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<document id="BB4BC3454FBE2C6E45363AC74B783D8F" ID-DOI="10.5281/zenodo.174573" ID-GBIF-Dataset="8c531d6d-9d32-49fd-8449-12126dac1d5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="174573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459785485270" checkinUser="plazi" docAuthor="Mesibov, Robert" docDate="2006" docId="038D879B2464FF89FEC0FAC179D9F8B1" docLanguage="en" docName="zt01354p044.pdf.imd" docOrigin="Zootaxa 1354" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tholerosoma corrugatum Mesibov, 2006, n. sp." docType="treatment" docVersion="12" lastPageNumber="43" masterDocId="FFB4FFE3246FFF85FFC8FFAE7B5AFFDC" masterDocTitle="Dirt-encrusted and dragon millipedes (Diplopoda: Polydesmida: Paradoxosomatidae) from Queensland, Australia" masterLastPageNumber="44" masterPageNumber="31" pageNumber="42" updateTime="1698218411109" updateUser="plazi">
<mods:mods id="2A45F059F0071DBC5FA6A396E735CF3A" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0BADE4BABE4A3457F7739A22AFBE9EAC">
<mods:title id="C952091AA8DBECE1A1589002E8DBAB74">Dirt-encrusted and dragon millipedes (Diplopoda: Polydesmida: Paradoxosomatidae) from Queensland, Australia</mods:title>
</mods:titleInfo>
<mods:name id="D69BC059ECD56A84AF3CBDD464B3A199" type="personal">
<mods:role id="F57C54EB647945C69089831C199B442A">
<mods:roleTerm id="A8499652E15FA51B0DEF42CF8A585995">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="70C54906DB87E6CD6C7F7219AE5AF981">Mesibov, Robert</mods:namePart>
</mods:name>
<mods:typeOfResource id="6DF4C35C315BB627495A0AFF6EC8922A">text</mods:typeOfResource>
<mods:relatedItem id="C3AA12415016FFFE2C7558D0DC4DF98E" type="host">
<mods:titleInfo id="F7D2C8D1DF3BDCF7662820A2F90E64DF">
<mods:title id="1ADD2E2BF5033236C8787C38513D2F00">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="21E2D509ECEC7B53326E5A21A41FED18">
<mods:date id="525AC8EE34D2EAAEB0D679E18E876E6B">2006</mods:date>
<mods:detail id="00836505C844CF9C2D3FE76F7F16EB13" type="volume">
<mods:number id="2ACC98F89CF3FBEE411F6D723BCAD33E">1354</mods:number>
</mods:detail>
<mods:extent id="ADE6D856F66CB4D3C85E31E7E79D670F" unit="page">
<mods:start id="E6493162728995233BC3B52FC64D7D55">31</mods:start>
<mods:end id="95551047A338A97DB0FCB7B2F389A593">44</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C95F04223597F8D79B381E81E2152037">journal article</mods:classification>
<mods:identifier id="594A7351F9729E6B0C91CBB7BEAFFDDA" type="DOI">10.5281/zenodo.174573</mods:identifier>
<mods:identifier id="3D7C8CFB97AA941045BD837FC1B514F2" type="GBIF-Dataset">8c531d6d-9d32-49fd-8449-12126dac1d5c</mods:identifier>
<mods:identifier id="B15CEDCB691D4E72D6C562FE11EA6360" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="919E5E37AAF756CC70AE228019CBDF9E" type="Zenodo-Dep">174573</mods:identifier>
</mods:mods>
<treatment id="038D879B2464FF89FEC0FAC179D9F8B1" ID-DOI="http://doi.org/10.5281/zenodo.6261707" ID-GBIF-Taxon="119348955" ID-Zenodo-Dep="6261707" LSID="urn:lsid:plazi:treatment:038D879B2464FF89FEC0FAC179D9F8B1" httpUri="http://treatment.plazi.org/id/038D879B2464FF89FEC0FAC179D9F8B1" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42">
<subSubSection id="C33E65062464FF8EFEC0FAC179E3FA70" pageId="11" pageNumber="42" type="nomenclature">
<paragraph id="8B9B368D2464FF8EFEC0FAC179D2FA55" blockId="11.[264,697,1391,1452]" box="[264,648,1391,1417]" pageId="11" pageNumber="42">
<heading id="D0D381E12464FF8EFEC0FAC179D2FA55" bold="true" box="[264,648,1391,1417]" fontSize="11" level="1" pageId="11" pageNumber="42" reason="1">
<emphasis id="B950EA9F2464FF8EFEC0FAC179D2FA55" bold="true" box="[264,648,1391,1417]" pageId="11" pageNumber="42">
<taxonomicName id="4C244D0E2464FF8EFEC0FAC17964FA55" ID-CoL="56JF5" box="[264,574,1391,1417]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="corrugatum" status="sp. nov.">
<emphasis id="B950EA9F2464FF8EFEC0FAC17964FA55" bold="true" box="[264,574,1391,1417]" italics="true" pageId="11" pageNumber="42">Tholerosoma corrugatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26357E42464FF8EFD8EFAC179D2FA55" box="[582,648,1391,1417]" pageId="11" pageNumber="42" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9B368D2464FF8EFEC0FA3C79E3FA70" blockId="11.[264,697,1391,1452]" box="[264,697,1426,1452]" pageId="11" pageNumber="42">
<figureCitation id="131F2A082464FF8EFEC0FA3C7A02FA70" box="[264,344,1426,1452]" captionStart="FIGURE 5" captionStartId="11.[264,367,1102,1126]" captionTargetBox="[275,1299,482,1066]" captionTargetId="figure@11.[264,1322,466,1078]" captionTargetPageId="11" captionText="FIGURE 5. Tholerosoma monteithi n. gen., n. sp. (A) medial view of right gonopod, (D) antenna, (E) leg 6 (male paratype, QM S 73974). Tholerosoma corrugatum n. sp. (B) medial view of right gonopod (holotype, QM S 73993), (C) medial view of right gonopod (male paratype, QM S 73994), (F) outline sketch of midbody metazonite, showing location of swellings on posterior edge (female paratype, QM S 73995). Setation not shown; dashed lines in (A) – (C) indicate course of prostatic groove. Scale bars for (A) – (C) and for (D) and (E) = 0.25 mm." httpUri="https://zenodo.org/record/174578/files/figure.png" pageId="11" pageNumber="42">Figs. 5</figureCitation>
B, 5C, 5F, 6A, 6C; map
<figureCitation id="131F2A082464FF8EFDBCFA3C79E3FA70" box="[628,697,1426,1452]" captionStart="FIGURE 7" captionStartId="13.[264,367,883,907]" captionTargetBox="[267,1307,331,850]" captionTargetId="figure@13.[264,1317,326,853]" captionTargetPageId="13" captionText="FIGURE 7. Localities for Desmoxytoides hasenpuschorum n. gen., n. sp. (left, squares), Tholerosoma monteithi n. gen., n. sp. (centre, triangles) and T. corrugatum n. sp. (centre, stars). Rectangle in Australia map (right) shows boundary of main maps." httpUri="https://zenodo.org/record/174580/files/figure.png" pageId="11" pageNumber="42">Fig. 7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33E65062464FF8EFEC0FA4C78B8F8E0" pageId="11" pageNumber="42" type="materials_examined">
<paragraph id="8B9B368D2464FF8EFEC0FA4C7828F9F8" blockId="11.[264,1323,1506,1892]" pageId="11" pageNumber="42">
<materialsCitation id="3B4C3CD02464FF8EFEC0FA4C7828F9F8" ID-GBIF-Occurrence="2285829273" collectedFrom="fern glade, pitfall &amp; intercept" collectingDate="1991-03-26" collectionCode="QM" collectorName="A. Graham" elevation="1150" latitude="-19.5" location="Mt Elliot summit" longLatPrecision="1271" longitude="146.95" pageId="11" pageNumber="42" specimenCode="S73993" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B950EA9F2464FF8EFEC0FA4C7A29FA27" box="[264,371,1506,1531]" italics="true" pageId="11" pageNumber="42">
<typeStatus id="549F882F2464FF8EFEC0FA4C7A29FA27" box="[264,371,1506,1531]" pageId="11" pageNumber="42" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="9D22FD042464FF8EFE4CFA4C7A99FA20" box="[388,451,1506,1532]" pageId="11" pageNumber="42" type="male">Male</specimenCount>
,
<location id="8EFB60562464FF8EFE18FA4C79C5FA20" LSID="urn:lsid:plazi:treatment:038D879B2464FF89FEC0FAC179D9F8B1:8EFB60562464FF8EFE18FA4C79C5FA20" box="[464,671,1506,1532]" latitude="-19.5" longLatPrecision="1271" longitude="146.95" name="Mt Elliot summit" pageId="11" pageNumber="42">Mt Elliot summit</location>
,
<geoCoordinate id="EE10504A2464FF8EFD64FA4C7856FA20" box="[684,780,1506,1532]" degrees="19" direction="south" minutes="30" orientation="latitude" pageId="11" pageNumber="42" precision="925" value="-19.5">19°30’S</geoCoordinate>
<geoCoordinate id="EE10504A2464FF8EFCDCFA4C78DDFA20" box="[788,903,1506,1532]" degrees="146" direction="east" minutes="57" orientation="longitude" pageId="11" pageNumber="42" precision="925" value="146.95">146°57’E</geoCoordinate>
,
<elevation id="0009D1BE2464FF8EFC5CFA4C78AAFA20" box="[916,1008,1506,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="42" unit="m" value="1150.0">
<quantity id="4CDC9B682464FF8EFC5CFA4C78AAFA20" box="[916,1008,1506,1532]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="11" pageNumber="42" unit="m" value="1150.0">1150 m</quantity>
</elevation>
,
<collectingDate id="EFDEE9A52464FF8EFC35FA4C7E5BFA20" box="[1021,1281,1506,1532]" pageId="11" pageNumber="42">
January to
<date id="FF9A104D2464FF8EFBB7FA4C7E5BFA20" box="[1151,1281,1506,1532]" pageId="11" pageNumber="42" value="1991-03-26">26.iii.1991</date>
</collectingDate>
,
<collectorName id="26D1535B2464FF8EFAC6FA4C7A33F9F8" pageId="11" pageNumber="42">A. Graham</collectorName>
,
<collectedFrom id="383875702464FF8EFEBDF9A47995F9F8" box="[373,719,1546,1572]" pageId="11" pageNumber="42">fern glade, pitfall &amp; intercept</collectedFrom>
,
<collectionCode id="ED35AE482464FF8EFD12F9A47853F9F8" box="[730,777,1546,1572]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="42" type="Museum">QM</collectionCode>
<specimenCode id="DB829EF62464FF8EFCD8F9A47834F9F8" box="[784,878,1546,1572]" pageId="11" pageNumber="42">
<accessionNumber id="9477AB6E2464FF8EFCD8F9A47834F9F8" box="[784,878,1546,1572]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S73993" pageId="11" pageNumber="42" type="EnaNcbi">S73993</accessionNumber>
</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8B9B368D2464FF8EFEF0F99C78B8F8E0" blockId="11.[264,1323,1506,1892]" pageId="11" pageNumber="42">
<materialsCitation id="3B4C3CD02464FF8EFEF0F99C7F98F9A8" ID-GBIF-Occurrence="2285829297" collectedFrom="rainforest" collectingDate="1999-10-04" collectingDateMax="1999-12-17" collectingDateMin="1999-10-04" collectingMethod="pitfall" collectionCode="QM" collectorName="D. &amp; I. Cook" elevation="280" latitude="-21.616667" location="track crossing" longLatPrecision="1263" longitude="148.98334" municipality="Stony Creek" pageId="11" pageNumber="42" specimenCode="S73994" specimenCount="1" typeStatus="paratype">
<emphasis id="B950EA9F2464FF8EFEF0F99C7AF5F997" box="[312,431,1586,1611]" italics="true" pageId="11" pageNumber="42">
<typeStatus id="549F882F2464FF8EFEF0F99C7AF5F997" box="[312,431,1586,1611]" pageId="11" pageNumber="42" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D22FD042464FF8EFE07F99C7954F990" box="[463,526,1586,1612]" pageId="11" pageNumber="42" type="male">Male</specimenCount>
,
<collectingMunicipality id="6BFFACF72464FF8EFDE2F99C7995F990" box="[554,719,1586,1612]" pageId="11" pageNumber="42">Stony Creek</collectingMunicipality>
,
<location id="8EFB60562464FF8EFD24F99C78F8F990" LSID="urn:lsid:plazi:treatment:038D879B2464FF89FEC0FAC179D9F8B1:8EFB60562464FF8EFD24F99C78F8F990" box="[748,930,1586,1612]" latitude="-21.616667" longLatPrecision="1263" longitude="148.98334" municipality="Stony Creek" name="track crossing" pageId="11" pageNumber="42">track crossing</location>
,
<geoCoordinate id="EE10504A2464FF8EFC77F99C7F45F990" box="[959,1055,1586,1612]" degrees="21" direction="south" minutes="37" orientation="latitude" pageId="11" pageNumber="42" precision="925" value="-21.616667">21°37’S</geoCoordinate>
<geoCoordinate id="EE10504A2464FF8EFBF0F99C7FF1F990" box="[1080,1195,1586,1612]" degrees="148" direction="east" minutes="59" orientation="longitude" pageId="11" pageNumber="42" precision="925" value="148.98334">148°59’E</geoCoordinate>
,
<elevation id="0009D1BE2464FF8EFB00F99C7E7CF990" box="[1224,1318,1586,1612]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="42" unit="m" value="280.0">
<quantity id="4CDC9B682464FF8EFB00F99C7E7CF990" box="[1224,1318,1586,1612]" metricMagnitude="2" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="42" unit="m" value="280.0">280 m</quantity>
</elevation>
,
<collectingDate id="EFDEE9A52464FF8EFEC0F9F47A91F9A8" box="[264,459,1626,1652]" pageId="11" pageNumber="42" value="1999-10-04" valueMax="1999-12-17" valueMin="1999-10-04">
<date id="FF9A104D2464FF8EFEC0F9F47A91F9A8" box="[264,459,1626,1652]" pageId="11" pageNumber="42" value="1999-10-04" valueMax="1999-12-17" valueMin="1999-10-04">4.x.–17.xii.1999</date>
</collectingDate>
,
<collectorName id="26D1535B2464FF8EFE14F9F479D0F9A8" box="[476,650,1626,1652]" pageId="11" pageNumber="42">D. &amp; I. Cook</collectorName>
,
<collectedFrom id="383875702464FF8EFD53F9F47854F9A8" box="[667,782,1626,1652]" pageId="11" pageNumber="42">rainforest</collectedFrom>
,
<collectingMethod id="52654E9A2464FF8EFCD7F9F47832F9A8" box="[799,872,1626,1652]" pageId="11" pageNumber="42">pitfall</collectingMethod>
, sample 9030,
<collectionCode id="ED35AE482464FF8EFBE0F9F47F0DF9A8" box="[1064,1111,1626,1652]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="42" type="Museum">QM</collectionCode>
<specimenCode id="DB829EF62464FF8EFBADF9F47F98F9A8" box="[1125,1218,1626,1652]" pageId="11" pageNumber="42">
<accessionNumber id="9477AB6E2464FF8EFBADF9F47F98F9A8" box="[1125,1218,1626,1652]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S73994" pageId="11" pageNumber="42" type="EnaNcbi">S73994</accessionNumber>
</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B4C3CD02464FF8EFB1BF9F47F74F918" ID-GBIF-Occurrence="2285829277" collectingDate="1986-12-06" collectingDateMax="1986-12-08" collectingDateMin="1986-12-06" collectionCode="QM" collectorName="G. Monteith &amp; G. Thompson &amp; Hamlet" elevation="650" latitude="-19.2" location="Bluewater Range" longLatPrecision="1273" longitude="146.4" pageId="11" pageNumber="42" specimenCode="S73995" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D22FD042464FF8EFB1BF9F47E7CF9A8" box="[1235,1318,1626,1652]" pageId="11" pageNumber="42" type="female">female</specimenCount>
,
<location id="8EFB60562464FF8EFEC0F92C7AB8F940" LSID="urn:lsid:plazi:treatment:038D879B2464FF89FEC0FAC179D9F8B1:8EFB60562464FF8EFEC0F92C7AB8F940" box="[264,482,1666,1692]" latitude="-19.2" longLatPrecision="1273" longitude="146.4" name="Bluewater Range" pageId="11" pageNumber="42">Bluewater Range</location>
,
<locationDeviation id="9A292FAF2464FF8EFE33F92C7822F940" box="[507,888,1666,1692]" location="Townsville" pageId="11" pageNumber="42">
<quantity id="4CDC9B682464FF8EFE33F92C790BF940" box="[507,593,1666,1692]" metricMagnitude="4" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="42" unit="km" value="45.0">45 km</quantity>
WNW of Townsville
</locationDeviation>
,
<geoCoordinate id="EE10504A2464FF8EFC59F92C78ABF940" box="[913,1009,1666,1692]" degrees="19" direction="south" minutes="12" orientation="latitude" pageId="11" pageNumber="42" precision="925" value="-19.2">19°12’S</geoCoordinate>
<geoCoordinate id="EE10504A2464FF8EFBCDF92C7F22F940" box="[1029,1144,1666,1692]" degrees="146" direction="east" minutes="24" orientation="longitude" pageId="11" pageNumber="42" precision="925" value="146.4">146°24’E</geoCoordinate>
,
<elevation id="0009D1BE2464FF8EFB59F92C7E7CF940" box="[1169,1318,1666,1692]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="11" pageNumber="42" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">
<quantity id="4CDC9B682464FF8EFB59F92C7E7CF940" box="[1169,1318,1666,1692]" metricMagnitude="2" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="11" pageNumber="42" unit="m" value="650.0" valueMax="700.0" valueMin="600.0">600–700 m</quantity>
</elevation>
,
<date id="FF9A104D2464FF8EFEC0F9047AC4F918" box="[264,414,1706,1732]" pageId="11" pageNumber="42" value="1986-12-06" valueMax="1986-12-08" valueMin="1986-12-06">
<collectingDate id="EFDEE9A52464FF8EFEC0F9047AC4F918" box="[264,414,1706,1732]" pageId="11" pageNumber="42" value="1986-12-06" valueMax="1986-12-08" valueMin="1986-12-06">6–8.xii.1986</collectingDate>
</date>
,
<collectorName id="26D1535B2464FF8EFE64F9047967F918" box="[428,573,1706,1732]" pageId="11" pageNumber="42">G. Monteith</collectorName>
,
<collectorName id="26D1535B2464FF8EFD82F90479B3F918" box="[586,745,1706,1732]" pageId="11" pageNumber="42">G. Thompson</collectorName>
&amp; S.
<collectorName id="26D1535B2464FF8EFCFBF90478D6F918" box="[819,908,1706,1732]" pageId="11" pageNumber="42">Hamlet</collectorName>
,
<collectionCode id="ED35AE482464FF8EFC51F9047892F918" box="[921,968,1706,1732]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="42" type="Museum">QM</collectionCode>
<specimenCode id="DB829EF62464FF8EFC1AF9047F74F918" box="[978,1070,1706,1732]" pageId="11" pageNumber="42">
<accessionNumber id="9477AB6E2464FF8EFC1AF9047F74F918" box="[978,1070,1706,1732]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S73995" pageId="11" pageNumber="42" type="EnaNcbi">S73995</accessionNumber>
</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B4C3CD02464FF8EFBF4F90479C1F8C8" ID-GBIF-Occurrence="2285829280" collectingDate="1992-11-19" collectingDateMax="1992-11-20" collectingDateMin="1992-11-19" collectionCode="QM" collectorName="G. Monteith &amp; G. Thompson &amp; D. Cook &amp; Janetzki" elevation="350" latitude="-20.333334" location="Mt Hayward" longLatPrecision="1268" longitude="148.75" pageId="11" pageNumber="42" specimenCode="S73996" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D22FD042464FF8EFBF4F9047E7CF918" box="[1084,1318,1706,1732]" pageId="11" pageNumber="42">stadium VII female</specimenCount>
,
<location id="8EFB60562464FF8EFEC0F97C7AF9F930" LSID="urn:lsid:plazi:treatment:038D879B2464FF89FEC0FAC179D9F8B1:8EFB60562464FF8EFEC0F97C7AF9F930" box="[264,419,1746,1772]" latitude="-20.333334" longLatPrecision="1268" longitude="148.75" name="Mt Hayward" pageId="11" pageNumber="42">Mt Hayward</location>
,
<geoCoordinate id="EE10504A2464FF8EFE7AF97C7949F930" box="[434,531,1746,1772]" degrees="20" direction="south" minutes="20" orientation="latitude" pageId="11" pageNumber="42" precision="925" value="-20.333334">20°20’S</geoCoordinate>
<geoCoordinate id="EE10504A2464FF8EFDD5F97C79CAF930" box="[541,656,1746,1772]" degrees="148" direction="east" minutes="45" orientation="longitude" pageId="11" pageNumber="42" precision="925" value="148.75">148°45’E</geoCoordinate>
,
<elevation id="0009D1BE2464FF8EFD57F97C79AAF930" box="[671,752,1746,1772]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="42" unit="m" value="350.0">
<quantity id="4CDC9B682464FF8EFD57F97C79AAF930" box="[671,752,1746,1772]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="42" unit="m" value="350.0">350 m</quantity>
</elevation>
,
<collectingDate id="EFDEE9A52464FF8EFD37F97C78F1F930" box="[767,939,1746,1772]" pageId="11" pageNumber="42" value="1992-11-19" valueMax="1992-11-20" valueMin="1992-11-19">
<date id="FF9A104D2464FF8EFD37F97C78F1F930" box="[767,939,1746,1772]" pageId="11" pageNumber="42" value="1992-11-19" valueMax="1992-11-20" valueMin="1992-11-19">19–20.xi.1992</date>
</collectingDate>
,
<collectorName id="26D1535B2464FF8EFC72F97C7F16F930" box="[954,1100,1746,1772]" pageId="11" pageNumber="42">G. Monteith</collectorName>
,
<collectorName id="26D1535B2464FF8EFB93F97C7FA5F930" box="[1115,1279,1746,1772]" pageId="11" pageNumber="42">G. Thompson</collectorName>
,
<collectorName id="26D1535B2464FF8EFAC6F97C7A1DF8C8" pageId="11" pageNumber="42">D. Cook</collectorName>
&amp; H.
<collectorName id="26D1535B2464FF8EFE5DF9547AA2F8C8" box="[405,504,1786,1812]" pageId="11" pageNumber="42">Janetzki</collectorName>
,
<collectionCode id="ED35AE482464FF8EFDCDF954796EF8C8" box="[517,564,1786,1812]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="42" type="Museum">QM</collectionCode>
<specimenCode id="DB829EF62464FF8EFDF5F95479C1F8C8" box="[573,667,1786,1812]" pageId="11" pageNumber="42">
<accessionNumber id="9477AB6E2464FF8EFDF5F95479C1F8C8" box="[573,667,1786,1812]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S73996" pageId="11" pageNumber="42" type="EnaNcbi">S73996</accessionNumber>
</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B4C3CD02464FF8EFD60F9547884F8E0" ID-GBIF-Occurrence="2285829289" collectedFrom="rainforest" collectingDate="2001-02-21" collectionCode="QM" collectorName="G. Monteith" elevation="450" latitude="-18.75" location="Palm Island site 1" longLatPrecision="1274" longitude="146.6" pageId="11" pageNumber="42" specimenCode="S73997" specimenCount="1" typeStatus="paratype">
<specimenCount id="9D22FD042464FF8EFD60F95479A1F8C8" box="[680,763,1786,1812]" pageId="11" pageNumber="42" type="female">female</specimenCount>
,
<location id="8EFB60562464FF8EFCC0F9547884F8C8" LSID="urn:lsid:plazi:treatment:038D879B2464FF89FEC0FAC179D9F8B1:8EFB60562464FF8EFCC0F9547884F8C8" box="[776,990,1786,1812]" latitude="-18.75" longLatPrecision="1274" longitude="146.6" name="Palm Island site 1" pageId="11" pageNumber="42">Palm Island site 1</location>
,
<geoCoordinate id="EE10504A2464FF8EFC25F9547F17F8C8" box="[1005,1101,1786,1812]" degrees="18" direction="south" minutes="45" orientation="latitude" pageId="11" pageNumber="42" precision="925" value="-18.75">18°45’S</geoCoordinate>
<geoCoordinate id="EE10504A2464FF8EFB9EF9547F93F8C8" box="[1110,1225,1786,1812]" degrees="146" direction="east" minutes="36" orientation="longitude" pageId="11" pageNumber="42" precision="925" value="146.6">146°36’E</geoCoordinate>
,
<elevation id="0009D1BE2464FF8EFB1FF9547E7CF8C8" box="[1239,1318,1786,1812]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="42" unit="m" value="450.0">
<quantity id="4CDC9B682464FF8EFB1FF9547E7CF8C8" box="[1239,1318,1786,1812]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="42" unit="m" value="450.0">450 m</quantity>
</elevation>
,
<collectingDate id="EFDEE9A52464FF8EFEC0F88C7ADBF8E0" box="[264,385,1826,1852]" pageId="11" pageNumber="42" value="2001-02-21">
<date id="FF9A104D2464FF8EFEC0F88C7ADBF8E0" box="[264,385,1826,1852]" pageId="11" pageNumber="42" value="2001-02-21">21.ii.2001</date>
</collectingDate>
,
<collectorName id="26D1535B2464FF8EFE46F88C7946F8E0" box="[398,540,1826,1852]" pageId="11" pageNumber="42">G. Monteith</collectorName>
,
<collectedFrom id="383875702464FF8EFDE0F88C79C1F8E0" box="[552,667,1826,1852]" pageId="11" pageNumber="42">rainforest</collectedFrom>
, sample 8323,
<collectionCode id="ED35AE482464FF8EFC82F88C7823F8E0" box="[842,889,1826,1852]" collectionName="Australia, Queensland, South Brisbane, Queensland Museum" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="11" pageNumber="42" type="Museum">QM</collectionCode>
<specimenCode id="DB829EF62464FF8EFC49F88C7884F8E0" box="[897,990,1826,1852]" pageId="11" pageNumber="42">
<accessionNumber id="9477AB6E2464FF8EFC49F88C7884F8E0" box="[897,990,1826,1852]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S73997" pageId="11" pageNumber="42" type="EnaNcbi">S73997</accessionNumber>
</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E65062464FF89FEF0F8E47958FE5A" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42" type="diagnosis">
<paragraph id="8B9B368D2464FF89FEF0F8E47958FE5A" blockId="11.[264,1323,1506,1892]" lastBlockId="12.[264,1323,284,390]" lastPageId="12" lastPageNumber="43" pageId="11" pageNumber="42">
<emphasis id="B950EA9F2464FF8EFEF0F8E47AF5F8BF" box="[312,431,1866,1891]" italics="true" pageId="11" pageNumber="42">Diagnosis</emphasis>
: Distinguished from
<taxonomicName id="4C244D0E2464FF8EFD66F8E47862F8BF" box="[686,824,1866,1891]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="11" pageNumber="42" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B950EA9F2464FF8EFD66F8E47862F8BF" box="[686,824,1866,1891]" italics="true" pageId="11" pageNumber="42">T. monteithi</emphasis>
</taxonomicName>
by having 6 small swellings on posterior edge of metatergite, rather than 2 large paramedian swellings on either side of transverse furrow; distal portion of telopodite curving towards base; and spiracles without plate-like structures on surface.
</paragraph>
</subSubSection>
<caption id="DF5B66052463FF89FEC0FC6D78BCFB90" httpUri="https://zenodo.org/record/3345166/files/figure.png" pageId="12" pageNumber="43" targetBox="[275,1311,432,939]" targetPageId="12">
<paragraph id="8B9B368D2463FF89FEC0FC6D78BCFB90" blockId="12.[264,1323,963,1100]" pageId="12" pageNumber="43">
<emphasis id="B950EA9F2463FF89FEC0FC6D7AD7FC07" bold="true" box="[264,397,963,987]" pageId="12" pageNumber="43">FIGURE 6.</emphasis>
<taxonomicName id="4C244D0E2463FF89FE50FC6A79F1FC07" box="[408,683,964,987]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="corrugatum" status="sp. nov.">
<emphasis id="B950EA9F2463FF89FE50FC6A79F1FC07" box="[408,683,964,987]" italics="true" pageId="12" pageNumber="43">Tholerosoma corrugatum</emphasis>
</taxonomicName>
<emphasis id="B950EA9F2463FF89FD7DFC6D79AEFC07" bold="true" box="[693,756,963,987]" pageId="12" pageNumber="43">
<taxonomicNameLabel id="A26357E42463FF89FD7DFC6D79AEFC07" box="[693,756,963,987]" pageId="12" pageNumber="43" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(A) dorsal view of midbody ring, (C) spiracles on midbody ring, anterior to left (holotype, QM
<accessionNumber id="9477AB6E2463FF89FD30FC477815FBDC" box="[760,847,1001,1024]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S73993" pageId="12" pageNumber="43" type="EnaNcbi">S73993</accessionNumber>
).
<taxonomicName id="4C244D0E2463FF89FCAAFC477F01FBDC" box="[866,1115,1001,1024]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="monteithi" status="gen. nov.">
<emphasis id="B950EA9F2463FF89FCAAFC477F01FBDC" box="[866,1115,1001,1024]" italics="true" pageId="12" pageNumber="43">Tholerosoma monteithi</emphasis>
</taxonomicName>
<emphasis id="B950EA9F2463FF89FBACFC467FA4FBDC" bold="true" box="[1124,1278,1000,1024]" pageId="12" pageNumber="43">
<taxonomicNameLabel id="A26357E42463FF89FBACFC467FE8FBDC" box="[1124,1202,1000,1024]" pageId="12" pageNumber="43" rank="genus">n. gen.</taxonomicNameLabel>
, n. sp.
</emphasis>
(B) anterior spiracle on midbody ring, anterior to left (male paratype partly cleaned with ultrasound, QM
<accessionNumber id="9477AB6E2463FF89FEF2FB9B7AD5FB90" box="[314,399,1077,1100]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S73975" pageId="12" pageNumber="43" type="EnaNcbi">S73975</accessionNumber>
). Scale bar for (A) = 0.5 mm, for (B) and (C) = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C33E65062463FF89FEF0FB2A78E1FA51" pageId="12" pageNumber="43" type="description">
<paragraph id="8B9B368D2463FF89FEF0FB2A78E1FA51" blockId="12.[264,1323,1155,1901]" pageId="12" pageNumber="43">
<emphasis id="B950EA9F2463FF89FEF0FB2A7A98FB41" box="[312,450,1156,1181]" italics="true" pageId="12" pageNumber="43">Description</emphasis>
: As for
<taxonomicName id="4C244D0E2463FF89FDEAFB2A79F7FB41" box="[546,685,1156,1181]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B950EA9F2463FF89FDEAFB2A79F7FB41" box="[546,685,1156,1181]" italics="true" pageId="12" pageNumber="43">T. monteithi</emphasis>
</taxonomicName>
, but in overall size at the lower end of the
<taxonomicName id="4C244D0E2463FF89FB57FB2A7E71FB41" box="[1183,1323,1156,1181]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B950EA9F2463FF89FB57FB2A7E71FB41" box="[1183,1323,1156,1181]" italics="true" pageId="12" pageNumber="43">T. monteithi</emphasis>
</taxonomicName>
range. Collum and metatergites with 6 small, oblong swellings on posterior edge, often hidden by encrusting dirt (
<figureCitation id="131F2A082463FF89FD88FB7D79CBFB31" box="[576,657,1235,1261]" captionStart="FIGURE 5" captionStartId="11.[264,367,1102,1126]" captionTargetBox="[275,1299,482,1066]" captionTargetId="figure@11.[264,1322,466,1078]" captionTargetPageId="11" captionText="FIGURE 5. Tholerosoma monteithi n. gen., n. sp. (A) medial view of right gonopod, (D) antenna, (E) leg 6 (male paratype, QM S 73974). Tholerosoma corrugatum n. sp. (B) medial view of right gonopod (holotype, QM S 73993), (C) medial view of right gonopod (male paratype, QM S 73994), (F) outline sketch of midbody metazonite, showing location of swellings on posterior edge (female paratype, QM S 73995). Setation not shown; dashed lines in (A) – (C) indicate course of prostatic groove. Scale bars for (A) – (C) and for (D) and (E) = 0.25 mm." httpUri="https://zenodo.org/record/174578/files/figure.png" pageId="12" pageNumber="43">Figs. 5</figureCitation>
F, 6A). Longitudinal corrugations at waist not as long or as deep as in
<taxonomicName id="4C244D0E2463FF89FE7FFB527912FAC9" box="[439,584,1276,1301]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B950EA9F2463FF89FE7FFB527912FAC9" box="[439,584,1276,1301]" italics="true" pageId="12" pageNumber="43">T. monteithi</emphasis>
</taxonomicName>
. Spiracles protruding, densely filled with extremely minute, blunt processes as in
<taxonomicName id="4C244D0E2463FF89FDC3FA8A79C3FAE1" box="[523,665,1316,1341]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B950EA9F2463FF89FDC3FA8A79C3FAE1" box="[523,665,1316,1341]" italics="true" pageId="12" pageNumber="43">T. monteithi</emphasis>
</taxonomicName>
, but without plate-like structures on surface (
<figureCitation id="131F2A082463FF89FB76FA8D7E5DFAE1" box="[1214,1287,1315,1341]" captionStart="FIGURE 6" captionStartId="12.[264,367,963,987]" captionTargetBox="[275,1311,432,939]" captionTargetId="figure@12.[275,1312,432,939]" captionTargetPageId="12" captionText="FIGURE 6. Tholerosoma corrugatum n. sp. (A) dorsal view of midbody ring, (C) spiracles on midbody ring, anterior to left (holotype, QM S 73993). Tholerosoma monteithi n. gen., n. sp. (B) anterior spiracle on midbody ring, anterior to left (male paratype partly cleaned with ultrasound, QM S 73975). Scale bar for (A) = 0.5 mm, for (B) and (C) = 0.1 mm." httpUri="https://zenodo.org/record/3345166/files/figure.png" pageId="12" pageNumber="43">Fig. 6</figureCitation>
C). Distal portion of gonopod telopodite tapering gradually, curving towards base after abrupt lateral turn; tip pointed towards base in
<typeStatus id="549F882F2463FF89FD12FADD781AFA51" box="[730,832,1395,1421]" pageId="12" pageNumber="43" type="holotype">holotype</typeStatus>
(
<figureCitation id="131F2A082463FF89FC98FADD78CDFA51" box="[848,919,1395,1421]" captionStart="FIGURE 5" captionStartId="11.[264,367,1102,1126]" captionTargetBox="[275,1299,482,1066]" captionTargetId="figure@11.[264,1322,466,1078]" captionTargetPageId="11" captionText="FIGURE 5. Tholerosoma monteithi n. gen., n. sp. (A) medial view of right gonopod, (D) antenna, (E) leg 6 (male paratype, QM S 73974). Tholerosoma corrugatum n. sp. (B) medial view of right gonopod (holotype, QM S 73993), (C) medial view of right gonopod (male paratype, QM S 73994), (F) outline sketch of midbody metazonite, showing location of swellings on posterior edge (female paratype, QM S 73995). Setation not shown; dashed lines in (A) – (C) indicate course of prostatic groove. Scale bars for (A) – (C) and for (D) and (E) = 0.25 mm." httpUri="https://zenodo.org/record/174578/files/figure.png" pageId="12" pageNumber="43">Fig. 5</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="C33E65062463FF89FEF0FA3279C7F9F1" pageId="12" pageNumber="43" type="distribution">
<paragraph id="8B9B368D2463FF89FEF0FA3279C7F9F1" blockId="12.[264,1323,1155,1901]" pageId="12" pageNumber="43">
<emphasis id="B950EA9F2463FF89FEF0FA32793BFA69" box="[312,609,1436,1461]" italics="true" pageId="12" pageNumber="43">Distribution and habitat</emphasis>
: Known from five sites in tropical rainforest from Palm Island north of Townsville to the Mackay area in north Queensland, an approximate linear range of
<quantity id="4CDC9B682463FF89FEA7FA457A9DF9D9" box="[367,455,1515,1541]" metricMagnitude="5" metricUnit="m" metricValue="4.3" pageId="12" pageNumber="43" unit="km" value="430.0">430 km</quantity>
and an elevation range of
<quantity id="4CDC9B682463FF89FD34FA4578CBF9D9" box="[764,913,1515,1541]" metricMagnitude="2" metricUnit="m" metricValue="7.15" metricValueMax="11.5" metricValueMin="2.8" pageId="12" pageNumber="43" unit="m" value="715.0" valueMax="1150.0" valueMin="280.0">280–1150 m</quantity>
. Sympatric with
<taxonomicName id="4C244D0E2463FF89FB9FFA427FB8F9D9" box="[1111,1250,1516,1541]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B950EA9F2463FF89FB9FFA427FB8F9D9" box="[1111,1250,1516,1541]" italics="true" pageId="12" pageNumber="43">T. monteithi</emphasis>
</taxonomicName>
in the Bluewater Range near Townsville.
</paragraph>
</subSubSection>
<subSubSection id="C33E65062463FF89FEF0F9927AE5F9A1" pageId="12" pageNumber="43" type="etymology">
<paragraph id="8B9B368D2463FF89FEF0F9927AE5F9A1" blockId="12.[264,1323,1155,1901]" pageId="12" pageNumber="43">
<emphasis id="B950EA9F2463FF89FEF0F9927AEFF989" box="[312,437,1596,1621]" italics="true" pageId="12" pageNumber="43">Etymology</emphasis>
: Latin
<emphasis id="B950EA9F2463FF89FDC3F99279D6F989" box="[523,652,1596,1621]" italics="true" pageId="12" pageNumber="43">corrugatus</emphasis>
(“ridged”), from the appearance of the posterior edge of the metatergite.
</paragraph>
</subSubSection>
<subSubSection id="C33E65062463FF89FEF0F92279D9F8B1" pageId="12" pageNumber="43" type="discussion">
<paragraph id="8B9B368D2463FF89FEF0F92279D9F8B1" blockId="12.[264,1323,1155,1901]" pageId="12" pageNumber="43">
<emphasis id="B950EA9F2463FF89FEF0F9227AC4F979" box="[312,414,1676,1701]" italics="true" pageId="12" pageNumber="43">Remarks</emphasis>
: Only two males of
<taxonomicName id="4C244D0E2463FF89FD5DF9227864F979" box="[661,830,1676,1701]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="corrugatum">
<emphasis id="B950EA9F2463FF89FD5DF9227864F979" box="[661,830,1676,1701]" italics="true" pageId="12" pageNumber="43">T. corrugatum</emphasis>
</taxonomicName>
are known, from sites more than
<quantity id="4CDC9B682463FF89FB18F9257E71F979" box="[1232,1323,1675,1701]" metricMagnitude="5" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="43" unit="km" value="350.0">350 km</quantity>
apart, and while the specimens agree in other details, the tip of the telopodite turns distally in the
<typeStatus id="549F882F2463FF89FE96F9757A99F929" box="[350,451,1755,1781]" pageId="12" pageNumber="43" type="paratype">paratype</typeStatus>
(
<figureCitation id="131F2A082463FF89FE12F975797DF929" box="[474,551,1755,1781]" captionStart="FIGURE 5" captionStartId="11.[264,367,1102,1126]" captionTargetBox="[275,1299,482,1066]" captionTargetId="figure@11.[264,1322,466,1078]" captionTargetPageId="11" captionText="FIGURE 5. Tholerosoma monteithi n. gen., n. sp. (A) medial view of right gonopod, (D) antenna, (E) leg 6 (male paratype, QM S 73974). Tholerosoma corrugatum n. sp. (B) medial view of right gonopod (holotype, QM S 73993), (C) medial view of right gonopod (male paratype, QM S 73994), (F) outline sketch of midbody metazonite, showing location of swellings on posterior edge (female paratype, QM S 73995). Setation not shown; dashed lines in (A) – (C) indicate course of prostatic groove. Scale bars for (A) – (C) and for (D) and (E) = 0.25 mm." httpUri="https://zenodo.org/record/174578/files/figure.png" pageId="12" pageNumber="43">Fig. 5</figureCitation>
C). The two nominal species of
<taxonomicName id="4C244D0E2463FF89FC0EF9727F3AF929" box="[966,1120,1756,1781]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9F2463FF89FC0EF9727F3AF929" box="[966,1120,1756,1781]" italics="true" pageId="12" pageNumber="43">Tholerosoma</emphasis>
</taxonomicName>
are most easily distinguished by the absence of large paramedian swellings in
<taxonomicName id="4C244D0E2463FF89FBFDF8AA7FBCF8C1" box="[1077,1254,1796,1821]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="corrugatum">
<emphasis id="B950EA9F2463FF89FBFDF8AA7FBCF8C1" box="[1077,1254,1796,1821]" italics="true" pageId="12" pageNumber="43">T. corrugatum</emphasis>
</taxonomicName>
. The sympatry of the two species is probably syntopy: both were taken at a single collecting site in the Bluewater Range in 1986.
</paragraph>
</subSubSection>
</treatment>
</document>
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