546 lines
73 KiB
XML
546 lines
73 KiB
XML
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<mods:title id="1A98642CF154C6D2B6B785A3D6B9E2B4">Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species</mods:title>
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<mods:affiliation id="58442A67800E087D8A6844522F2D542A">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<mods:affiliation id="769217210F5CB71B3033EF32D42AEDE9">Red de Ecoetología, Instituto de Ecología, A. C. Carretera antigua a Coatepec 351, El Haya, Xalapa, Veracruz 91073, Mexico.</mods:affiliation>
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<treatment id="03CA87F2FFD8FFE6FD8D4386BE983DD2" ID-DOI="http://doi.org/10.5281/zenodo.4726237" ID-GBIF-Taxon="181753775" ID-Zenodo-Dep="4726237" LSID="urn:lsid:plazi:treatment:03CA87F2FFD8FFE6FD8D4386BE983DD2" httpUri="http://treatment.plazi.org/id/03CA87F2FFD8FFE6FD8D4386BE983DD2" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
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<emphasis id="B917EAF6FFD8FFE7FD8D4386BD8F3B9D" bold="true" box="[549,765,397,424]" italics="true" pageId="38" pageNumber="39">Phanaeus porioni</emphasis>
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<bibRefCitation id="EFF24B15FFD8FFE7FCAB4386BCD73B92" author="Arnaud P." box="[771,933,397,424]" pageId="38" pageNumber="39" pagination="2 - 8" refId="ref35808" refString="Arnaud P. 2001. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 6: 2 - 8." type="journal article" year="2001">Arnaud, 2001</bibRefCitation>
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<figureCitation id="13582A61FFD8FFE7FDC043BEBDBF3BF5" box="[616,717,437,463]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="38" pageNumber="39">Figs 1M</figureCitation>
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,
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<figureCitation id="13582A61FFD8FFE7FD7143BEBD873BF5" box="[729,757,437,463]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="38" pageNumber="39">2I</figureCitation>
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<figureCitation id="13582A61FFD8FFE7FCA943BEBC6D3BF5" box="[769,799,437,463]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="38" pageNumber="39">11</figureCitation>
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,
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<figureCitation id="13582A61FFD8FFE7FC8443BEBC393BEA" box="[812,843,437,464]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="38" pageNumber="39">15</figureCitation>
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,
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<figureCitation id="13582A61FFD8FFE7FCF043BEBCFE3BF5" box="[856,908,437,463]" captionStart="Fig" captionStartId="55.[189,232,1681,1707]" captionTargetBox="[258,1329,264,1646]" captionTargetId="figure-157@55.[258,1329,264,1646]" captionTargetPageId="55" captionText="Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836802" httpUri="https://zenodo.org/record/4836802/files/figure.png" pageId="38" pageNumber="39">18N</figureCitation>
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,
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<figureCitation id="13582A61FFD8FFE7FC3143BEBCB93BF5" box="[921,971,437,463]" captionStart="Fig" captionStartId="56.[189,231,1678,1704]" captionTargetBox="[306,1282,265,1646]" captionTargetId="figure-158@56.[306,1282,265,1646]" captionTargetPageId="56" captionText="Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017." figureDoi="http://doi.org/10.5281/zenodo.4836804" httpUri="https://zenodo.org/record/4836804/files/figure.png" pageId="38" pageNumber="39">19N</figureCitation>
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<emphasis id="B917EAF6FFD8FFE7FF1543F0BE43382F" box="[189,305,507,533]" italics="true" pageId="38" pageNumber="39">Phanaeus</emphasis>
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(
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<emphasis id="B917EAF6FFD8FFE7FEEA43F0BE86382F" box="[322,500,507,533]" italics="true" pageId="38" pageNumber="39">Notiophanaeus</emphasis>
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)
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<emphasis id="B917EAF6FFD8FFE7FDAC43F0BDA6382F" box="[516,724,507,533]" italics="true" pageId="38" pageNumber="39">endymion porioni</emphasis>
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<emphasis id="B917EAF6FFD8FFE7FF15404ABE433861" box="[189,305,577,603]" italics="true" pageId="38" pageNumber="39">Phanaeus</emphasis>
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(
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<emphasis id="B917EAF6FFD8FFE7FEEA404ABE863861" box="[322,500,577,603]" italics="true" pageId="38" pageNumber="39">Notiophanaeus</emphasis>
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)
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<emphasis id="B917EAF6FFD8FFE7FDAC404ABDA63861" box="[516,724,577,603]" italics="true" pageId="38" pageNumber="39">endymion porioni</emphasis>
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–
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<bibRefCitation id="EFF24B15FFD8FFE7FD58404ABCB73866" author="Arnaud P." box="[752,965,577,604]" pageId="38" pageNumber="39" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 94</bibRefCitation>
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(as subspecies of
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||
<taxonomicName id="4C634D67FFD8FFE7FB334049BA583861" ID-CoL="4FSJL" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[1179,1322,577,603]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD8FFE7FB334049BA583861" box="[1179,1322,577,603]" italics="true" pageId="38" pageNumber="39">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C634D67FFD8FFE7FF154062BD7A38B9" ID-CoL="874DL" authorityName="Arnaud" authorityYear="2001" box="[189,520,617,643]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="subSpecies" species="endymion" subSpecies="porioni">
|
||
<emphasis id="B917EAF6FFD8FFE7FF154062BD7A38B9" box="[189,520,617,643]" italics="true" pageId="38" pageNumber="39">Phanaeus endymion porioni</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FD8D4062BD8938BE" author="Arnaud P." box="[549,763,617,644]" page="95" pageId="38" pageNumber="39" year="2002">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FD8D4062BD8938BE" author="Arnaud P." box="[549,763,617,644]" pageId="38" pageNumber="39" refId="ref35859" refString="Arnaud P. 2002 b. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus. Hillside Books, Canterbury." type="book" year="2002">Arnaud 2002b: 95</bibRefCitation>
|
||
</treatmentCitation>
|
||
(as subspecies of
|
||
<taxonomicName id="4C634D67FFD8FFE7FC7B4061BB1038B9" ID-CoL="4FSJL" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[979,1122,617,643]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD8FFE7FC7B4061BB1038B9" box="[979,1122,617,643]" italics="true" pageId="38" pageNumber="39">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
). —
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FB084062BA0438BE" author="Krajcik M." box="[1184,1398,617,644]" page="150" pageId="38" pageNumber="39" year="2006">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FB084062BA0438BE" author="Krajcik M." box="[1184,1398,617,644]" pageId="38" pageNumber="39" pagination="1 - 189" refId="ref39668" refString="Krajcik M. 2006. Checklist of Scarabaeoidea of the World. 1. Scarabaeinae (Coleoptera: Scarabaeidae: Scarabaeinae). Animma. X. Suplement 3: 1 - 189." type="journal article" year="2006">Krajcik 2006: 150</bibRefCitation>
|
||
</treatmentCitation>
|
||
(as subspecies of
|
||
<taxonomicName id="4C634D67FFD8FFE7FE104099BD353891" ID-CoL="4FSJL" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[440,583,657,683]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD8FFE7FE104099BD353891" box="[440,583,657,683]" italics="true" pageId="38" pageNumber="39">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
). —
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FD2C409ABA183896" box="[644,1386,657,684]" pageId="38" pageNumber="39">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FD2C409ABCCC3891" author="Edmonds W. D. & Zidek J." box="[644,958,657,683]" pageId="38" pageNumber="39" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 1</bibRefCitation>
|
||
, 8, 13 (as synonym of
|
||
<taxonomicName id="4C634D67FFD8FFE7FB624099BA2B3891" ID-CoL="4FSJL" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[1226,1369,657,683]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD8FFE7FB624099BA2B3891" box="[1226,1369,657,683]" italics="true" pageId="38" pageNumber="39">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF1540D3BEF83953" blockId="38.[189,1399,727,953]" pageId="38" pageNumber="39">
|
||
<treatmentCitationGroup id="AB7311CAFFD8FFE7FF1540D3BEF83953" pageId="38" pageNumber="39">
|
||
Non
|
||
<taxonomicName id="4C634D67FFD8FFE7FF5340DCBDCF38CB" ID-CoL="4FSJL" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[251,701,727,754]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion" subGenus="Notiophanaeus">
|
||
<emphasis id="B917EAF6FFD8FFE7FF5340DCBE1D38CB" box="[251,367,727,753]" italics="true" pageId="38" pageNumber="39">Phanaeus</emphasis>
|
||
(
|
||
<emphasis id="B917EAF6FFD8FFE7FE2C40DCBD4438CB" box="[388,566,727,753]" italics="true" pageId="38" pageNumber="39">Notiophanaeus</emphasis>
|
||
)
|
||
<emphasis id="B917EAF6FFD8FFE7FDE340DCBDCF38CB" box="[587,701,727,753]" italics="true" pageId="38" pageNumber="39">endymion</emphasis>
|
||
</taxonomicName>
|
||
(error) –
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FC9340DCBC5C3923" pageId="38" pageNumber="39">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FC9340DCBB6738CB" author="Edmonds W. D." box="[827,1045,727,753]" pageId="38" pageNumber="39" pagination="1 - 105" refId="ref37487" refString="Edmonds W. D. 1994. Revision of Phanaeus Macleay, a New World genus of Scarabaeinae dung beetles (Coleoptera: Scarabaeidae, Scarabaeinae). Contributions in Science, Natural History Museum of Los Angeles County 443: 1 - 105." type="journal article" year="1994">Edmonds 1994: 2</bibRefCitation>
|
||
, 8–9, 12, 19, 36, 39, 41–46, 54, 74, 101, figs 17, 209, 215–216, 221 (in part).
|
||
</treatmentCitation>
|
||
—
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FCF440F4BC30397A" pageId="38" pageNumber="39">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FCF440F4BBE83923" author="Edmonds W. D. & Zidek J." box="[860,1178,767,793]" pageId="38" pageNumber="39" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 3</bibRefCitation>
|
||
, 13, figs 132–133, 136, 140, 143, 152–155 (in part).
|
||
<taxonomicName id="4C634D67FFD8FFE7FDD9412CBC30397A" ID-CoL="874DL" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[625,834,807,833]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD8FFE7FDD9412CBC30397A" box="[625,834,807,833]" italics="true" pageId="38" pageNumber="39">Phanaeus porioni</emphasis>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
–
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FCC9412CBEF83953" pageId="38" pageNumber="39">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FCC9412CBBEF397B" author="Edmonds W. D. & Zidek J." box="[865,1181,807,833]" pageId="38" pageNumber="39" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 3</bibRefCitation>
|
||
, 6 (as synonym of
|
||
<taxonomicName id="4C634D67FFD8FFE7FF434144BE083952" ID-CoL="4FSJL" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[235,378,846,872]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD8FFE7FF434144BE083952" box="[235,378,846,872]" italics="true" pageId="38" pageNumber="39">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF15417CBE5B3983" blockId="38.[189,1399,727,953]" pageId="38" pageNumber="39">
|
||
<treatmentCitationGroup id="AB7311CAFFD8FFE7FF15417CBE5B3983" pageId="38" pageNumber="39">
|
||
Non
|
||
<taxonomicName id="4C634D67FFD8FFE7FF5F417DBE9139AA" ID-CoL="4FSJL" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[247,483,886,912]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD8FFE7FF5F417DBE9139AA" box="[247,483,886,912]" italics="true" pageId="38" pageNumber="39">Phanaeus endymion</emphasis>
|
||
</taxonomicName>
|
||
(error) –
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FDFB417DBCF339AA" author="Creedy T. J. & Mann D. J." box="[595,897,886,913]" page="34" pageId="38" pageNumber="39" year="2011">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FDFB417DBCF339AA" author="Creedy T. J. & Mann D. J." box="[595,897,886,913]" pageId="38" pageNumber="39" refId="ref36744" refString="Creedy T. J. & Mann D. J. 2011. Identification Guide to the Scarabaeinae Dung Beetles of Cusuco National Park, Honduras. Version 1.0. Operation Wallacea, Lincolnshire." type="book" year="2011">Creedy & Mann 2011: 34</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 51. —
|
||
<treatmentCitation id="0AC210F5FFD8FFE7FC75417DBA6739AA" author="Edmonds W. D. & Zidek J." box="[989,1301,886,912]" page="1" pageId="38" pageNumber="39" year="2012">
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FC75417DBA6739AA" author="Edmonds W. D. & Zidek J." box="[989,1301,886,912]" pageId="38" pageNumber="39" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek 2012: 1</bibRefCitation>
|
||
</treatmentCitation>
|
||
, 5–6 (in part).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFD8FFE7FF1541EFBBE43EEB" pageId="38" pageNumber="39" type="diagnosis">
|
||
<paragraph id="8BDC36E4FFD8FFE7FF1541EFBE4439C4" blockId="38.[189,310,996,1022]" box="[189,310,996,1022]" pageId="38" pageNumber="39">
|
||
<heading id="D0948188FFD8FFE7FF1541EFBE4439C4" bold="true" box="[189,310,996,1022]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
|
||
<emphasis id="B917EAF6FFD8FFE7FF1541EFBE4439C4" bold="true" box="[189,310,996,1022]" pageId="38" pageNumber="39">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF15461CBBE43EEB" blockId="38.[187,1399,1047,1233]" pageId="38" pageNumber="39">
|
||
<taxonomicName id="4C634D67FFD8FFE7FF15461CBEFD3E0B" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[189,399,1047,1073]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD8FFE7FF15461CBEFD3E0B" box="[189,399,1047,1073]" italics="true" pageId="38" pageNumber="39">Phanaeus porioni</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D67FFD8FFE7FE654613BD2F3E0B" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[461,605,1047,1073]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD8FFE7FE654613BD2F3E0B" box="[461,605,1047,1073]" italics="true" pageId="38" pageNumber="39">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
are closely related, but the former is easily separated by the major female with the pronotal process weakly or not concave posteriorly and pronotal tubercles nearly aligned (
|
||
<figureCitation id="13582A61FFD8FFE7FF6C466CBE403EBB" box="[196,306,1127,1153]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="38" pageNumber="39">Fig. 11B</figureCitation>
|
||
), while the males are distinguished by the endophallite copulatrix (
|
||
<figureCitation id="13582A61FFD8FFE7FBDD466CBBCD3EBB" box="[1141,1215,1127,1153]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="38" pageNumber="39">Fig. 1</figureCitation>
|
||
). Additionally,
|
||
<taxonomicName id="4C634D67FFD8FFE7FF154684BE433E92" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[189,305,1167,1192]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD8FFE7FF154684BE433E92" box="[189,305,1167,1192]" italics="true" pageId="38" pageNumber="39">P. porioni</emphasis>
|
||
</taxonomicName>
|
||
is recognized by the elytral striae always distinctly punctate (
|
||
<figureCitation id="13582A61FFD8FFE7FBBD4684BB193E93" box="[1045,1131,1167,1193]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="38" pageNumber="39">Fig. 11</figureCitation>
|
||
), but the strial surface never roughened as in
|
||
<taxonomicName id="4C634D67FFD8FFE7FE6F46BCBD393EEB" authority="Moctezuma & Halffter, 2021" authorityName="Moctezuma & Halffter" authorityYear="2021" box="[455,587,1207,1233]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="pacificus" status="sp. nov.">
|
||
<emphasis id="B917EAF6FFD8FFE7FE6F46BCBD393EEB" box="[455,587,1207,1233]" italics="true" pageId="38" pageNumber="39">P. pacificus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFD8FFE7FDFB46BCBDDC3EEB" box="[595,686,1207,1233]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13582A61FFD8FFE7FD1546BDBC713EEB" box="[701,771,1206,1233]" captionStart="Fig" captionStartId="34.[189,232,1997,2023]" captionTargetBox="[211,1370,749,1965]" captionTargetId="figure-104@34.[189,1399,749,1965]" captionTargetPageId="34" captionText="Fig. 9. Phanaeus pacificus sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836782" httpUri="https://zenodo.org/record/4836782/files/figure.png" pageId="38" pageNumber="39">Fig. 9</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C634D67FFD8FFE7FCED46BCBCBC3EEA" authority="Moctezuma & Halffter, 2021" authorityName="Moctezuma & Halffter" authorityYear="2021" box="[837,974,1206,1232]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="jackenioi" status="sp. nov.">
|
||
<emphasis id="B917EAF6FFD8FFE7FCED46BCBCBC3EEA" box="[837,974,1206,1232]" italics="true" pageId="38" pageNumber="39">P. jackenioi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A224578DFFD8FFE7FC7D46BCBB423EEB" box="[981,1072,1207,1233]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13582A61FFD8FFE7FB9746BDBBF53EEB" box="[1087,1159,1206,1233]" captionStart="Fig" captionStartId="24.[189,232,1998,2024]" captionTargetBox="[217,1394,753,1972]" captionTargetId="figure-104@24.[189,1399,753,1972]" captionTargetPageId="24" captionText="Fig. 6. Phanaeus jackenioi sp. nov. A. Holotype, ♂ (UVGC). B. ♀ (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836768" httpUri="https://zenodo.org/record/4836768/files/figure.png" pageId="38" pageNumber="39">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFD8FFE7FF1546F6BD343DD2" pageId="38" pageNumber="39" type="materials_examined">
|
||
<paragraph id="8BDC36E4FFD8FFE7FF1546F6BE1E3F2C" blockId="38.[189,364,1276,1303]" box="[189,364,1276,1303]" pageId="38" pageNumber="39">
|
||
<heading id="D0948188FFD8FFE7FF1546F6BE1E3F2C" bold="true" box="[189,364,1276,1303]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
|
||
<emphasis id="B917EAF6FFD8FFE7FF1546F6BE1E3F2C" bold="true" box="[189,364,1276,1303]" pageId="38" pageNumber="39">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF42473BBDAD3F70" blockId="38.[189,1399,1328,1474]" box="[234,735,1328,1354]" pageId="38" pageNumber="39">
|
||
<emphasis id="B917EAF6FFD8FFE7FF42473BBE2E3F70" bold="true" box="[234,348,1328,1354]" pageId="38" pageNumber="39">
|
||
<typeStatus id="54D88846FFD8FFE7FF42473BBE2E3F70" box="[234,348,1328,1354]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
(studied from photographs,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FD02473BBDA53F70" box="[682,727,1328,1354]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF154753BDBB3FFB" blockId="38.[189,1399,1328,1474]" pageId="38" pageNumber="39">
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FF154753BDB73FFB" ID-GBIF-Occurrence="3093980609" collectionCode="CPFA" country="Honduras" elevation="250" location="Rco Bonito" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Atlantida" typeStatus="holotype">
|
||
<collectingCountry id="F3747674FFD8FFE7FF154753BE123F48" box="[189,352,1368,1394]" name="Honduras" pageId="38" pageNumber="39">HONDURAS</collectingCountry>
|
||
•
|
||
<specimenCount id="9D65FD6DFFD8FFE7FE2C4753BEEE3F48" box="[388,412,1368,1394]" pageId="38" pageNumber="39" type="male">♂</specimenCount>
|
||
,
|
||
<bibRefCitation id="EFF24B15FFD8FFE7FE05475CBD023F48" author="Arnaud P." box="[429,624,1367,1394]" pageId="38" pageNumber="39" pagination="2 - 8" refId="ref35808" refString="Arnaud P. 2001. Description de nouvelles especes de Phanaeides. (Col. Scarabaeidae). Besoiro 6: 2 - 8." type="journal article" year="2001">Arnaud 2001: 4</bibRefCitation>
|
||
(
|
||
<figureCitation id="13582A61FFD8FFE7FD2E475CBD873F48" box="[646,757,1367,1394]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="38" pageNumber="39">Fig. 11C</figureCitation>
|
||
);
|
||
<collectingRegion id="49A7F806FFD8FFE7FCA7475CBC0D3F4B" box="[783,895,1367,1393]" country="Honduras" name="Atlantida" pageId="38" pageNumber="39">Atlántida</collectingRegion>
|
||
,
|
||
<location id="8EBC603FFFD8FFE7FC394753BB533F4B" LSID="urn:lsid:plazi:treatment:03CA87F2FFD8FFE6FD8D4386BE983DD2:8EBC603FFFD8FFE7FC394753BB533F4B" box="[913,1057,1367,1394]" country="Honduras" name="Rco Bonito" pageId="38" pageNumber="39" stateProvince="Atlantida">Rco Bonito</location>
|
||
; “
|
||
<emphasis id="B917EAF6FFD8FFE7FB97475CBA663F4B" box="[1087,1300,1367,1393]" italics="true" pageId="38" pageNumber="39">
|
||
<taxonomicName id="4C634D67FFD8FFE7FB97475CBA7D3F4B" box="[1087,1295,1367,1393]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="endym">Phanaeus endym</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
<taxonomicName id="4C634D67FFD8FFE7FA884753BE9E3FA3" authority="P. ARNAUD DET 2001" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD8FFE7FA884753BA043F4B" box="[1312,1398,1368,1393]" italics="true" pageId="38" pageNumber="39">porioni</emphasis>
|
||
P.ARNAUD DET 2001
|
||
</taxonomicName>
|
||
HOLOTYPE ♂ /HONDURAS -ATLANTIDA, Rco Bonito (
|
||
<quantity id="4C9B9B01FFD8FFE7FAA44774BA1B3FA0" box="[1292,1385,1407,1434]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="m" value="250.0">
|
||
<elevation id="004ED1D7FFD8FFE7FAA44774BA1B3FA0" box="[1292,1385,1407,1434]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
), juilliet-1995, Thierry PORION Leg”;
|
||
<collectionCode id="ED72AE21FFD8FFE7FDD247ACBDB73FFB" box="[634,709,1447,1473]" collectionName="P and F Arnaud" pageId="38" pageNumber="39">CPFA</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF4247E6BD383C32" blockId="38.[189,1395,1517,1584]" box="[234,586,1517,1544]" pageId="38" pageNumber="39">
|
||
<emphasis id="B917EAF6FFD8FFE7FF4247E6BEBA3C3D" bold="true" box="[234,456,1517,1543]" pageId="38" pageNumber="39">
|
||
<typeStatus id="54D88846FFD8FFE7FF4247E6BE153C3D" box="[234,359,1517,1543]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
|
||
revised
|
||
</emphasis>
|
||
(
|
||
<specimenCount id="9D65FD6DFFD8FFE7FE7347E6BD753C32" box="[475,519,1517,1544]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FDBC47E6BD4C3C32" box="[532,574,1517,1544]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF15441EBA013C15" blockId="38.[189,1395,1517,1584]" box="[189,1395,1556,1584]" pageId="38" pageNumber="39">
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FF15441EBA1D3C15" ID-GBIF-Occurrence="3093980405" box="[189,1391,1556,1584]" collectionCode="VMC" collectorName="Thierry Porion" country="Honduras" elevation="250" location="Rco Bonito" pageId="38" pageNumber="39" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Atlantida">
|
||
<collectingCountry id="F3747674FFD8FFE7FF15441EBE123C15" box="[189,352,1557,1583]" name="Honduras" pageId="38" pageNumber="39">HONDURAS</collectingCountry>
|
||
–
|
||
<emphasis id="B917EAF6FFD8FFE7FED5441FBE863C15" bold="true" box="[381,500,1556,1583]" pageId="38" pageNumber="39">
|
||
<collectingRegion id="49A7F806FFD8FFE7FED5441FBE863C15" box="[381,500,1556,1583]" country="Honduras" name="Atlantida" pageId="38" pageNumber="39">Atlántida</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D65FD6DFFD8FFE7FDA5441EBD493C15" box="[525,571,1557,1583]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FDE0441EBD073C15" box="[584,629,1557,1583]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
|
||
; “
|
||
<location id="8EBC603FFFD8FFE7FD26441EBC653C15" LSID="urn:lsid:plazi:treatment:03CA87F2FFD8FFE6FD8D4386BE983DD2:8EBC603FFFD8FFE7FD26441EBC653C15" box="[654,791,1557,1583]" country="Honduras" name="Rco Bonito" pageId="38" pageNumber="39" stateProvince="Atlantida">Rco Bonito</location>
|
||
(
|
||
<quantity id="4C9B9B01FFD8FFE7FC8E441EBC073C15" box="[806,885,1557,1584]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="m" value="250.0">
|
||
<elevation id="004ED1D7FFD8FFE7FC8E441EBC073C15" box="[806,885,1557,1584]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="38" pageNumber="39" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
), juilliet-1995,
|
||
<collectorName id="26965332FFD8FFE7FB82441EBBA53C15" box="[1066,1239,1557,1583]" pageId="38" pageNumber="39">Thierry Porion</collectorName>
|
||
Leg”;
|
||
<collectionCode id="ED72AE21FFD8FFE7FA80441EBA1D3C15" box="[1320,1391,1557,1583]" pageId="38" pageNumber="39">VMC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF154450BDCD3C4F" blockId="38.[189,703,1627,1653]" box="[189,703,1627,1653]" pageId="38" pageNumber="39">
|
||
<emphasis id="B917EAF6FFD8FFE7FF154450BD703C4F" bold="true" box="[189,514,1627,1653]" pageId="38" pageNumber="39">Non-type material revised</emphasis>
|
||
(
|
||
<specimenCount id="9D65FD6DFFD8FFE7FDBB4450BD143C4F" box="[531,614,1627,1653]" pageId="38" pageNumber="39" type="male">14 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FDDB4450BDC13C4F" box="[627,691,1627,1653]" pageId="38" pageNumber="39" type="female">9 ♀♀</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF154485BDF53D55" blockId="38.[189,1399,1677,1904]" pageId="38" pageNumber="39">
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FF154485BD8E3CEA" ID-GBIF-Occurrence="3093980458" collectingDate="1996-07" collectingMethod="intercept trap in cocoa plantation" collectionCode="TAMU" collectorName="Coll. R Lehmann. Flight" country="Honduras" location="Atlantida" pageId="38" pageNumber="39" specimenCount="155" specimenCount-female="2" specimenCount-male="3" stateProvince="Atlantida">
|
||
<collectingCountry id="F3747674FFD8FFE7FF154485BE123C92" box="[189,352,1678,1704]" name="Honduras" pageId="38" pageNumber="39">HONDURAS</collectingCountry>
|
||
–
|
||
<emphasis id="B917EAF6FFD8FFE7FE2B4486BE883C92" bold="true" box="[387,506,1677,1704]" pageId="38" pageNumber="39">
|
||
<collectingRegion id="49A7F806FFD8FFE7FE2B4486BE883C92" box="[387,506,1677,1704]" country="Honduras" name="Atlantida" pageId="38" pageNumber="39">Atlántida</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D65FD6DFFD8FFE7FDBC4485BD283C93" box="[532,602,1678,1705]" pageId="38" pageNumber="39" type="male">3 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FDC04485BDDF3C93" box="[616,685,1678,1705]" pageId="38" pageNumber="39" type="female">2 ♀♀</specimenCount>
|
||
; “
|
||
<locationDeviation id="9A6E2FC6FFD8FFE7FD614485BCC83C92" box="[713,954,1678,1705]" pageId="38" pageNumber="39">
|
||
<quantity id="4C9B9B01FFD8FFE7FD614485BC683C92" box="[713,794,1678,1704]" metricMagnitude="4" metricUnit="m" metricValue="1.3" pageId="38" pageNumber="39" unit="km" value="13.0">13 km</quantity>
|
||
. W La Ceiba
|
||
</locationDeviation>
|
||
.
|
||
<date id="FFDD1024FFD8FFE7FC6F4485BB453C92" box="[967,1079,1678,1704]" pageId="38" pageNumber="39" value="1996-07">
|
||
<collectingDate id="EF99E9CCFFD8FFE7FC6F4485BB453C92" box="[967,1079,1678,1704]" pageId="38" pageNumber="39" value="1996-07">
|
||
VII-
|
||
<quantity id="4C9B9B01FFD8FFE7FC534485BB453C92" box="[1019,1079,1678,1704]" metricMagnitude="6" metricUnit="m" metricValue="1.99615" pageId="38" pageNumber="39" unit="m" value="1996150.0">1996</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FBEC4485BBEA3C93" box="[1092,1176,1678,1705]" pageId="38" pageNumber="39" type="generic">150 m.</specimenCount>
|
||
<collectorName id="26965332FFD8FFE7FB094485BE713CEA" pageId="38" pageNumber="39">Coll. R Lehmann. Flight</collectorName>
|
||
<collectingMethod id="52224EF3FFD8FFE7FEA244BDBDFC3CEA" box="[266,654,1718,1744]" pageId="38" pageNumber="39">intercept trap in cocoa plantation</collectingMethod>
|
||
”;
|
||
<collectionCode id="ED72AE21FFD8FFE7FD0044BDBD8E3CEA" box="[680,764,1718,1744]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="38" pageNumber="39">TAMU</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FCBC44BDBA173CEA" ID-GBIF-Occurrence="3093980466" box="[788,1381,1718,1744]" collectingDate="1996-07" collectingMethod="intercept trap in cocoa plantation" collectionCode="UVGC" collectorName="Coll. R Lehmann. Flight" country="Honduras" location="Atlantida" pageId="38" pageNumber="39" specimenCount="1" specimenCount-female="1" stateProvince="Atlantida">
|
||
<specimenCount id="9D65FD6DFFD8FFE7FCBC44BDBC333CEA" box="[788,833,1718,1744]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFD8FFE7FABA44BDBA173CEA" box="[1298,1381,1718,1744]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="38" pageNumber="39">UVGC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FF1544D5BC733CC2" ID-GBIF-Occurrence="3093980436" box="[189,769,1758,1784]" collectingDate="1996-07" collectingMethod="intercept trap in cocoa plantation" collectionCode="VMC" collectorName="Coll. R Lehmann. Flight" country="Honduras" location="Atlantida" pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Atlantida">
|
||
<specimenCount id="9D65FD6DFFD8FFE7FF1544D5BF9E3CC2" box="[189,236,1758,1784]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFD8FFE7FD1644D5BC733CC2" box="[702,769,1758,1784]" pageId="38" pageNumber="39">VMC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FCB244D5BC8E3D1A" ID-GBIF-Occurrence="3093980461" collectedFrom="tropical rainforest" collectingDate="1996-06-15" collectingDateMax="1996-06-19" collectingDateMin="1996-06-15" collectingMethod="intercept trap" collectionCode="IEXA" collectorName="Coll. R. Lehmann. Flight" country="Honduras" location="La Ceiba." pageId="38" pageNumber="39" specimenCount="177" specimenCount-female="1" specimenCount-male="1" stateProvince="Atlantida">
|
||
<specimenCount id="9D65FD6DFFD8FFE7FCB244D5BC3A3CC2" box="[794,840,1758,1784]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FCFD44D5BCF13CC2" box="[853,899,1758,1784]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
|
||
; “
|
||
<locationDeviation id="9A6E2FC6FFD8FFE7FC3544D5BBFB3CC2" box="[925,1161,1758,1785]" pageId="38" pageNumber="39">
|
||
<quantity id="4C9B9B01FFD8FFE7FC3544D5BC9E3CC2" box="[925,1004,1758,1785]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="38" pageNumber="39" unit="km" value="15.0">15 km</quantity>
|
||
. W La Ceiba
|
||
</locationDeviation>
|
||
.
|
||
<date id="FFDD1024FFD8FFE7FB3C44D5BA4C3CC3" box="[1172,1342,1758,1785]" pageId="38" pageNumber="39" value="1996-06-15" valueMax="1996-06-19" valueMin="1996-06-15">
|
||
<collectingDate id="EF99E9CCFFD8FFE7FB3C44D5BA4C3CC3" box="[1172,1342,1758,1785]" pageId="38" pageNumber="39" value="1996-06-15" valueMax="1996-06-19" valueMin="1996-06-15">
|
||
VI15-19-
|
||
<quantity id="4C9B9B01FFD8FFE7FB5744D5BA4C3CC3" box="[1279,1342,1758,1785]" metricMagnitude="6" metricUnit="m" metricValue="1.996175" pageId="38" pageNumber="39" unit="m" value="1996175.0">1996</quantity>
|
||
</collectingDate>
|
||
</date>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FAE244D5BFA93D1A" pageId="38" pageNumber="39" type="generic">175 m.</specimenCount>
|
||
<collectorName id="26965332FFD8FFE7FF4C450EBD7D3D25" box="[228,527,1797,1824]" pageId="38" pageNumber="39">Coll. R. Lehmann. Flight</collectorName>
|
||
<collectingMethod id="52224EF3FFD8FFE7FDB0450EBDCA3D1A" box="[536,696,1797,1824]" pageId="38" pageNumber="39">intercept trap</collectingMethod>
|
||
,
|
||
<collectedFrom id="387F7519FFD8FFE7FD6D450EBCEC3D25" box="[709,926,1797,1823]" pageId="38" pageNumber="39">tropical rainforest</collectedFrom>
|
||
”;
|
||
<collectionCode id="ED72AE21FFD8FFE7FC1E450DBC8E3D1A" box="[950,1020,1798,1824]" pageId="38" pageNumber="39">IEXA</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FBBE450DBD693D72" ID-GBIF-Occurrence="3093980305" collectedFrom="tropical rainforest" collectingDate="1996-06-15" collectingDateMax="1996-06-19" collectingDateMin="1996-06-15" collectingMethod="intercept trap" collectionCode="TAMU" collectorName="Coll. R. Lehmann. Flight" country="Honduras" location="La Ceiba." pageId="38" pageNumber="39" specimenCount="10" specimenCount-female="3" specimenCount-male="7" stateProvince="Atlantida">
|
||
<specimenCount id="9D65FD6DFFD8FFE7FBBE450DBB293D1A" box="[1046,1115,1798,1824]" pageId="38" pageNumber="39" type="male">7 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FBC1450DBBDC3D1A" box="[1129,1198,1798,1824]" pageId="38" pageNumber="39" type="female">3 ♀♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFD8FFE7FE6F4525BD693D72" box="[455,539,1838,1864]" collectionName="USA, Texas, College Station, Texas A & M University" pageId="38" pageNumber="39">TAMU</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FD9F4526BBFE3D7D" ID-GBIF-Occurrence="3093980413" box="[567,1164,1837,1864]" collectedFrom="tropical rainforest" collectingDate="1996-06-15" collectingDateMax="1996-06-19" collectingDateMin="1996-06-15" collectingMethod="intercept trap" collectionCode="UVGC" collectorName="Coll. R. Lehmann. Flight" country="Honduras" location="La Ceiba." pageId="38" pageNumber="39" specimenCount="1" specimenCount-male="1" stateProvince="Atlantida">
|
||
<specimenCount id="9D65FD6DFFD8FFE7FD9F4526BD163D72" box="[567,612,1837,1864]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
|
||
same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFD8FFE7FB914526BBFE3D7D" box="[1081,1164,1837,1863]" collectionName="Guatemala, Guatemala City, Universidad del Valle de Guatemala, Collecion de Artropodos" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="38" pageNumber="39">UVGC</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B0B3CB9FFD8FFE7FB004526BDF13D55" ID-GBIF-Occurrence="3093980515" collectedFrom="tropical rainforest" collectingDate="1996-06-15" collectingDateMax="1996-06-19" collectingDateMin="1996-06-15" collectingMethod="intercept trap" collectionCode="VMC" collectorName="Coll. R. Lehmann. Flight" country="Honduras" location="La Ceiba." pageId="38" pageNumber="39" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Atlantida">
|
||
<specimenCount id="9D65FD6DFFD8FFE7FB004526BBAA3D72" box="[1192,1240,1837,1864]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D65FD6DFFD8FFE7FB4E4526BA583D72" box="[1254,1322,1837,1864]" pageId="38" pageNumber="39" type="female">2 ♀♀</specimenCount>
|
||
; same collection data as for preceding;
|
||
<collectionCode id="ED72AE21FFD8FFE7FD94455EBDF13D55" box="[572,643,1877,1903]" pageId="38" pageNumber="39">VMC</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF154590BE2F3D8F" blockId="38.[189,349,1947,1973]" box="[189,349,1947,1973]" pageId="38" pageNumber="39">
|
||
<heading id="D0948188FFD8FFE7FF154590BE2F3D8F" bold="true" box="[189,349,1947,1973]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
|
||
<emphasis id="B917EAF6FFD8FFE7FF154590BE2F3D8F" bold="true" box="[189,349,1947,1973]" pageId="38" pageNumber="39">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD8FFE7FF1545C5BD343DD2" blockId="38.[189,582,1998,2025]" box="[189,582,1998,2025]" pageId="38" pageNumber="39">
|
||
<collectingCountry id="F3747674FFD8FFE7FF1545C5BE403DD2" box="[189,306,1998,2024]" name="Honduras" pageId="38" pageNumber="39">Honduras</collectingCountry>
|
||
,
|
||
<collectingRegion id="49A7F806FFD8FFE7FE9445C5BEDE3DD2" box="[316,428,1998,2024]" country="Honduras" name="Atlantida" pageId="38" pageNumber="39">Atlántida</collectingRegion>
|
||
, Rco Bonito.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFD9FFE6FF154303BC863C74" pageId="39" pageNumber="40" type="description">
|
||
<paragraph id="8BDC36E4FFD9FFE6FF154303BE193B18" blockId="39.[187,1399,264,432]" box="[189,363,264,290]" pageId="39" pageNumber="40">
|
||
<heading id="D0948188FFD9FFE6FF154303BE193B18" bold="true" box="[189,363,264,290]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
|
||
<emphasis id="B917EAF6FFD9FFE6FF154303BE193B18" bold="true" box="[189,363,264,290]" pageId="39" pageNumber="40">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF42433FBE0F3B74" blockId="39.[187,1399,264,432]" box="[234,381,308,334]" pageId="39" pageNumber="40">
|
||
<emphasis id="B917EAF6FFD9FFE6FF42433FBE0F3B74" bold="true" box="[234,381,308,334]" pageId="39" pageNumber="40">Major male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF15435EBCF73B8A" blockId="39.[187,1399,264,432]" pageId="39" pageNumber="40">
|
||
HEAD. Clypeus bidentate, black anteriorly, dark metallic blue, green or blue-green posteriorly; roughened sculpture. Genae dark metallic blue, green or blue-green, with roughened sculpture. Front black. Cephalic horn black, curved posteriorly over pronotum (
|
||
<figureCitation id="13582A61FFD9FFE6FD48439EBC453B8A" box="[736,823,405,432]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="39" pageNumber="40">Figs 2I</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFD9FFE6FCEB439DBC0A3B8A" box="[835,888,406,432]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="39" pageNumber="40">11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF1543DEBB89388B" blockId="39.[187,1399,468,689]" pageId="39" pageNumber="40">
|
||
PRONOTUM. Keel absent in the middle of anterior pronotal margin. Disc triangular, flat, with two distinctly developed tubercles on anterior portion. Triangle uniformly dark metallic blue, green, or bluegreen; becoming black on posterior margin and beneath the posterolateral angles, lightly granulate, scabriculous, impunctate. Sides dark metallic blue, green, or blue-green; smooth sculpture, scabriculous, with superficially impressed punctures. Lateral lines of pronotal triangle straight. Posterolateral angles widened or acute, projected posteriorly. Lateral fossae distinctly impressed. Basal fossae obtusely oval, distinctly impressed. Posterior margin with superficially impressed punctures (
|
||
<figureCitation id="13582A61FFD9FFE6FBFE409CBBDE388B" box="[1110,1196,663,689]" captionStart="Fig" captionStartId="9.[189,232,1786,1812]" captionTargetBox="[260,1330,265,1758]" captionTargetId="figure-104@9.[257,1330,265,1765]" captionTargetPageId="9" captionText="Fig. 2. Lateral view of males within the Phanaeus endymion species group. A. Phanaeus chiapanecus sp. nov. (TAMU). B. Phanaeus endymion Harold, 1863 (VMC). C. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). D. Phanaeus jackenioi sp. nov. (UVGC). E. Phanaeus malyi Arnaud, 2002 (VMC). F. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). G. Phanaeus pacificus sp. nov. (UVGC). H. Phanaeus panamensis sp. nov. (TAMU). I. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). J. Phanaeus pyrois Bates, 1887, green morph (VMC). K. Phanaeus rzedowskii sp. nov. (TAMU). L. Phanaeus zapotecus Edmonds, 2006 (VMC)." figureDoi="http://doi.org/10.5281/zenodo.4836752" httpUri="https://zenodo.org/record/4836752/files/figure.png" pageId="39" pageNumber="40">Figs 2I</figureCitation>
|
||
,
|
||
<figureCitation id="13582A61FFD9FFE6FB10409CBB9C388B" box="[1208,1262,663,689]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="39" pageNumber="40">11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF1540DDBCB6390A" blockId="39.[189,1398,725,817]" pageId="39" pageNumber="40">
|
||
ELYTRA. Striae fine, smooth, scabriculous, impressed basally as distinct fossae; dark blue, green, or bluegreen; always with distinctly impressed punctation. Interstriae black, smooth, scabriculous, with almost effaced punctures. Sutural margin without apical tooth (
|
||
<figureCitation id="13582A61FFD9FFE6FCE3411DBCC5390A" box="[843,951,790,816]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="39" pageNumber="40">Fig. 11A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF15415EBDCA3955" blockId="39.[189,696,853,879]" box="[189,696,853,879]" pageId="39" pageNumber="40">PROTIBIAE. Quadridentate with apical spine.</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF15419FBC6539F4" blockId="39.[189,1398,915,975]" pageId="39" pageNumber="40">TERGITE VIII. Dark metallic blue, green, or blue-green; scabriculous; with rough, superficially impressed punctures. Basal margin with setae variable in size.</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF1541F8BDD43E74" blockId="39.[189,1398,1011,1102]" pageId="39" pageNumber="40">
|
||
GENITALIA. Right and left lobes of endophallite copulatrix similar in size. Right lobe obtusely triangular in shape, weakly developed superiorly. Left lobe obtusely lobed, strongly developed. Central ridge less developed than central column (
|
||
<figureCitation id="13582A61FFD9FFE6FD9E4638BDEB3E74" box="[566,665,1075,1102]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="39" pageNumber="40">Fig. 1M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF424679BE0F3EB6" blockId="39.[189,1398,1138,1230]" box="[234,381,1138,1164]" pageId="39" pageNumber="40">
|
||
<emphasis id="B917EAF6FFD9FFE6FF424679BE0F3EB6" bold="true" box="[234,381,1138,1164]" pageId="39" pageNumber="40">Minor male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF154699BCA03EF4" blockId="39.[189,1398,1138,1230]" pageId="39" pageNumber="40">Like the major male, except for the reduction of the secondary sexual characters (i.e., cephalic horn, pronotal triangle and tubercles, and pronotal posterolateral angles).</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF4246F9BE373F36" blockId="39.[187,1400,1266,1519]" box="[234,325,1266,1292]" pageId="39" pageNumber="40">
|
||
<emphasis id="B917EAF6FFD9FFE6FF4246F9BE373F36" bold="true" box="[234,325,1266,1292]" pageId="39" pageNumber="40">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF154719BBE13FD5" blockId="39.[187,1400,1266,1519]" pageId="39" pageNumber="40">
|
||
Similar to the male, except for the head showing a cephalic trituberculate carina; with weakly developed, nearly aligned tubercles; carinate middle tubercle slightly more developed than lateral tubercles; frons with distinctly impressed punctures; pronotal sculpture smooth, with almost completely effaced punctures; pronotum almost completely black, becoming dark metallic blue, green, or blue-green posteriorly and laterally; pronotal process trituberculate, weakly concave posteriorly; pronotal tubercles nearly aligned; with rounded to dentiform middle tubercle, more developed than lateral tubercles; lateral tubercles rounded; posterior pronotal midline almost completely effaced (
|
||
<figureCitation id="13582A61FFD9FFE6FBB347DEBBF43FD5" box="[1051,1158,1493,1519]" captionStart="Fig" captionStartId="40.[189,232,1862,1888]" captionTargetBox="[233,1379,306,1775]" captionTargetId="figure-24@40.[189,1399,306,1813]" captionTargetPageId="40" captionText="Fig. 11. Phanaeus porioni Arnaud, 2001 stat. rev. A. ♂ (TAMU). B. ♀ (TAMU). C. Holotype, ♂ (by Patrick Arnaud) (CPFA)." figureDoi="http://doi.org/10.5281/zenodo.4836788" httpUri="https://zenodo.org/record/4836788/files/figure.png" pageId="39" pageNumber="40">Fig. 11B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF424418BE123C17" blockId="39.[189,1012,1555,1614]" box="[234,352,1555,1581]" pageId="39" pageNumber="40">
|
||
<emphasis id="B917EAF6FFD9FFE6FF424418BE123C17" bold="true" box="[234,352,1555,1581]" pageId="39" pageNumber="40">Variation</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF15443FBC863C74" blockId="39.[189,1012,1555,1614]" box="[189,1012,1588,1614]" pageId="39" pageNumber="40">
|
||
Mean length
|
||
<quantity id="4C9B9B01FFD9FFE6FEF0443FBED83C74" box="[344,426,1588,1614]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="39" pageNumber="40" unit="mm" value="17.0">17 mm</quantity>
|
||
(
|
||
<quantity id="4C9B9B01FFD9FFE6FE12443FBD193C74" box="[442,619,1588,1614]" metricMagnitude="-2" metricUnit="m" metricValue="1.66" metricValueMax="2.01" metricValueMin="1.31" pageId="39" pageNumber="40" unit="mm" value="16.6" valueMax="20.1" valueMin="13.1">13.1–20.1 mm</quantity>
|
||
). Colour variants were not found.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFD9FFE6FF154479BB063C83" box="[189,1140,1650,1721]" pageId="39" pageNumber="40" type="distribution">
|
||
<paragraph id="8BDC36E4FFD9FFE6FF154479BE243CB6" blockId="39.[189,1140,1650,1721]" box="[189,342,1650,1676]" pageId="39" pageNumber="40">
|
||
<heading id="D0948188FFD9FFE6FF154479BE243CB6" bold="true" box="[189,342,1650,1676]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
|
||
<emphasis id="B917EAF6FFD9FFE6FF154479BE243CB6" bold="true" box="[189,342,1650,1676]" pageId="39" pageNumber="40">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF154495BB063C83" blockId="39.[189,1140,1650,1721]" box="[189,1140,1694,1721]" pageId="39" pageNumber="40">
|
||
Caribbean
|
||
<collectingCountry id="F3747674FFD9FFE6FE954495BEC03C82" box="[317,434,1694,1720]" name="Honduras" pageId="39" pageNumber="40">Honduras</collectingCountry>
|
||
,
|
||
<collectingCountry id="F3747674FFD9FFE6FE164495BD7E3C82" box="[446,524,1694,1720]" name="Belize" pageId="39" pageNumber="40">Belize</collectingCountry>
|
||
, and probably north-Caribbean
|
||
<collectingCountry id="F3747674FFD9FFE6FC284495BC8D3C82" box="[896,1023,1694,1720]" name="Guatemala" pageId="39" pageNumber="40">Guatemala</collectingCountry>
|
||
(
|
||
<figureCitation id="13582A61FFD9FFE6FBA74495BB143C83" box="[1039,1126,1694,1721]" captionStart="Fig" captionStartId="50.[189,232,1890,1916]" captionTargetBox="[193,1392,927,1854]" captionTargetId="figure-235@50.[189,1398,924,1857]" captionTargetPageId="50" captionText="Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta." figureDoi="http://doi.org/10.5281/zenodo.4836796" httpUri="https://zenodo.org/record/4836796/files/figure.png" pageId="39" pageNumber="40">Fig. 15</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C379656FFFD9FFE6FF1544D6BE983DD2" pageId="39" pageNumber="40" type="discussion">
|
||
<paragraph id="8BDC36E4FFD9FFE6FF1544D6BE5D3CCD" blockId="39.[188,1399,1757,2024]" box="[189,303,1757,1783]" pageId="39" pageNumber="40">
|
||
<heading id="D0948188FFD9FFE6FF1544D6BE5D3CCD" bold="true" box="[189,303,1757,1783]" fontSize="11" level="3" pageId="39" pageNumber="40" reason="3">
|
||
<emphasis id="B917EAF6FFD9FFE6FF1544D6BE5D3CCD" bold="true" box="[189,303,1757,1783]" pageId="39" pageNumber="40">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BDC36E4FFD9FFE6FF154503BE983DD2" blockId="39.[188,1399,1757,2024]" pageId="39" pageNumber="40">
|
||
<taxonomicName id="4C634D67FFD9FFE6FF154503BEE43D18" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[189,406,1800,1826]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD9FFE6FF154503BEE43D18" box="[189,406,1800,1826]" italics="true" pageId="39" pageNumber="40">Phanaeus porioni</emphasis>
|
||
</taxonomicName>
|
||
was considered as a synonym of
|
||
<taxonomicName id="4C634D67FFD9FFE6FCFF4502BC9F3D18" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[855,1005,1800,1826]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD9FFE6FCFF4502BC9F3D18" box="[855,1005,1800,1826]" italics="true" pageId="39" pageNumber="40">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFF24B15FFD9FFE6FB814503BA013D19" author="Edmonds W. D. & Zidek J." box="[1065,1395,1800,1827]" pageId="39" pageNumber="40" pagination="1 - 108" refId="ref37762" refString="Edmonds W. D. & Zidek J. 2012. Taxonomy of Phanaeus revisited: Revised keys to and comments on species of the New World dung beetle genus Phanaeus Macleay, 1819 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini). Insecta Mundi 274: 1 - 108." type="journal article" year="2012">Edmonds & Zídek (2012)</bibRefCitation>
|
||
. Nevertheless, differences in external and genital morphology were found by us. As a consequence,
|
||
<taxonomicName id="4C634D67FFD9FFE6FF154541BE403D59" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[189,306,1866,1891]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD9FFE6FF154541BE403D59" box="[189,306,1866,1891]" italics="true" pageId="39" pageNumber="40">P. porioni</emphasis>
|
||
</taxonomicName>
|
||
is resurrected from previous synonymy and full species status is given to it. Apparently, a significant area of sympatry is found between
|
||
<taxonomicName id="4C634D67FFD9FFE6FD7F4560BC173DBE" authorityName="Harold. Ex-Mus" authorityYear="1863" box="[727,869,1898,1924]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="endymion">
|
||
<emphasis id="B917EAF6FFD9FFE6FD7F4560BC173DBE" box="[727,869,1898,1924]" italics="true" pageId="39" pageNumber="40">P. endymion</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C634D67FFD9FFE6FC354560BB7D3DBE" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[925,1039,1899,1924]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD9FFE6FC354560BB7D3DBE" box="[925,1039,1899,1924]" italics="true" pageId="39" pageNumber="40">P. porioni</emphasis>
|
||
</taxonomicName>
|
||
. The males of both species are strongly mimetic, but females are easily diagnosed by external morphology. The endophallite copulatrix will help to confidently separate males of both species (
|
||
<figureCitation id="13582A61FFD9FFE6FC9645A0BCF13DFF" box="[830,899,1963,1989]" captionStart="Fig" captionStartId="8.[189,232,1681,1707]" captionTargetBox="[334,1251,265,1642]" captionTargetId="figure-153@8.[333,1254,265,1655]" captionTargetPageId="8" captionText="Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA)." figureDoi="http://doi.org/10.5281/zenodo.4836748" httpUri="https://zenodo.org/record/4836748/files/figure.png" pageId="39" pageNumber="40">Fig. 1</figureCitation>
|
||
). We were not able to study any specimens of
|
||
<taxonomicName id="4C634D67FFD9FFE6FF7545C4BE223DD2" authorityName="P. ARNAUD DET" authorityYear="2001" baseAuthorityName="Arnaud" baseAuthorityYear="2001" box="[221,336,1999,2024]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="porioni">
|
||
<emphasis id="B917EAF6FFD9FFE6FF7545C4BE223DD2" box="[221,336,1999,2024]" italics="true" pageId="39" pageNumber="40">P. porioni</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F3747674FFD9FFE6FE3045C5BE953DD2" box="[408,487,1998,2024]" name="Belize" pageId="39" pageNumber="40">Belize</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |