treatments-xml/data/03/82/87/038287BBFFBDFF8E6A8AFBF0579FFE5E.xml

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<mods:title id="2583875E67F6CA6F66209DED7C416FB9">The genus Haplidia Hope in Lebanon (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini)</mods:title>
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<mods:namePart id="82A9A4702166E8E94F7C977BA67002F7">Uliana, Marco</mods:namePart>
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<mods:namePart id="685ED8937835F8E9569FDD6B0A09423B">Sehnal, Richard</mods:namePart>
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<paragraph id="8B9436ADFFBDFF8D6A8AFBF051D5FBC6" blockId="1.[151,1437,1052,2016]" box="[151,749,1052,1079]" pageId="1" pageNumber="537">
<heading id="D0DC81C1FFBDFF8D6A8AFBF051D5FBC6" bold="true" box="[151,749,1052,1079]" fontSize="11" level="1" pageId="1" pageNumber="537" reason="1">
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<taxonomicName id="4C2B4D2EFFBDFF8D6A8AFBF053C4FBC6" authorityName="Hope" authorityYear="1837" box="[151,252,1052,1079]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEABFFFBDFF8D6A8AFBF053C4FBC6" bold="true" box="[151,252,1052,1079]" italics="true" pageId="1" pageNumber="537">Haplidia</emphasis>
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and 
<taxonomicName id="4C2B4D2EFFBDFF8D6B2AFBF052E1FBC6" authorityName="Burmeister" authorityYear="1855" box="[311,473,1052,1079]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBDFF8D6B2AFBF052E1FBC6" bold="true" box="[311,473,1052,1079]" italics="true" pageId="1" pageNumber="537">Haplidia fissa</emphasis>
</taxonomicName>
, historical background
</emphasis>
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<paragraph id="8B9436ADFFBDFF8D6A8AFBAD5079F9DE" blockId="1.[151,1437,1052,2016]" pageId="1" pageNumber="537">
The genus name 
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<emphasis id="B95FEABFFFBDFF8D6B46FBAD5295FBAA" box="[347,429,1089,1115]" italics="true" pageId="1" pageNumber="537">Aplidia</emphasis>
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was introduced by 
<bibRefCitation id="EFBA4B5CFFBDFF8D688CFBAD5019FBAA" author="Hope, F. W." box="[657,801,1089,1115]" pageId="1" pageNumber="537" pagination="15 - 121" refId="ref7163" refString="Hope, F. W. (1837) The Coleopterist's Manual, containing the lamellicorn insects of Linneus and Fabricius. Henry G. Bohn, London, xiii + [2] + 15 - 121 + [4] pp., 3 pls." type="book chapter" year="1837">Hope (1837)</bibRefCitation>
to accommodate a single taxon, 
<taxonomicName id="4C2B4D2EFFBDFF8D6E81FBAD527FFB8E" authority="Fabricius, 1801" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="transversa">
<emphasis id="B95FEABFFFBDFF8D6E81FBAD56A4FBAA" box="[1180,1436,1089,1115]" italics="true" pageId="1" pageNumber="537">Melolontha transversa</emphasis>
<bibRefCitation id="EFBA4B5CFFBDFF8D6A8AFB89527FFB8E" author="Fabricius, J. C." box="[151,327,1125,1151]" pageId="1" pageNumber="537" refId="ref7030" refString="Fabricius, J. C. (1801) Systema Eleutheratorum secundum ordines, genera, species: adiectis synonymis, locis, observationibus, descriptionibus. Tomus II. Bibliopoli Academici Novi, Kiliae, 687 pp." type="book" year="1801">Fabricius, 1801</bibRefCitation>
</taxonomicName>
, 
<typeStatus id="5490880FFFBDFF8D6B4FFB8952BAFB8E" box="[338,386,1125,1151]" pageId="1" pageNumber="537">type</typeStatus>
species of the genus. 
<bibRefCitation id="EFBA4B5CFFBDFF8D6869FB89507AFB8E" author="Burmeister, H. C. C." box="[628,834,1124,1151]" pageId="1" pageNumber="537" refId="ref6743" refString="Burmeister, H. C. C. (1855) Handbuch der Entomologie. Vierter Band. Besondere Entomologie. Fortsetzung. Zweite Abtheilung. Coleoptera Lamellicornia Phyllophaga chaenochela. Theod. Chr. Friedr. Enslin, Berlin, x + 569 pp." type="book" year="1855">Burmeister (1855)</bibRefCitation>
recognized a second species, 
<taxonomicName id="4C2B4D2EFFBDFF8D6E93FB895622FB8E" authorityName="Burmeister" authorityYear="1855" box="[1166,1306,1125,1151]" class="Insecta" family="Scarabaeidae" genus="Aplidia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBDFF8D6E93FB895622FB8E" box="[1166,1306,1125,1151]" italics="true" pageId="1" pageNumber="537">Aplidia fissa</emphasis>
</taxonomicName>
, based on a Syrian—Mesopotamian female specimen whose re-examination was apparently neglected by all following authors. As a matter of fact, various subsequent identifications, descriptions of new species, and compilations of systematics catalogues for the genus seem to have been often based on literature alone, without re-examination of specimens, namely 
<typeStatus id="5490880FFFBDFF8D6AEEFB19520AFAFE" box="[243,306,1269,1295]" pageId="1" pageNumber="537">types</typeStatus>
. One of such catalogues (
<bibRefCitation id="EFBA4B5CFFBDFF8D6844FB1951D2FAE1" author="Harold, E. von" box="[601,746,1269,1296]" pageId="1" pageNumber="537" pagination="979 - 1346" refId="ref7107" refString="Harold, E. von (1869) [new names]. In: Gemminger, M. &amp; Harold, B. von: Catalogus coleopterorum hucusque descriptorum synonymicus et systematicus. IV. Scarabaeidae. E. H. Gummi, Monachii, pp. 979 - 1346 + [7]." type="journal article" year="1869">Harold 1869</bibRefCitation>
) also introduced the incorrect subsequent spelling 
<taxonomicName id="4C2B4D2EFFBDFF8D6F2FFB1956AEFAFE" authorityName="Hope" authorityYear="1837" box="[1330,1430,1269,1295]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEABFFFBDFF8D6F2FFB1956AEFAFE" box="[1330,1430,1269,1295]" italics="true" pageId="1" pageNumber="537">Haplidia</emphasis>
</taxonomicName>
, which was retained for over a century (
<emphasis id="B95FEABFFFBDFF8D684DFAF65140FAC2" box="[592,632,1306,1331]" italics="true" pageId="1" pageNumber="537">e.g.</emphasis>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D689BFAF5502DFAC2" author="Reitter, E." box="[646,789,1305,1331]" pageId="1" pageNumber="537" pagination="93 - 303" refId="ref7798" refString="Reitter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europaischen fauna und angrenzenden landern, enthaltend die gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn, 40, 93 - 303. [1901]" type="journal article" year="1902">Reitter 1902</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D693CFAF55089FAC2" author="Grandi, G." box="[801,945,1304,1331]" pageId="1" pageNumber="537" pagination="159 - 224" refId="ref7069" refString="Grandi, G. (1925) Contribute alla conoscenza biologica e morfologica di alcuni Lamellicorni fillofagi. Bollettino del Laboratorio di Zoologia Generale e Agraria della Facolta Agraria in Portici, 18, 159 - 224." type="journal article" year="1925">Grandi 1925</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D69A0FAF5575CFAC2" author="Petrovitz, R." box="[957,1124,1304,1331]" pageId="1" pageNumber="537" pagination="204 - 206" refId="ref7648" refString="Petrovitz, R. (1958) Eine neue Haplidia - Art aus Kleinasien (Coleopt., Scarabaeidae). Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 97, 204 - 206." type="journal article" year="1958">Petrovitz 1958</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6E72FAF55792FAC2" author="Petrovitz, R." box="[1135,1194,1305,1331]" pageId="1" pageNumber="537" pagination="325 - 343" refId="ref7692" refString="Petrovitz, R. (1967) Ergebnisse zoologischer sammelreisen in der Turkei. Annalen des Naturhistorischen Museums in Wien, 70, 325 - 343." type="journal article" year="1967">1967</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6EA8FAF557C8FAC2" author="Petrovitz, R." box="[1205,1264,1305,1331]" pageId="1" pageNumber="537" pagination="38 - 45" refId="ref7719" refString="Petrovitz, R. (1970) Scarabaeidae aus Iran. Entomologie et Phytopathologie Appliquees, 29, 38 - 45." type="journal article" year="1970">1970</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6EE1FAF5567AFAC2" author="Petrovitz, R." box="[1276,1346,1305,1331]" pageId="1" pageNumber="537" pagination="215 - 238" refId="ref7741" refString="Petrovitz, R. (1971 a) Scarabaeidae from the Near East. Israel Journal of Entomology, 6, 215 - 238." type="journal article" year="1971">1971a</bibRefCitation>
, b; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6F76FAF5522BFAA6" author="Viggiani, G." pageId="1" pageNumber="537" pagination="89 - 152" refId="ref8037" refString="Viggiani, G. (1969) Richerche sulla entomofauna del Nocciulo. 1. Contributo alla conoscenza della Haplidia etrusca Kr. (Col. Scarabaeidae). Note morfoetologiche e mezzi di lotta. Bollettino del Laboratorio di Entomologia Agraria Filippo Silvestri, 27, 89 - 152." type="journal article" year="1969">Viggiani 1969</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6B02FAD15262FAA6" author="Viggiani, G." box="[287,346,1341,1367]" pageId="1" pageNumber="537" pagination="11 - 15" refId="ref8088" refString="Viggiani, G. (1977) Nuovi reperti su alcuni coleotteri radicicoli (Haplidia etrusca Kr., Otiorrhynchus armatus Boh., O. trophonius Reitt.). Bollettino del Laboratorio di Entomologia Agraria Filippi Silvestri, 34, 11 - 15." type="journal article" year="1977">1977</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6B7AFAD15126FAA6" author="Carpaneto, G. M." box="[359,542,1341,1367]" pageId="1" pageNumber="537" pagination="111 - 126" refId="ref6789" refString="Carpaneto, G. M. (1979) Observazioni sui coleotteri Scarabaeoidea delle Isole Ponziane. Fragmenta Entomologica, 15, 111 - 126." type="journal article" year="1979">Carpaneto 1979</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6836FAD15187FAA6" author="Baraud, J." box="[555,703,1340,1367]" pageId="1" pageNumber="537" pagination="191 - 196" refId="ref6465" refString="Baraud, J. (1975) Descriptions de cinq nouvelles especes palearctiques de Coleopteres Scarabaeoidea. Nouvelle Revue d'Entomologie, 5, 191 - 196." type="journal article" year="1975">Baraud 1975</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D68D1FAD1503FFAA6" author="Baraud, J." box="[716,775,1341,1367]" pageId="1" pageNumber="537" pagination="1 - 352" refId="ref6492" refString="Baraud, J. (1977) Coleopteres Scarabaeoidea. Faune de l'Europe occidentale Belgique-France-Grande-Bretagne-Italie- Peninsule Iberique. Nouvelle Revue d'Entomologie, 7 (Supplement 4), 1 - 352." type="journal article" year="1977">1977</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D690EFAD15076FAA6" author="Baraud, J." box="[787,846,1341,1367]" pageId="1" pageNumber="537" pagination="539 - 580" refId="ref6524" refString="Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672" type="journal article" year="1988">1988</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6947FAD150ADFAA6" author="Baraud, J." box="[858,917,1341,1367]" pageId="1" pageNumber="537" pagination="159 - 162" refId="ref6569" refString="Baraud, J. (1990) Description de deux nouvelles especes du genre Haplidia Hope (Coleoptera, Scarabaeoidea, Melolonthidae). Nouvelle Revue d'Entomologie, 7, 159 - 162." type="journal article" year="1990">1990</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D69BFFAD150E5FAA6" author="Baraud, J." box="[930,989,1341,1367]" pageId="1" pageNumber="537" refId="ref6603" refString="Baraud, J. (1992) Coleopteres Scarabaeoidea d'Europe. Faune de France 78. Societe Lineenne de Lyon, Lyon, xi + 856 pp. https: // doi. org / 10.3406 / linly. 1992.11007" type="book" year="1992">1992</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D69F7FAD1579DFAA6" author="Sabatinelli, G." box="[1002,1189,1341,1367]" pageId="1" pageNumber="537" pagination="5 - 50" refId="ref7842" refString="Sabatinelli, G. (1991) Note sul genere Haplidia Burm. con descizione di H. baraudi n. sp. Bollettino della Societa Entomologica Italiana, 123, 4 5 - 50." type="journal article" year="1991">Sabatinelli 1991</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6EAEFAD1522CFA8A" author="Coca Abia, M. M. &amp; Martin Piera, F." pageId="1" pageNumber="537" pagination="69 - 77" refId="ref6903" refString="Coca Abia, M. M. &amp; Martin Piera, F. (1993) The genus Haplidia Hope, 1837 (Coleoptera, Melolonthidae) in the Iberian Peninsula. Nouvelle Revue d'Entomologie, 10, 69 - 77." type="journal article" year="1993">Coca Abia &amp; Martìn Piera 1993</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6B3CFA8D5297FA8A" author="Keith, D." box="[289,431,1377,1403]" pageId="1" pageNumber="537" pagination="380 - 387" refId="ref7237" refString="Keith, D. (2000 a) Contribution a la connaissance des Scarabaeoidea du Moyen Orient: taxons nouveaux ou peu connus (Coleoptera). Lambillionea, 100, 380 - 387." type="journal article" year="2000">Keith 2000a</bibRefCitation>
, b; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6BC7FA8D51ADFA8A" author="Coca Abia, M. M." box="[474,661,1377,1403]" pageId="1" pageNumber="537" pagination="220 - 227" refId="ref6852" refString="Coca Abia, M. M. (2004) External morphology and genitalia of Melolontha melolontha (L., 1758) (Coleoptera, Scarabaeidae, Melolonthinae). A comparative study with other genera of the Iberian Peninsula. Laimburg Journal, 1, 220 - 227." type="journal article" year="2004">Coca Abia 2004</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D68BCFA8D5007FA8A" author="Pesarini, C." box="[673,831,1377,1403]" pageId="1" pageNumber="537" pagination="1 - 132" refId="ref7626" refString="Pesarini, C. (2004) Insetti della Fauna Italiana Coleotteri Lamellicorni. Natura, 93, 1 - 132." type="journal article" year="2004">Pesarini 2004</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6951FA8D5751FA8A" author="Keith, D. &amp; Lo Cascio, P." box="[844,1129,1376,1403]" pageId="1" pageNumber="537" pagination="139 - 142" refId="ref7375" refString="Keith, D. &amp; Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142." type="journal article" year="2005">Keith &amp; Lo Cascio 2005</bibRefCitation>
). The original spelling was reintroduced by recent authors such as 
<bibRefCitation id="EFBA4B5CFFBDFF8D685AFA6950A7FA6E" author="Chikatunov, V. &amp; Pavlicek, T." box="[583,927,1413,1439]" pageId="1" pageNumber="537" pagination="37 - 65" refId="ref6815" refString="Chikatunov, V. &amp; Pavlicek, T. (1997) Catalogue of the beetles (Coleoptera) in Israel and adjacent areas: 1. Scarabaeoidea. Klapalekiana, 33, 37 - 65." type="journal article" year="1997">Chikatunov &amp; Pavlíček (1997)</bibRefCitation>
, followed by 
<bibRefCitation id="EFBA4B5CFFBDFF8D6E2BFA695606FA51" author="Smetana, A. &amp; Kral, D." box="[1078,1342,1413,1440]" pageId="1" pageNumber="537" pagination="207 - 228" refId="ref7942" refString="Smetana, A. &amp; Kral, D. (2006) Rhizotrogini. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 207 - 228." type="book chapter" year="2006">Smetana &amp; Král (2006)</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6F55FA6953DAFA35" author="Bezdek, A." pageId="1" pageNumber="537" pagination="249 - 280" refId="ref6643" refString="Bezdek, A. (2016) Rhizotrogini. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, Boston, pp. 249 - 280." type="book chapter" year="2016">Bezděk (2016)</bibRefCitation>
and in recent contributions (
<emphasis id="B95FEABFFFBDFF8D6834FA46516AFA32" box="[553,594,1450,1475]" italics="true" pageId="1" pageNumber="537">e.g.</emphasis>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D687CFA4551D1FA32" author="Boyer, J. - C." box="[609,745,1449,1475]" pageId="1" pageNumber="537" pagination="39 - 40" refId="ref6703" refString="Boyer, J. - C. (2014) Aplidia transversa transversa (F. 1801), nouveau pour la faune de France (Coleoptera Scarabaeidae Melolonthinae Rhizotrogini). L'Entomologiste, 70, 39 - 40." type="journal article" year="2014">Boyer 2014</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D68E8FA4550AEFA32" author="Miessen, G." box="[757,918,1449,1475]" pageId="1" pageNumber="537" pagination="57 - 73" refId="ref7539" refString="Miessen, G. (2016) Essais sur le genre Aplidia Hope 1837 a Chypre et commentaires sur quelques especes limitrophes (Coleoptera, Scarabaeoidea, Melolonthidae, Rhizotrogini). Entomologie Faunistique-Faunistic Entomology, 69, 57 - 73." type="journal article" year="2016">Miessen 2016</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D69BFFA45574EFA32" author="Soldati, F. &amp; Noblecourt, T. &amp; Barnouin, T." box="[930,1142,1449,1475]" pageId="1" pageNumber="537" pagination="371 - 372" refId="ref7993" refString="Soldati, F., Noblecourt, T. &amp; Barnouin, T. (2016) Presence d' Aplidia transversa (F., 1801) dans les Alpes-Maritimes (Coleoptera Scarabaeidae Melolonthinae). L'Entomologiste, 72, 371 - 372." type="journal article" year="2016">
Soldati 
<emphasis id="B95FEABFFFBDFF8D69E4FA46570BFA32" box="[1017,1075,1449,1475]" italics="true" pageId="1" pageNumber="537">et al.</emphasis>
2016
</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6E9FFA45562BFA32" author="Mienis, H. K." box="[1154,1299,1449,1475]" pageId="1" pageNumber="537" pagination="3 - 4" refId="ref7502" refString="Mienis, H. K. (2017) Activity pattern of the Cockchafer Aplidia chaifensis in Kibbutz Netzer Sereni, Israel, in 2017. Natuurhistorische en Andere Notities, 14, 3 - 4." type="journal article" year="2017">Mienis 2017</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6F3DFA455201FA16" author="Lambert, G. &amp; Lemaire, J. - M." pageId="1" pageNumber="537" pagination="13 - 14" refId="ref7453" refString="Lambert, G. &amp; Lemaire, J. - M. (2018) Confirmation de la presence du hanneton Aplidia transversa (Fabricius, 1801) (Coleoptera Scarabaeidae Melolonthinae) dans les Alpes-Maritimes (France). Biocosme Mesogeen, 35, 13 - 14." type="journal article" year="2018">Lambert &amp; Lemaire 2018</bibRefCitation>
). However, based on Article 33.2.3.1 (
<bibRefCitation id="EFBA4B5CFFBDFF8D68F7FA215049FA16" author="ICZN" box="[746,881,1485,1511]" pageId="1" pageNumber="537" refId="ref7211" refString="ICZN, (1999) International Code of Zoological Nomenclature. 4 th Edition. International trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
), the spelling 
<taxonomicName id="4C2B4D2EFFBDFF8D6E12FA21574BFA16" authorityName="Hope" authorityYear="1837" box="[1039,1139,1485,1511]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEABFFFBDFF8D6E12FA21574BFA16" box="[1039,1139,1485,1511]" italics="true" pageId="1" pageNumber="537">Haplidia</emphasis>
</taxonomicName>
should be retained as correct, as being in prevailing usage and attributed to the original author and date. Consequently, 
<taxonomicName id="4C2B4D2EFFBDFF8D6EA7FA1D5660F9FA" authorityName="Burmeister" authorityYear="1855" box="[1210,1368,1521,1547]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBDFF8D6EA7FA1D5660F9FA" box="[1210,1368,1521,1547]" italics="true" pageId="1" pageNumber="537">Haplidia fissa</emphasis>
</taxonomicName>
is the original combination for Burmeister’s taxon (Article 33.3.1).
</paragraph>
</subSubSection>
<subSubSection id="C3316526FFBDFF8E6ADAF9D5579FFE5E" lastPageId="2" lastPageNumber="538" pageId="1" pageNumber="537" type="description">
<paragraph id="8B9436ADFFBDFF8D6ADAF9D55641F8F6" blockId="1.[151,1437,1052,2016]" pageId="1" pageNumber="537">
Forty-five years after the introduction of 
<taxonomicName id="4C2B4D2EFFBDFF8D688FF9D550A6F9A2" authority=", Kraatz (1882)" authorityName=", Kraatz" authorityYear="1882" box="[658,926,1593,1619]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEABFFFBDFF8D688FF9D551CDF9A2" box="[658,757,1593,1619]" italics="true" pageId="1" pageNumber="537">Haplidia</emphasis>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D691FF9D550A6F9A2" author="Kraatz, G." box="[770,926,1593,1619]" pageId="1" pageNumber="537" pagination="33 - 42" refId="ref7415" refString="Kraatz, G. (1882) Revision der europaisch-syrischen Arten der Melolonthiden-Gattung Haplidia Hope. Deutsche Entomologische Zeitschrift, 26, 33 - 42. https: // doi. org / 10.1002 / mmnd. 48018820202" type="journal article" year="1882">Kraatz (1882)</bibRefCitation>
</taxonomicName>
published the first comprehensive revision of the genus. Regarding 
<taxonomicName id="4C2B4D2EFFBDFF8D6B97F9B1511FF986" authorityName="Burmeister" authorityYear="1855" box="[394,551,1629,1655]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBDFF8D6B97F9B1511FF986" box="[394,551,1629,1655]" italics="true" pageId="1" pageNumber="537">Haplidia fissa</emphasis>
</taxonomicName>
, he identified as such some female specimens from 
<collectingCountry id="F33C763DFFBDFF8D6E70F9B157F6F986" box="[1133,1230,1629,1655]" name="Lebanon" pageId="1" pageNumber="537">Lebanon</collectingCountry>
(
<collectingRegion id="49EFF84FFFBDFF8D6EC1F9B1567FF989" box="[1244,1351,1629,1656]" country="Lebanon" name="Beyrouth" pageId="1" pageNumber="537">Beyrouth</collectingRegion>
) that in his opinion best agreed with the description of the Syrian—Mesopotamian female of 
<taxonomicName id="4C2B4D2EFFBDFF8D6E21F96E57B7F96A" authorityName="Burmeister" authorityYear="1855" box="[1084,1167,1665,1691]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBDFF8D6E21F96E57B7F96A" box="[1084,1167,1665,1691]" italics="true" pageId="1" pageNumber="537">H. fissa</emphasis>
</taxonomicName>
, and used the associated male specimens for a supplemental description of the species, thus providing the basis for the misidentifications that followed. Subsequent authors of descriptive and faunistic articles, revisions and catalogues followed 
<bibRefCitation id="EFBA4B5CFFBDFF8D6EEDF92556A4F912" author="Kraatz, G." box="[1264,1436,1737,1763]" pageId="1" pageNumber="537" pagination="33 - 42" refId="ref7415" refString="Kraatz, G. (1882) Revision der europaisch-syrischen Arten der Melolonthiden-Gattung Haplidia Hope. Deutsche Entomologische Zeitschrift, 26, 33 - 42. https: // doi. org / 10.1002 / mmnd. 48018820202" type="journal article" year="1882">Kraatz’s (1882)</bibRefCitation>
interpretation (
<emphasis id="B95FEABFFFBDFF8D6B20F902525DF8F6" box="[317,357,1774,1799]" italics="true" pageId="1" pageNumber="537">e.g.</emphasis>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6B6EF9015138F8F6" author="Reitter, E." box="[371,512,1773,1799]" pageId="1" pageNumber="537" pagination="93 - 303" refId="ref7798" refString="Reitter, E. (1902) Bestimmungs-Tabelle der Melolonthidae aus der europaischen fauna und angrenzenden landern, enthaltend die gruppen der Pachydemini, Sericini und Melolonthini. Verhandlungen des Naturforschenden Vereines in Brunn, 40, 93 - 303. [1901]" type="journal article" year="1902">Reitter 1902</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6816F90151F3F8F6" author="Dalla Torre, K. W. von" box="[523,715,1773,1799]" pageId="1" pageNumber="537" pagination="135 - 290" refId="ref6945" refString="Dalla Torre, K. W. von. (1912) Pars 49. Scarabaeidae: Melolonthidae III. In: Junk W. &amp; Schenkling S. (Eds): Coleopterorum Catalogus. Vol. XX. W. Junk, Berlin, pp. 135 - 290." type="book chapter" year="1912">Dalla Torre 1912</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D68CBF9015043F8F6" author="Petrovitz, R." box="[726,891,1773,1799]" pageId="1" pageNumber="537" pagination="325 - 343" refId="ref7692" refString="Petrovitz, R. (1967) Ergebnisse zoologischer sammelreisen in der Turkei. Annalen des Naturhistorischen Museums in Wien, 70, 325 - 343." type="journal article" year="1967">Petrovitz 1967</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D699BF90150F8F8F6" author="Petrovitz, R." box="[902,960,1773,1799]" pageId="1" pageNumber="537" pagination="38 - 45" refId="ref7719" refString="Petrovitz, R. (1970) Scarabaeidae aus Iran. Entomologie et Phytopathologie Appliquees, 29, 38 - 45." type="journal article" year="1970">1970</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D69D1F901572AF8F6" author="Petrovitz, R." box="[972,1042,1773,1799]" pageId="1" pageNumber="537" pagination="215 - 238" refId="ref7741" refString="Petrovitz, R. (1971 a) Scarabaeidae from the Near East. Israel Journal of Entomology, 6, 215 - 238." type="journal article" year="1971">1971a</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6E03F9015788F8F6" author="Baraud, J." box="[1054,1200,1772,1799]" pageId="1" pageNumber="537" pagination="191 - 196" refId="ref6465" refString="Baraud, J. (1975) Descriptions de cinq nouvelles especes palearctiques de Coleopteres Scarabaeoidea. Nouvelle Revue d'Entomologie, 5, 191 - 196." type="journal article" year="1975">Baraud 1975</bibRefCitation>
; 
<bibRefCitation id="EFBA4B5CFFBDFF8D6EA6F9015655F8F6" author="Montreuil, O." box="[1211,1389,1772,1799]" pageId="1" pageNumber="537" pagination="129 - 131" refId="ref7581" refString="Montreuil, O. (2005) Contribution a l'etude du genre Haplidia Hope: description de deux nouvelles especes du Liban (Coleoptera, Melolonthinae). Bulletin de la Societe entomologique de France, 110 (2), 129 - 131." type="journal article" year="2005">Montreuil 2005</bibRefCitation>
).
</paragraph>
<paragraph id="8B9436ADFFBDFF8D6ADAF8FD5276F811" blockId="1.[151,1437,1052,2016]" pageId="1" pageNumber="537">
Also 
<bibRefCitation id="EFBA4B5CFFBDFF8D6B1BF8FD5285F8DA" author="Baraud, J." box="[262,445,1809,1835]" pageId="1" pageNumber="537" pagination="539 - 580" refId="ref6524" refString="Baraud, J. (1988) Contribution a l'etude du genre Haplidia Hope (Coleoptera Scarabaeoidea Melolonthidae). Revue Suisse de Zoologie, 95, 539 - 580. https: // doi. org / 10.5962 / bhl. part. 79672" type="journal article" year="1988">Baraud’s (1988)</bibRefCitation>
revision of the whole genus maintained the identification of the species sensu 
<bibRefCitation id="EFBA4B5CFFBDFF8D6F4FF8FD53DDF8BE" author="Kraatz, G." pageId="1" pageNumber="537" pagination="33 - 42" refId="ref7415" refString="Kraatz, G. (1882) Revision der europaisch-syrischen Arten der Melolonthiden-Gattung Haplidia Hope. Deutsche Entomologische Zeitschrift, 26, 33 - 42. https: // doi. org / 10.1002 / mmnd. 48018820202" type="journal article" year="1882">Kraatz (1882)</bibRefCitation>
. This revision included the study of the Kraatz and Petrovitz collections and the redescription of 
<taxonomicName id="4C2B4D2EFFBDFF8D6F24F8D953F2F882" authorityName="Burmeister" authorityYear="1855" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBDFF8D6F24F8D953F2F882" italics="true" pageId="1" pageNumber="537">Haplidia fissa</emphasis>
</taxonomicName>
male (sensu Kraatz), but not an examination of the 
<typeStatus id="5490880FFFBDFF8D690BF8B5507EF882" box="[790,838,1881,1907]" pageId="1" pageNumber="537">type</typeStatus>
in Burmeister’s collection. Kraatz’s concept of 
<taxonomicName id="4C2B4D2EFFBDFF8D6F7EF8B65230F866" authorityName="Burmeister" authorityYear="1855" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="537" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBDFF8D6F7EF8B65230F866" italics="true" pageId="1" pageNumber="537">Haplidia fissa</emphasis>
</taxonomicName>
was therefore reconfirmed and, as such, accepted also by 
<bibRefCitation id="EFBA4B5CFFBDFF8D69AFF89157B9F866" author="Sabatinelli, G." box="[946,1153,1917,1943]" pageId="1" pageNumber="537" pagination="5 - 50" refId="ref7842" refString="Sabatinelli, G. (1991) Note sul genere Haplidia Burm. con descizione di H. baraudi n. sp. Bollettino della Societa Entomologica Italiana, 123, 4 5 - 50." type="journal article" year="1991">Sabatinelli (1991)</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6E92F89153DCF84A" author="Chikatunov, V. &amp; Pavlicek, T." pageId="1" pageNumber="537" pagination="37 - 65" refId="ref6815" refString="Chikatunov, V. &amp; Pavlicek, T. (1997) Catalogue of the beetles (Coleoptera) in Israel and adjacent areas: 1. Scarabaeoidea. Klapalekiana, 33, 37 - 65." type="journal article" year="1997">Chikatunov &amp; Pavlíček (1997)</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6AF2F84D52BDF84A" author="Keith, D." box="[239,389,1953,1979]" pageId="1" pageNumber="537" pagination="380 - 387" refId="ref7237" refString="Keith, D. (2000 a) Contribution a la connaissance des Scarabaeoidea du Moyen Orient: taxons nouveaux ou peu connus (Coleoptera). Lambillionea, 100, 380 - 387." type="journal article" year="2000">Keith (2000a</bibRefCitation>
, b), 
<bibRefCitation id="EFBA4B5CFFBDFF8D6BA9F84D51E3F84A" author="Keith, D. &amp; Lo Cascio, P." box="[436,731,1952,1979]" pageId="1" pageNumber="537" pagination="139 - 142" refId="ref7375" refString="Keith, D. &amp; Lo Cascio, P. (2005) A new Haplidia Hope from Nisyros Island (Dodecanese, Greece) with a catalogue of the genus. Lambillionea, 105, 139 - 142." type="journal article" year="2005">Keith &amp; Lo Cascio (2005)</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D68FBF84D5091F84A" author="Montreuil, O." box="[742,937,1952,1979]" pageId="1" pageNumber="537" pagination="129 - 131" refId="ref7581" refString="Montreuil, O. (2005) Contribution a l'etude du genre Haplidia Hope: description de deux nouvelles especes du Liban (Coleoptera, Melolonthinae). Bulletin de la Societe entomologique de France, 110 (2), 129 - 131." type="journal article" year="2005">Montreuil (2005)</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D69A8F84D5785F84D" author="Smetana, A. &amp; Kral, D." box="[949,1213,1953,1980]" pageId="1" pageNumber="537" pagination="207 - 228" refId="ref7942" refString="Smetana, A. &amp; Kral, D. (2006) Rhizotrogini. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera, Volume 3. Scarabaeoidea-Scirtoidea-Dascilloidea-Buprestoidea-Byrrhoidea. Apollo Books, Stenstrup, pp. 207 - 228." type="book chapter" year="2006">Smetana &amp; Král (2006)</bibRefCitation>
, 
<bibRefCitation id="EFBA4B5CFFBDFF8D6ED5F84D5656F84D" author="Bezdek, A." box="[1224,1390,1953,1980]" pageId="1" pageNumber="537" pagination="249 - 280" refId="ref6643" refString="Bezdek, A. (2016) Rhizotrogini. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 3. Scarabaeoidea, Scirtoidea, Dascilloidea, Buprestoidea, Byrrhoidea. Revised and updated edition. Brill, Leiden, Boston, pp. 249 - 280." type="book chapter" year="2016">Bezděk (2016)</bibRefCitation>
and 
<bibRefCitation id="EFBA4B5CFFBDFF8D6A8AF8295271F811" author="Miessen, G." box="[151,329,1989,2016]" pageId="1" pageNumber="537" pagination="57 - 73" refId="ref7539" refString="Miessen, G. (2016) Essais sur le genre Aplidia Hope 1837 a Chypre et commentaires sur quelques especes limitrophes (Coleoptera, Scarabaeoidea, Melolonthidae, Rhizotrogini). Entomologie Faunistique-Faunistic Entomology, 69, 57 - 73." type="journal article" year="2016">Miessen (2016)</bibRefCitation>
.
</paragraph>
<paragraph id="8B9436ADFFBEFF8E6ADAFF7551A5FE96" blockId="2.[151,1437,153,431]" pageId="2" pageNumber="538">
After examination of the female 
<typeStatus id="5490880FFFBEFF8E682FFF75515AFF42" box="[562,610,153,179]" pageId="2" pageNumber="538">type</typeStatus>
of 
<taxonomicName id="4C2B4D2EFFBEFF8E689AFF75501CFF42" authorityName="Burmeister" authorityYear="1855" box="[647,804,153,179]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="538" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBEFF8E689AFF75501CFF42" box="[647,804,153,179]" italics="true" pageId="2" pageNumber="538">Haplidia fissa</emphasis>
</taxonomicName>
that we could locate in Burmeister’s collection we could ascertain (Sehnal, in press) that it belongs to the genus 
<taxonomicName id="4C2B4D2EFFBEFF8E691EFF51571DFF26" authority="Keith, 2001" authorityName="Keith" authorityYear="2001" box="[771,1061,189,215]" class="Insecta" family="Melolonthidae" genus="Medeotrogus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEABFFFBEFF8E691EFF5150AEFF26" box="[771,918,189,215]" italics="true" pageId="2" pageNumber="538">Medeotrogus</emphasis>
<bibRefCitation id="EFBA4B5CFFBEFF8E6980FF51571DFF26" author="Keith, D." box="[925,1061,189,215]" pageId="2" pageNumber="538" pagination="75 - 78" refId="ref7308" refString="Keith, D. (2001) Contribution a la connaissance des Scarabaeoidea du Proche et du Moyen-Orient (9 e note). Description de Medeotrogus linnavuorii n. gen., n. sp. d'Irak et d'Iran (Col. Melolonthidae) et d' Adoretus kashmarensis, n. sp. d'Iran. Symbioses, Nouvelle Serie, 5, 75 - 78." type="journal article" year="2001">Keith, 2001</bibRefCitation>
</taxonomicName>
, recently described on specimens from 
<collectingCountry id="F33C763DFFBEFF8E6AC9FF0D5239FF0A" box="[212,257,225,251]" name="Iraq" pageId="2" pageNumber="538">Iraq</collectingCountry>
and 
<collectingCountry id="F33C763DFFBEFF8E6B2AFF0D5250FF0A" box="[311,360,225,251]" name="Iran" pageId="2" pageNumber="538">Iran</collectingCountry>
. Among characters of the genus are some that are similar to 
<taxonomicName id="4C2B4D2EFFBEFF8E6E14FF0D5755FF0A" authorityName=", Kraatz" authorityYear="1882" box="[1033,1133,225,251]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="538" phylum="Arthropoda" rank="genus">
<emphasis id="B95FEABFFFBEFF8E6E14FF0D5755FF0A" box="[1033,1133,225,251]" italics="true" pageId="2" pageNumber="538">Haplidia</emphasis>
</taxonomicName>
, such as emarginated clypeus, integuments ranging from glossy to matte chestnut brown, the male body symmetrically elongate, whereas in females the posterior part of the elytra is markedly wider. Such similarity, and the lack of study of the primary 
<typeStatus id="5490880FFFBEFF8E6F78FEC556A0FEB2" box="[1381,1432,297,323]" pageId="2" pageNumber="538">type</typeStatus>
, caused the misinterpretation of 
<taxonomicName id="4C2B4D2EFFBEFF8E6BE5FEA151AEFE96" authorityName="Burmeister" authorityYear="1855" box="[504,662,333,359]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="538" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBEFF8E6BE5FEA151AEFE96" box="[504,662,333,359]" italics="true" pageId="2" pageNumber="538">Haplidia fissa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B9436ADFFBEFF8E6ADAFE9D579FFE5E" blockId="2.[151,1437,153,431]" pageId="2" pageNumber="538">
As no name is available for 
<taxonomicName id="4C2B4D2EFFBEFF8E681CFE9D50B1FE7A" authority="sensu Kraatz (1882)" authorityName="sensu Kraatz" authorityYear="1882" box="[513,905,369,395]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="538" phylum="Arthropoda" rank="species" species="fissa">
<emphasis id="B95FEABFFFBEFF8E681CFE9D51A7FE7A" box="[513,671,369,395]" italics="true" pageId="2" pageNumber="538">Haplidia fissa</emphasis>
sensu 
<bibRefCitation id="EFBA4B5CFFBEFF8E68F6FE9D50B1FE7A" author="Kraatz, G." box="[747,905,369,395]" pageId="2" pageNumber="538" pagination="33 - 42" refId="ref7415" refString="Kraatz, G. (1882) Revision der europaisch-syrischen Arten der Melolonthiden-Gattung Haplidia Hope. Deutsche Entomologische Zeitschrift, 26, 33 - 42. https: // doi. org / 10.1002 / mmnd. 48018820202" type="journal article" year="1882">Kraatz (1882)</bibRefCitation>
</taxonomicName>
, we propose for the species commonly referred as such the new name 
<taxonomicName id="4C2B4D2EFFBEFF8E6B8FFE795736FE5E" authority="Sehnal, Montreuil &amp; Uliana" authorityName="Sehnal, Montreuil &amp; Uliana" authorityYear="2020" box="[402,1038,405,431]" class="Insecta" family="Melolonthidae" genus="Haplidia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="538" phylum="Arthropoda" rank="species" species="karlaeschneiderae" status="sp. nov.">
<emphasis id="B95FEABFFFBEFF8E6B8FFE7951F4FE5E" box="[402,716,405,431]" italics="true" pageId="2" pageNumber="538">Haplidia karlaeschneiderae</emphasis>
Sehnal, Montreuil &amp; Uliana
</taxonomicName>
, 
<taxonomicNameLabel id="A26C57C4FFBEFF8E6E04FE795798FE5E" box="[1049,1184,405,431]" pageId="2" pageNumber="538" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
</treatment>
</document>