguage="en" docName="zootaxa.4563.2.1.pdf" docOrigin="Zootaxa 4563 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sphaerobelum denticulatum Wesener 2019, new species" docType="treatment" docVersion="9" lastPageNumber="221" masterDocId="FFD22D7EE30A8C34A27AFFCFFFE9FFFC" masterDocTitle="First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)" masterLastPageNumber="248" masterPageNumber="201" pageNumber="216" updateTime="1698703176884" updateUser="ExternalLinkService">
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<mods:title id="8E4DEAA1830772FB5D15708A1D72C7AD">First records of giant pill-millipedes from Laos (Diplopoda, Sphaerotheriida, Zephroniidae)</mods:title>
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<mods:name id="1EAAEA7370B55E2B026AA0745AD8D532" type="personal">
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<mods:namePart id="8BEEA2325C236D0CB8475EC225CC795D">Wesener, Thomas</mods:namePart>
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<mods:title id="627123207C1D11F3D41CDD790584362F">Zootaxa</mods:title>
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<mods:part id="E341E4C16FE744DC04E3130FFEC9E4D4">
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<mods:date id="9416C1CBF8D16AAA32D8E98B4164B56E">2019</mods:date>
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<mods:number id="7F497D926AB4976A0AFD71E519BFEDC1">2019-03-01</mods:number>
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<mods:number id="CBFA623DDC2741211373339697B26BF2">4563</mods:number>
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<mods:identifier id="2DA2169A9671B5DA16DD422AE747D323" type="DOI">10.11646/zootaxa.4563.2.1</mods:identifier>
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<mods:identifier id="ECB44779F742EC7E487DB91B3F77A821" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="376C243FD967709383D94BEE725AFB05" type="Zenodo-Dep">2601072</mods:identifier>
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<treatment id="03EB5506E3058C20A2EDFD33FEEDFCC4" ID-DOI="http://doi.org/10.5281/zenodo.5934526" ID-GBIF-Taxon="154474555" ID-Zenodo-Dep="5934526" LSID="urn:lsid:plazi:treatment:03EB5506E3058C20A2EDFD33FEEDFCC4" httpUri="http://treatment.plazi.org/id/03EB5506E3058C20A2EDFD33FEEDFCC4" lastPageId="20" lastPageNumber="221" pageId="15" pageNumber="216">
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<subSubSection id="C358B79BE3058C3BA2EDFD33FD6CFCEA" box="[151,645,764,790]" pageId="15" pageNumber="216" type="nomenclature">
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<paragraph id="8BFDE410E3058C3BA2EDFD33FD6CFCEA" blockId="15.[151,645,764,823]" box="[151,645,764,790]" pageId="15" pageNumber="216">
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<heading id="D0B5537CE3058C3BA2EDFD33FD6CFCEA" bold="true" box="[151,645,764,790]" fontSize="11" level="1" pageId="15" pageNumber="216" reason="1">
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<emphasis id="B9363802E3058C3BA2EDFD33FD6CFCEA" bold="true" box="[151,645,764,790]" pageId="15" pageNumber="216">
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<taxonomicName id="4C429F93E3058C3BA2EDFD33FE07FCEA" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[151,494,764,790]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
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<emphasis id="B9363802E3058C3BA2EDFD33FE07FCEA" bold="true" box="[151,494,764,790]" italics="true" pageId="15" pageNumber="216">Sphaerobelum denticulatum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2058579E3058C3BA38FFD33FD6CFCEA" box="[501,645,764,790]" pageId="15" pageNumber="216" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C358B79BE3058C3BA2EDFCD1FED4FCCB" box="[151,317,798,823]" pageId="15" pageNumber="216" type="description">
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<paragraph id="8BFDE410E3058C3BA2EDFCD1FED4FCCB" blockId="15.[151,645,764,823]" box="[151,317,798,823]" pageId="15" pageNumber="216">
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<figureCitation id="1379F895E3058C3BA2EDFCD1FEFCFCCB" box="[151,277,798,823]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="15" pageNumber="216">Figures 6C</figureCitation>
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,
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<figureCitation id="1379F895E3058C3BA35AFCD0FED4FCCB" box="[288,317,799,823]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="15" pageNumber="216">13</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C358B79BE3058C3BA2EDFCA9FE0CFC3B" pageId="15" pageNumber="216" type="materials_examined">
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<paragraph id="8BFDE410E3058C3BA2EDFCA9FE0CFC3B" blockId="15.[151,1437,870,2011]" pageId="15" pageNumber="216">
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<emphasis id="B9363802E3058C3BA2EDFCA9FE6AFC83" bold="true" box="[151,387,870,895]" pageId="15" pageNumber="216">Material examined:</emphasis>
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<materialsCitation id="3B2AEE4DE3058C3BA3F1FCA8FE0BFC3B" ID-GBIF-Occurrence="2013952572" collectingDate="2010-09-18" collectionCode="MHNG" collectorName="P. Schwendinger" country="Laos" elevation="1020" latitude="20.605556" location="Tad Lak" longLatPrecision="21" longitude="102.02112" pageId="15" pageNumber="216" specimenCount="1" stateProvince="Oudomxai" typeStatus="holotype">
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1 M
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<typeStatus id="54F95AB2E3058C3BA3BBFCA9FDCAFC83" box="[449,547,870,895]" pageId="15" pageNumber="216" type="holotype">holotype</typeStatus>
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(
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<collectionCode id="ED537CD5E3058C3BA048FCA8FD60FC83" box="[562,649,871,895]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="15" pageNumber="216" type="Museum">MHNG</collectionCode>
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LT-10/12) from
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<collectingCountry id="F355A480E3058C3BA13CFCA8FC96FC82" box="[838,895,871,894]" name="Laos" pageId="15" pageNumber="216">Laos</collectingCountry>
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,
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<collectingRegion id="49862AF2E3058C3BA1F6FCA9FB99FC83" box="[908,1136,870,895]" country="Laos" name="Oudomxai" pageId="15" pageNumber="216">Oudomxai Province</collectingRegion>
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, ca
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<quantity id="4CBA49F5E3058C3BA6E6FCA8FB3FFC83" box="[1180,1238,870,895]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="216" unit="km" value="3.0">3 km</quantity>
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E of
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<location id="8E9DB2CBE3058C3BA76FFCA9FA9AFC83" LSID="urn:lsid:plazi:treatment:03EB5506E3058C20A2EDFD33FEEDFCC4:8E9DB2CBE3058C3BA76FFCA9FA9AFC83" box="[1301,1395,870,895]" country="Laos" latitude="20.605556" longLatPrecision="21" longitude="102.02112" name="Tad Lak" pageId="15" pageNumber="216" stateProvince="Oudomxai">Tad Lak</location>
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11, SE of
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<collectingRegion id="49862AF2E3058C3BA2A5FC45FEBCFC5F" box="[223,341,906,931]" country="Laos" name="Oudomxai" pageId="15" pageNumber="216">Oudomxai</collectingRegion>
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city (= Muang Xai) (
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<geoCoordinate id="EE7682D7E3058C3BA031FC45FD3AFC5F" box="[587,723,906,931]" direction="north" orientation="latitude" pageId="15" pageNumber="216" precision="15" value="20.605556">N20°36'20"</geoCoordinate>
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;
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||
<geoCoordinate id="EE7682D7E3058C3BA0A5FC45FC90FC5F" box="[735,889,906,931]" direction="east" orientation="longitude" pageId="15" pageNumber="216" precision="15" value="102.02112">102°01'16" E</geoCoordinate>
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),
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||
<quantity id="4CBA49F5E3058C3BA1F5FC44FC0EFC5E" box="[911,999,907,931]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="15" pageNumber="216" unit="m" value="1020.0">
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<elevation id="006F0323E3058C3BA1F5FC44FC0EFC5E" box="[911,999,907,931]" metricMagnitude="3" metricUnit="m" metricValue="1.02" pageId="15" pageNumber="216" unit="m" value="1020.0">1020 m</elevation>
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</quantity>
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||
, undisturbed evergreen hill forest, leg.
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<date id="FFFCC2D0E3058C3BA2EDFC60FEF0FC3B" box="[151,281,943,967]" pageId="15" pageNumber="216" value="2010-09-18">
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<collectingDate id="EFB83B38E3058C3BA2EDFC60FEF0FC3B" box="[151,281,943,967]" pageId="15" pageNumber="216" value="2010-09-18">18.IX.2010</collectingDate>
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</date>
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,
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<collectorName id="26B781C6E3058C3BA35EFC60FE0BFC3B" box="[292,482,942,967]" pageId="15" pageNumber="216">P. Schwendinger</collectorName>
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</materialsCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C358B79BE3058C3BA2BDFC1DFDDDFBAB" pageId="15" pageNumber="216" type="diagnosis">
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<paragraph id="8BFDE410E3058C3BA2BDFC1DFDDDFBAB" blockId="15.[151,1437,870,2011]" pageId="15" pageNumber="216">
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||
<emphasis id="B9363802E3058C3BA2BDFC1DFEAAFC17" bold="true" box="[199,323,978,1003]" pageId="15" pageNumber="216">Diagnosis:</emphasis>
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<taxonomicName id="4C429F93E3058C3BA336FC1CFE12FC17" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[332,507,979,1003]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
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<emphasis id="B9363802E3058C3BA336FC1CFE12FC17" box="[332,507,979,1003]" italics="true" pageId="15" pageNumber="216">S. denticulatum</emphasis>
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</taxonomicName>
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<emphasis id="B9363802E3058C3BA07EFC1DFDACFC17" bold="true" box="[516,581,978,1003]" pageId="15" pageNumber="216">
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<taxonomicNameLabel id="A2058579E3058C3BA07EFC1DFDACFC17" box="[516,581,978,1003]" pageId="15" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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belongs to a group of
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<taxonomicName id="4C429F93E3058C3BA130FC1CFC02FC17" authorityName="Verhoeff" authorityYear="1924" box="[842,1003,979,1003]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="216" phylum="Arthropoda" rank="genus">
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<emphasis id="B9363802E3058C3BA130FC1CFC02FC17" box="[842,1003,979,1003]" italics="true" pageId="15" pageNumber="216">Sphaerobelum</emphasis>
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</taxonomicName>
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species in which the mesal margin of the femur is extended with several teeth (
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<figureCitation id="1379F895E3058C3BA009FC39FD37FBF3" box="[627,734,1014,1039]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="15" pageNumber="216">Fig. 13A</figureCitation>
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).
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<taxonomicName id="4C429F93E3058C3BA089FC38FC4AFBF2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[755,931,1014,1038]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
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||
<emphasis id="B9363802E3058C3BA089FC38FC4AFBF2" box="[755,931,1014,1038]" italics="true" pageId="15" pageNumber="216">S. denticulatum</emphasis>
|
||
</taxonomicName>
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||
<emphasis id="B9363802E3058C3BA1D7FC39FC07FBF3" bold="true" box="[941,1006,1014,1039]" pageId="15" pageNumber="216">
|
||
<taxonomicNameLabel id="A2058579E3058C3BA1D7FC39FC07FBF3" box="[941,1006,1014,1039]" pageId="15" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
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||
</emphasis>
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||
shares only with
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<taxonomicName id="4C429F93E3058C3BA6C6FC38FAD5FBF2" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[1212,1340,1014,1038]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="216" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
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||
<emphasis id="B9363802E3058C3BA6C6FC38FAD5FBF2" box="[1212,1340,1014,1038]" italics="true" pageId="15" pageNumber="216">S. phouloei</emphasis>
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||
</taxonomicName>
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||
<emphasis id="B9363802E3058C3BA73FFC39FA6FFBF3" bold="true" box="[1349,1414,1014,1039]" pageId="15" pageNumber="216">
|
||
<taxonomicNameLabel id="A2058579E3058C3BA73FFC39FA6FFBF3" box="[1349,1414,1014,1039]" pageId="15" pageNumber="216" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
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a curved telopoditomere 4 of the posterior telopods which overlaps the immovable finger (
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<figureCitation id="1379F895E3058C3BA6FCFBD5FAFAFBCF" box="[1158,1299,1050,1075]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="15" pageNumber="216">Figs 13D, E</figureCitation>
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). See above for a separation of these two species.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C358B79BE3058C20A2BDFBADFD04FD13" lastPageId="20" lastPageNumber="221" pageId="15" pageNumber="216" type="description">
|
||
<paragraph id="8BFDE410E3058C3BA2BDFBADFC13FB1B" blockId="15.[151,1437,870,2011]" pageId="15" pageNumber="216">
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||
<emphasis id="B9363802E3058C3BA2BDFBADFDF4FB87" bold="true" box="[199,541,1122,1147]" pageId="15" pageNumber="216">Description. Measurements:</emphasis>
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||
Body length: length ca
|
||
<quantity id="4CBA49F5E3058C3BA13EFBACFC59FB86" box="[836,944,1123,1147]" metricMagnitude="-2" metricUnit="m" metricValue="2.27" pageId="15" pageNumber="216" unit="mm" value="22.7">22.7 mm</quantity>
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||
. Width, of thoracic shield =
|
||
<quantity id="4CBA49F5E3058C3BA774FBADFA9EFB86" box="[1294,1399,1122,1147]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="15" pageNumber="216" unit="mm" value="11.5">11.5 mm</quantity>
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||
(= broadest). Height, of thoracic shield =
|
||
<quantity id="4CBA49F5E3058C3BA03CFB49FD77FB62" box="[582,670,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" pageId="15" pageNumber="216" unit="mm" value="6.3">6.3 mm</quantity>
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||
, tergite 7 =
|
||
<quantity id="4CBA49F5E3058C3BA159FB49FC90FB62" box="[803,889,1158,1183]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="15" pageNumber="216" unit="mm" value="6.9">6.9 mm</quantity>
|
||
(= highest).
|
||
<emphasis id="B9363802E3058C3BA67DFB49FB61FB63" bold="true" box="[1031,1160,1158,1183]" pageId="15" pageNumber="216">Coloration</emphasis>
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||
: in preserved specimens tergites black, thoracic shield grooves and paratergite depressions dark green. Head and collum black (
|
||
<figureCitation id="1379F895E3058C3BA74EFB65FA66FB3F" box="[1332,1423,1194,1219]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="15" pageNumber="216">Fig. 6C</figureCitation>
|
||
). Antennae dirty green, antennomere 6 yellow. Legs light brown, tarsus darker.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3058C3BA2BDFB3DFB05FA67" blockId="15.[151,1437,870,2011]" pageId="15" pageNumber="216">
|
||
<emphasis id="B9363802E3058C3BA2BDFB3DFEEEFAF7" bold="true" box="[199,263,1266,1291]" pageId="15" pageNumber="216">Head</emphasis>
|
||
: Eyes with ca. 55 ocelli. Antennae short, extending back to leg 3, with rounded joints. Lengths of antennomeres: 1>2=3=4=5<<6, 6 as large as 3–5 combined. All antennomeres densely pubescent, sensilla basiconica surrounding apical disc. Last antennomere thickened, cylindrical. Apical disc with 68/74 apical cones. All antennomeres covered by long setae. Organ of Tömösváry located inside antennal groove.
|
||
<emphasis id="B9363802E3058C3BA6BFFA91FA7DFA8B" bold="true" box="[1221,1428,1374,1399]" pageId="15" pageNumber="216">Gnathochilarium</emphasis>
|
||
: structure typical of the order. Palpi sensory cones located in single field.
|
||
<emphasis id="B9363802E3058C3BA1BCFA4DFBADFA67" bold="true" box="[966,1092,1410,1435]" pageId="15" pageNumber="216">Mandibles</emphasis>
|
||
: not dissected.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3058C3BA2BDFA69FA74F9D7" blockId="15.[151,1437,870,2011]" pageId="15" pageNumber="216">
|
||
<emphasis id="B9363802E3058C3BA2BDFA69FE6CFA43" bold="true" box="[199,389,1446,1471]" pageId="15" pageNumber="216">Stigmatic plates</emphasis>
|
||
: first stigmatic plate rounded, apex well-rounded, straight towards coxa 1.
|
||
<emphasis id="B9363802E3058C3BA6AFFA69FA97FA43" bold="true" box="[1237,1406,1446,1471]" pageId="15" pageNumber="216">Laterotergites</emphasis>
|
||
: 1 and 2 with a short, well-rounded projection.
|
||
<emphasis id="B9363802E3058C3BA0D3FA05FCE8FA1F" bold="true" box="[681,769,1482,1507]" pageId="15" pageNumber="216">Collum</emphasis>
|
||
: with glabrous surface, margins with few isolated setae.
|
||
<emphasis id="B9363802E3058C3BA2EDFA21FEBAF9FB" bold="true" box="[151,339,1518,1543]" pageId="15" pageNumber="216">Thoracic shield</emphasis>
|
||
: surface glabrous, surface chagrinated unlike tergites, setae only in grooves. Shallow grooves beset with numerous long setae, slope towards groove with 2 or 3 weak anterior and 2 or 3 posterior keels.
|
||
</paragraph>
|
||
<paragraph id="8BFDE410E3058C3BA2EDF9F9FBD5F827" blockId="15.[151,1437,870,2011]" pageId="15" pageNumber="216">
|
||
<emphasis id="B9363802E3058C3BA2EDF9F9FF1FF9B3" bold="true" box="[151,246,1590,1615]" pageId="15" pageNumber="216">Tergites</emphasis>
|
||
: glabrous, but entirely covered by tiny pits. Tips of paratergites of midbody tergites projecting posteriorly (
|
||
<figureCitation id="1379F895E3058C3BA2E5F995FF11F98F" box="[159,248,1626,1651]" captionStart="FIGURE 6" captionStartId="11.[151,250,1916,1938]" captionTargetBox="[150,1423,987,1897]" captionTargetId="figure@11.[150,1436,983,1903]" captionTargetPageId="11" captionText="FIGURE 6. Habitus multi-layer stacked photographs of Sphaerobelum species. A: Sphaerobelum bolavensis n. sp.; B: Sphaerobelum phouloei n. sp.; C: Sphaerobelum denticulatum n. sp.; D: Sphaerobelum spinatum n. sp.; E: Sphaerobelum lachneeis n. sp.; F: Sphaerobelum peterjaegeri n. sp.; G: Sphaerobelum nigrum n. sp.; H: Sphaerobelum splendidum n. sp.; I: Sphaerobelum laoticum n. sp. Not to scale." httpUri="https://zenodo.org/record/2601084/files/figure.png" pageId="15" pageNumber="216">Fig. 6C</figureCitation>
|
||
).
|
||
<emphasis id="B9363802E3058C3BA376F995FE4BF98F" bold="true" box="[268,418,1626,1651]" pageId="15" pageNumber="216">Endotergum</emphasis>
|
||
: inner section lacking any spines or setae. Middle area with a single row of small, circular cuticular impressions. Distance between impressions as wide as their diameter. Apically, 1–2 sparse rows of short marginal bristles, ending well before tergite margin. Bristles not smooth, but with numerous small spinicles.
|
||
<emphasis id="B9363802E3058C3BA718F96DFF34F923" bold="true" pageId="15" pageNumber="216">Anal shield</emphasis>
|
||
: slightly bell shaped, surface sparsely covered by tiny setae each standing in a tiny pit. Underside with a single, black locking carina, located close to tergite margin. Locking carinae
|
||
<date id="FFFCC2D0E3058C3BA18FF924FBF8F8FF" box="[1013,1041,1771,1795]" pageId="15" pageNumber="216">3x</date>
|
||
size of those of tergites.
|
||
<emphasis id="B9363802E3058C3BA750F925FA8BF8FF" bold="true" box="[1322,1378,1770,1795]" pageId="15" pageNumber="216">Legs</emphasis>
|
||
: legpair 1 with 3 or 4 ventral spines, leg-pair 2 with 5, leg-pair 3 with 6 or 7. First two leg-pairs without an apical spine. Leg-pairs 4–21 with 10 or 12 ventral spines and one dorso-apical spine. In leg 9 femur 1.2 times, tarsus 3.7 times longer than wide (
|
||
<figureCitation id="1379F895E3058C3BA311F899FE3EF893" box="[363,471,1878,1903]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="15" pageNumber="216">Fig. 13A</figureCitation>
|
||
). All podomeres densely setose. Coxa with a large and marginally toothed process. Coxa process absent at first leg and sharply projecting at second. Prefemur at apical margin with a projection laterally and mesally. Lateral projection triangular and sharply edged, juxtaposed to coxal process. Femur extended mesally into a dentate margin featuring 12–14 teeth. Prefemur also with ca. 6 teeth.
|
||
</paragraph>
|
||
<caption id="DF3DB498E31A8C24A2EDF855FB17F811" httpUri="https://zenodo.org/record/2601094/files/figure.png" pageId="16" pageNumber="217" startId="16.[151,250,1946,1968]" targetBox="[177,1416,193,1934]" targetPageId="16">
|
||
<paragraph id="8BFDE410E31A8C24A2EDF855FB17F811" blockId="16.[151,1436,1946,2029]" pageId="16" pageNumber="217">
|
||
<emphasis id="B9363802E31A8C24A2EDF855FEF3F84C" bold="true" box="[151,282,1946,1968]" pageId="16" pageNumber="217">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E31A8C24A352F855FDC2F84C" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[296,555,1946,1968]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="217" phylum="Arthropoda" rank="species" species="bolavensis" status="sp. nov.">
|
||
<emphasis id="B9363802E31A8C24A352F855FDC2F84C" box="[296,555,1946,1968]" italics="true" pageId="16" pageNumber="217">Sphaerobelum bolavensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E31A8C24A049F855FD84F84C" bold="true" box="[563,621,1946,1968]" pageId="16" pageNumber="217">
|
||
<taxonomicNameLabel id="A2058579E31A8C24A049F855FD84F84C" box="[563,621,1946,1968]" pageId="16" pageNumber="217" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (ZMUC);
|
||
<emphasis id="B9363802E31A8C24A139F855FCBCF84C" bold="true" box="[835,853,1946,1968]" pageId="16" pageNumber="217">A</emphasis>
|
||
: coxa of second left leg with gonopore;
|
||
<emphasis id="B9363802E31A8C24A68DF855FAE1F84C" bold="true" box="[1271,1288,1946,1968]" pageId="16" pageNumber="217">B</emphasis>
|
||
: right anterior telopod, anterior view;
|
||
<emphasis id="B9363802E31A8C24A3FBF877FE75F832" bold="true" box="[385,412,1976,1998]" pageId="16" pageNumber="217">C:</emphasis>
|
||
right anterior telopod, posterior view;
|
||
<emphasis id="B9363802E31A8C24A15BF877FCDAF832" bold="true" box="[801,819,1976,1998]" pageId="16" pageNumber="217">D</emphasis>
|
||
: right anterior telopod, lateral view;
|
||
<emphasis id="B9363802E31A8C24A6DFF877FB5CF832" bold="true" box="[1189,1205,1976,1998]" pageId="16" pageNumber="217">E</emphasis>
|
||
: left posterior telopod, anterior view;
|
||
<emphasis id="B9363802E31A8C24A350F818FED0F811" bold="true" box="[298,313,2007,2029]" pageId="16" pageNumber="217">F</emphasis>
|
||
: left posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E31A8C24A11BF818FC58F811" bold="true" box="[865,945,2007,2029]" pageId="16" pageNumber="217">Syn-Cx</emphasis>
|
||
= syncoxite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3DB498E31B8C25A2EDF87FFCB6F819" httpUri="https://zenodo.org/record/2601096/files/figure.png" pageId="17" pageNumber="218" startId="17.[151,250,1968,1990]" targetBox="[150,1436,250,1894]" targetPageId="17">
|
||
<paragraph id="8BFDE410E31B8C25A2EDF87FFCB6F819" blockId="17.[151,1437,1968,2021]" pageId="17" pageNumber="218">
|
||
<emphasis id="B9363802E31B8C25A2EDF87FFEF3F83A" bold="true" box="[151,282,1968,1990]" pageId="17" pageNumber="218">FIGURE 11</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E31B8C25A352F87FFDF2F83A" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[296,539,1968,1990]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="17" pageNumber="218" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
|
||
<emphasis id="B9363802E31B8C25A352F87FFDF2F83A" box="[296,539,1968,1990]" italics="true" pageId="17" pageNumber="218">Sphaerobelum phouloei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E31B8C25A059F87FFDB6F83A" bold="true" box="[547,607,1968,1990]" pageId="17" pageNumber="218">
|
||
<taxonomicNameLabel id="A2058579E31B8C25A059F87FFDB6F83A" box="[547,607,1968,1990]" pageId="17" pageNumber="218" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male holotype (ZMUC);
|
||
<emphasis id="B9363802E31B8C25A108F87FFC6DF83A" bold="true" box="[882,900,1968,1990]" pageId="17" pageNumber="218">A</emphasis>
|
||
: left first coxa with stigmatic plate;
|
||
<emphasis id="B9363802E31B8C25A779F87FFAFDF83A" bold="true" box="[1283,1300,1968,1990]" pageId="17" pageNumber="218">B</emphasis>
|
||
: 9th left leg. Abbreviations:
|
||
<emphasis id="B9363802E31B8C25A34EF800FEBAF819" bold="true" box="[308,339,1999,2021]" pageId="17" pageNumber="218">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E31B8C25A3D1F800FE37F819" bold="true" box="[427,478,1999,2021]" pageId="17" pageNumber="218">St-Pl</emphasis>
|
||
= stigmatic plate. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3DB498E3188C26A2EDF859FA6AF816" httpUri="https://zenodo.org/record/2601098/files/figure.png" pageId="18" pageNumber="219" startId="18.[151,250,1942,1964]" targetBox="[151,1436,247,1876]" targetPageId="18">
|
||
<paragraph id="8BFDE410E3188C26A2EDF859FA6AF816" blockId="18.[151,1436,1942,2026]" pageId="18" pageNumber="219">
|
||
<emphasis id="B9363802E3188C26A2EDF859FEF5F850" bold="true" box="[151,284,1942,1964]" pageId="18" pageNumber="219">FIGURE 12</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E3188C26A351F859FDF6F850" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[299,543,1942,1964]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="18" pageNumber="219" phylum="Arthropoda" rank="species" species="phouloei" status="sp. nov.">
|
||
<emphasis id="B9363802E3188C26A351F859FDF6F850" box="[299,543,1942,1964]" italics="true" pageId="18" pageNumber="219">Sphaerobelum phouloei</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3188C26A05DF859FD8AF850" bold="true" box="[551,611,1942,1964]" pageId="18" pageNumber="219">
|
||
<taxonomicNameLabel id="A2058579E3188C26A05DF859FD8AF850" box="[551,611,1942,1964]" pageId="18" pageNumber="219" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (ZMUC);
|
||
<emphasis id="B9363802E3188C26A145F859FCB8F850" bold="true" box="[831,849,1942,1964]" pageId="18" pageNumber="219">A</emphasis>
|
||
: coxa of second left leg with gonopore;
|
||
<emphasis id="B9363802E3188C26A77BF859FAFBF850" bold="true" box="[1281,1298,1942,1964]" pageId="18" pageNumber="219">B</emphasis>
|
||
: left anterior telopod, anterior view;
|
||
<emphasis id="B9363802E3188C26A3FEF87AFE77F837" bold="true" box="[388,414,1973,1995]" pageId="18" pageNumber="219">C:</emphasis>
|
||
left anterior telopod, posterior view;
|
||
<emphasis id="B9363802E3188C26A160F87AFCC5F837" bold="true" box="[794,812,1973,1995]" pageId="18" pageNumber="219">D</emphasis>
|
||
: left anterior telopod, lateral view;
|
||
<emphasis id="B9363802E3188C26A6EEF87AFB4CF837" bold="true" box="[1172,1189,1973,1995]" pageId="18" pageNumber="219">E</emphasis>
|
||
: right posterior telopod, anterior view;
|
||
<emphasis id="B9363802E3188C26A350F81BFED0F816" bold="true" box="[298,313,2004,2026]" pageId="18" pageNumber="219">F</emphasis>
|
||
: right posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E3188C26A115F81BFC67F816" bold="true" box="[879,910,2004,2026]" pageId="18" pageNumber="219">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E3188C26A19CF81BFBDFF816" bold="true" box="[998,1078,2004,2026]" pageId="18" pageNumber="219">Syn-Cx</emphasis>
|
||
= syncoxite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3DB498E3198C27A2EDF850FCAEF80E" httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="19" pageNumber="220" startId="19.[151,250,1951,1973]" targetBox="[191,1401,195,1934]" targetPageId="19">
|
||
<paragraph id="8BFDE410E3198C27A2EDF850FCAEF80E" blockId="19.[151,1437,1951,2034]" pageId="19" pageNumber="220">
|
||
<emphasis id="B9363802E3198C27A2EDF850FEF0F849" bold="true" box="[151,281,1951,1973]" pageId="19" pageNumber="220">FIGURE 13</emphasis>
|
||
.
|
||
<taxonomicName id="4C429F93E3198C27A35CF850FDABF849" authority="Wesener, 2019" authorityName="Wesener" authorityYear="2019" box="[294,578,1951,1973]" class="Diplopoda" family="Zephronidae" genus="Sphaerobelum" kingdom="Animalia" order="Sphaerotheriida" pageId="19" pageNumber="220" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
|
||
<emphasis id="B9363802E3198C27A35CF850FDABF849" box="[294,578,1951,1973]" italics="true" pageId="19" pageNumber="220">Sphaerobelum denticulatum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9363802E3198C27A033F850FD6AF849" bold="true" box="[585,643,1951,1973]" pageId="19" pageNumber="220">
|
||
<taxonomicNameLabel id="A2058579E3198C27A033F850FD6AF849" box="[585,643,1951,1973]" pageId="19" pageNumber="220" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype (MHNG);
|
||
<emphasis id="B9363802E3198C27A121F850FC84F849" bold="true" box="[859,877,1951,1973]" pageId="19" pageNumber="220">A</emphasis>
|
||
: 9th left leg;
|
||
<emphasis id="B9363802E3198C27A18DF850FBE1F849" bold="true" box="[1015,1032,1951,1973]" pageId="19" pageNumber="220">B</emphasis>
|
||
: left anterior telopod, anterior view;
|
||
<emphasis id="B9363802E3198C27A7FBF850FA75F849" bold="true" box="[1409,1436,1951,1973]" pageId="19" pageNumber="220">C:</emphasis>
|
||
left anterior telopod, posterior view;
|
||
<emphasis id="B9363802E3198C27A056F872FDD7F82F" bold="true" box="[556,574,1981,2003]" pageId="19" pageNumber="220">D</emphasis>
|
||
: left posterior telopod, anterior view;
|
||
<emphasis id="B9363802E3198C27A19CF872FC1EF82F" bold="true" box="[998,1015,1981,2003]" pageId="19" pageNumber="220">E</emphasis>
|
||
: left posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B9363802E3198C27A34EF813FEBAF80E" bold="true" box="[308,339,2012,2034]" pageId="19" pageNumber="220">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B9363802E3198C27A3D1F813FE12F80E" bold="true" box="[427,507,2012,2034]" pageId="19" pageNumber="220">Syn-Cx</emphasis>
|
||
= syncoxite. Scale bars = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFDE410E31E8C20A2BDFF58FE79FF4C" blockId="20.[151,1437,151,824]" box="[199,400,151,176]" pageId="20" pageNumber="221">Female unknown.</paragraph>
|
||
<paragraph id="8BFDE410E31E8C20A2BDFF73FD04FD13" blockId="20.[151,1437,151,824]" pageId="20" pageNumber="221">
|
||
<emphasis id="B9363802E31E8C20A2BDFF73FE0AFF29" bold="true" box="[199,483,188,213]" pageId="20" pageNumber="221">Male sexual characters</emphasis>
|
||
: gonopore covered with a single, undivided, circular, sclerotized plate.
|
||
<emphasis id="B9363802E31E8C20A74EFF73FF13FF04" bold="true" pageId="20" pageNumber="221">Anterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E31E8C20A373FF10FE7CFF04" box="[265,405,223,248]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="20" pageNumber="221">Figs 13B, C</figureCitation>
|
||
): consisting of 4 telopoditomeres above syncoxite. Telopoditomere 1 rectangular, as long as wide. Telopoditomere 2 large, almost as long as telopoditomere 3 and 4 combined. Process of telopoditomere 2 located posteriorly, visible mesally and apically in anterior view. Process wide, projecting to half of telopoditomere 4, apically with a well-rounded tip. Telopoditomere 3 cylindrical, straight, slightly shorter than telopoditomere 4. Telopoditomere 4 cylindrical, well-rounded, posterior side with a black sclerotized spot and three small, triangular spine. Telopoditomeres
|
||
<quantity id="4CBA49F5E31E8C20A3DDFE5BFE18FE51" box="[423,497,404,429]" metricMagnitude="-2" metricUnit="m" metricValue="6.35" metricValueMax="10.16" metricValueMin="2.54" pageId="20" pageNumber="221" unit="in" value="2.5" valueMax="4.0" valueMin="1.0">1–4 in</quantity>
|
||
anterior view covered with long setae. In posterior view all telopoditomeres with fewer setae.
|
||
<emphasis id="B9363802E31E8C20A356FE78FDE3FE2C" bold="true" box="[300,522,439,464]" pageId="20" pageNumber="221">Posterior telopods</emphasis>
|
||
(
|
||
<figureCitation id="1379F895E31E8C20A067FE78FD58FE2C" box="[541,689,439,464]" captionStart="FIGURE 13" captionStartId="19.[151,250,1951,1973]" captionTargetBox="[191,1401,195,1934]" captionTargetId="figure@19.[191,1401,193,1934]" captionTargetPageId="19" captionText="FIGURE 13. Sphaerobelum denticulatum n. sp., holotype (MHNG); A: 9th left leg; B: left anterior telopod, anterior view; C: left anterior telopod, posterior view; D: left posterior telopod, anterior view; E: left posterior telopod, posterior view. Abbreviations: Cx = coxa; Syn-Cx = syncoxite. Scale bars = 1 mm." httpUri="https://zenodo.org/record/2601100/files/figure.png" pageId="20" pageNumber="221">Figs 13D, E</figureCitation>
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): telopoditomere 1 slightly longer than wide. Immovable finger (process of telopoditomere 2) longer than movable finger, consisting of telopoditomeres 3 and 4. Immovable finger with a characteristic, distally swollen apex, well rounded, apex therefore wider than base. Immovable finger in anterior view with a large spine, at tip opposite to tip of telopoditomere 4 with sclerotized spots. Telopoditomere 3 rectangular, towards immoveable finger with a triangular expansion carrying two spines. Telopoditomere 4 reaching 2/3 of length of telopoditomere 3, much more slender, apically strongly tapering into acute tip, curved towards immovable finger. Inner margin with single spine towards immovable finger. Telopoditomere 1 at both sides covered by setae. Telopoditomeres 2 and
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||
<quantity id="4CBA49F5E31E8C20A0D5FD7BFD32FD31" box="[687,731,692,717]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="20" pageNumber="221" unit="in" value="3.0">3 in</quantity>
|
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anterior view covered by long setae, in posterior view mostly glabrous. Telopoditomere 4 only with marginal setae.
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C358B79BE31E8C20A2BDFD33FEEDFCC4" pageId="20" pageNumber="221" type="etymology">
|
||
<paragraph id="8BFDE410E31E8C20A2BDFD33FEEDFCC4" blockId="20.[151,1437,151,824]" pageId="20" pageNumber="221">
|
||
<emphasis id="B9363802E31E8C20A2BDFD33FE4AFCE9" bold="true" box="[199,419,764,789]" pageId="20" pageNumber="221">Derivatio nominis</emphasis>
|
||
: denticulatum, noun in apposition, after the dentate mesal margin of the femur and prefemur.
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||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |