165 lines
30 KiB
XML
165 lines
30 KiB
XML
<document id="8E3B021D66749ED8255089D3F8BEBF3C" ID-CLB-Dataset="293303" ID-DOI="10.1080/14772019.2021.1960911" ID-GBIF-Dataset="53749253-12aa-41cc-9eec-e28c0b075738" ID-ISSN="1478-0941" ID-Zenodo-Dep="10949919" ID-ZooBank="F8991F09-B5FB-40EF-B4CC-474D925085B8" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1711484962835" checkinUser="felipe" docAuthor="Gale, Andrew Scott" docDate="2021" docId="B9207C419A69FFE40E20FC79FB45FC34" docLanguage="en" docName="JSystPalaeontol.19.10.691-741.pdf" docOrigin="Journal of Systematic Palaeontology 19 (10)" docSource="http://dx.doi.org/10.1080/14772019.2021.1960911" docStyle="DocumentStyle:6400638016C1F4AE0D0B6502E032D571.1:JSystPalaeontol.2016-.journal_article.closed" docStyleId="6400638016C1F4AE0D0B6502E032D571" docStyleName="JSystPalaeontol.2016-.journal_article.closed" docStyleVersion="1" docTitle="Pouzaster pocknotata Gale 2021, sp. nov." docType="treatment" docVersion="5" lastPageNumber="721" masterDocId="451904399A76FFFB0F0FFF8CFF93FF93" masterDocTitle="Taxonomy and phylogeny of the ‘ football stars’ (Asteroidea, Sphaerasteridae)" masterLastPageNumber="741" masterPageNumber="691" pageNumber="721" updateTime="1712755325394" updateUser="ExternalLinkService">
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<mods:title id="210A877CD3BA2E0B7E3AF6291806909B">Taxonomy and phylogeny of the ‘ football stars’ (Asteroidea, Sphaerasteridae)</mods:title>
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<mods:name id="8AF6540E0489E3E2DEAD0CCDA9038BCB" type="personal">
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<mods:namePart id="9D8C859A4D6FB1602F2B39317015F568">Gale, Andrew Scott</mods:namePart>
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<mods:affiliation id="89C8774CAF454F413C22986D8954A159">School of the Environment, Geography and Geological Sciences, University of Portsmouth, Burnaby Building, Burnaby Road, Portsmouth PO 1 3 QL UK; & Department of Earth Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:title id="0FCB9566F9DBB753CF145B41949960B3">Journal of Systematic Palaeontology</mods:title>
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<mods:part id="41F9863B4FB51FAD90701D91C4628EDC">
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<mods:date id="1E76129A883344FB14125E13F37F1C18">2021</mods:date>
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<mods:number id="3AEABDB3CACE773D25AE5085AD28A0D3">2021-10-01</mods:number>
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<mods:url id="A08B43659110F0608A890A4FBA4984B6">http://dx.doi.org/10.1080/14772019.2021.1960911</mods:url>
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<mods:classification id="6F07DFAE27244C35B58158991BE726DF">journal article</mods:classification>
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<mods:identifier id="02332B1996B6F80CC0B1CEC3061335CB" type="DOI">10.1080/14772019.2021.1960911</mods:identifier>
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<treatment id="B9207C419A69FFE40E20FC79FB45FC34" ID-DOI="http://doi.org/10.5281/zenodo.10955115" ID-Zenodo-Dep="10955115" LSID="urn:lsid:plazi:treatment:B9207C419A69FFE40E20FC79FB45FC34" httpUri="http://treatment.plazi.org/id/B9207C419A69FFE40E20FC79FB45FC34" lastPageNumber="721" pageId="31" pageNumber="721">
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<subSubSection id="79939EDC9A69FFE40E20FC79FDF7FBBC" pageId="31" pageNumber="721" type="nomenclature">
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<paragraph id="3136CD579A69FFE40E20FC79FDEEFB9F" blockId="31.[303,637,1013,1071]" box="[303,637,1013,1036]" pageId="31" pageNumber="721">
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<heading id="6A7E7A3B9A69FFE40E20FC79FDEEFB9F" box="[303,637,1013,1036]" centered="true" fontSize="10" level="2" pageId="31" pageNumber="721" reason="8">
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<taxonomicName id="F689B6D49A69FFE40E20FC79FD8EFB9F" ID-CoL="8RW8L" authority="Gale, 2021" authorityName="Gale" authorityYear="2021" box="[303,541,1013,1036]" class="Asteroidea" family="Sphaerasteridae" genus="Pouzaster" kingdom="Animalia" order="Valvatida" pageId="31" pageNumber="721" phylum="Echinodermata" rank="species" species="pocknotata" status="sp. nov.">
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<emphasis id="03FD11459A69FFE40E20FC79FD8EFB9F" bold="true" box="[303,541,1013,1036]" italics="true" pageId="31" pageNumber="721">Pouzaster pocknotata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="18CEAC3E9A69FFE40D29FC79FDEEFB9F" box="[550,637,1013,1036]" pageId="31" pageNumber="721" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<paragraph id="3136CD579A69FFE40E48FB94FDF7FBBC" blockId="31.[303,637,1013,1071]" box="[327,612,1047,1071]" pageId="31" pageNumber="721">
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(
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<figureCitation id="A9B2D1D29A69FFE40E5FFB94FD99FBBC" box="[336,522,1047,1071]" captionStart="Figure 13" captionStartId="24.[159,226,1587,1608]" captionTargetId="figure-1@23.[159,1467,188,1921]" captionTargetPageId="23" captionText="Figure 13. A–H, P, Sphaeraster tabulatus (Goldfuss, 1833); A, actinal ossicle, to show crater-rimmed spine pits (NHMUK EE 17680); B, G, abactinal ossicles of small individuals which have smooth external faces (NHMUK EE 17682, 17683); C, D, abactinal ossicles of very small individuals (NHMUK EE 17684, 17685); E, lateral view of small abactinal plate (NHMUK EE 17686); F, abactinal ossicle with weak radial sculpture (NHMUK EE 17681); H, enlargement of sculpture of abactinal ossicle A; P, lateral view of abactinal plates of small individual, to show lack of papular pores and articulation structures (compare with Fig. 9M, N; NHMUK EE 17687). I–O, Q–S, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles; I–O, paratype abactinals (I, J, NHMUK EE 17689; K, NHMUK EE 17690; L, M, NHMUK EE 17691; N, NHMUK EE 17692; O, NHMUK EE 17693; R, S, NHMUK EE 17695). Q, holotype, abactinal (NHMUK EE 17688); S, enlargement of surface detail of K. Provenance: A–H, P are from the upper Oxfordian, Hypsilum ammonite Zone, Plettenberg, near Balingen, Germany. I–O, Q–S are from the lower Bathonian, Zigzagiceras zigzag ammonite Zone at La Pouza, near La-Voulte-sur-Rhone, Ard̀eche, France. Scale bars: A, F, O = 1 mm; B–E, G, I–N, P–R = 0.5mm; H, S = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.10955087" httpUri="https://zenodo.org/record/10955087/files/figure.png" pageId="31" pageNumber="721">Figs 13I–O, Q–S</figureCitation>
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,
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<figureCitation id="A9B2D1D29A69FFE40D16FB94FDCEFBBC" box="[537,605,1048,1071]" captionStart="Figure 22" captionStartId="39.[159,226,1644,1665]" captionTargetId="figure-1@38.[159,1466,188,1920]" captionTargetPageId="38" captionText="Figure 22. A–D, G, J, Valettaster digitatus (Quenstedt, 1858), abactinal ossicles, in external view (A, C, D, G, J) and lateral view (B) (GPIT/A5/50). E, Valettaster ocellatus (Forbes, 1848), abactinal ossicle, in external view (NHMUK EE 17696). F, I, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles in external view (MHNML 2020.1.2, 1.3). H, K, Valettaster granulatus (Brunnich Nielsen, 1943), abactinal ossicles, external view (SNSB-BSGP 2020 XLV1, 2). L, Valettaster argus (Spencer, 1907), articulated group of abactinal ossicles (NHMUK EE 17637f). M, N, Valettaster thuyi sp. nov., holotype, abactinal ossicle in external and lateral views, respectively (MHNML 2020.1.1). Provenance: A–D, G, J are from the Oxfordian of Oerlingen, south Germany. B is from the Santonian of East Kent, UK. F, I are from the serpentinus ammonite Zone, Toarcian at Fougerolles, France. H, K are from the lower Maastrichtian of R¨ugen, Germany. M, N are from the bifrons ammonite Zone, Toarcian, Fougerolles, France. L is from the coranguinum Zone, Santonian, Broadstairs, Kent, UK. Scale bars: A–E, G, H, J, K = 5 mm; F, I, L = 0.5mm; M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10955101" httpUri="https://zenodo.org/record/10955101/files/figure.png" pageId="31" pageNumber="721">22F, I</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="79939EDC9A69FFE40F90FBE4FE72FAB3" pageId="31" pageNumber="721" type="description">
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<paragraph id="3136CD579A69FFE40F90FBE4FE72FAB3" blockId="31.[159,781,1128,1312]" pageId="31" pageNumber="721">
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<emphasis id="03FD11459A69FFE40F90FBE4FF79FBEC" bold="true" box="[159,234,1128,1151]" pageId="31" pageNumber="721">Types.</emphasis>
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<materialsCitation id="81E1C70A9A69FFE40FF5FBE4FDCEFB4C" collectionCode="NHMUK" collectorName="La Voulte-sur-Rhone" country="France" location="lower Bathonian" municipality="Zone of La Pouza" pageId="31" pageNumber="721" specimenCode="EE 17688, EE 17689-17695" specimenCount="2" typeStatus="holotype">
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The
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<typeStatus id="EE3273F59A69FFE40E3BFBE4FE02FBEC" box="[308,401,1128,1151]" pageId="31" pageNumber="721" type="holotype">holotype</typeStatus>
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(
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<figureCitation id="A9B2D1D29A69FFE40EA4FBE4FD87FBEC" box="[427,532,1128,1151]" captionStart="Figure 13" captionStartId="24.[159,226,1587,1608]" captionTargetId="figure-1@23.[159,1467,188,1921]" captionTargetPageId="23" captionText="Figure 13. A–H, P, Sphaeraster tabulatus (Goldfuss, 1833); A, actinal ossicle, to show crater-rimmed spine pits (NHMUK EE 17680); B, G, abactinal ossicles of small individuals which have smooth external faces (NHMUK EE 17682, 17683); C, D, abactinal ossicles of very small individuals (NHMUK EE 17684, 17685); E, lateral view of small abactinal plate (NHMUK EE 17686); F, abactinal ossicle with weak radial sculpture (NHMUK EE 17681); H, enlargement of sculpture of abactinal ossicle A; P, lateral view of abactinal plates of small individual, to show lack of papular pores and articulation structures (compare with Fig. 9M, N; NHMUK EE 17687). I–O, Q–S, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles; I–O, paratype abactinals (I, J, NHMUK EE 17689; K, NHMUK EE 17690; L, M, NHMUK EE 17691; N, NHMUK EE 17692; O, NHMUK EE 17693; R, S, NHMUK EE 17695). Q, holotype, abactinal (NHMUK EE 17688); S, enlargement of surface detail of K. Provenance: A–H, P are from the upper Oxfordian, Hypsilum ammonite Zone, Plettenberg, near Balingen, Germany. I–O, Q–S are from the lower Bathonian, Zigzagiceras zigzag ammonite Zone at La Pouza, near La-Voulte-sur-Rhone, Ard̀eche, France. Scale bars: A, F, O = 1 mm; B–E, G, I–N, P–R = 0.5mm; H, S = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.10955087" httpUri="https://zenodo.org/record/10955087/files/figure.png" pageId="31" pageNumber="721">Fig. 13Q</figureCitation>
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) is a primary radial ossicle (
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<collectionCode id="579855929A69FFE40FF7FB04FECCFB0C" box="[248,351,1160,1183]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="31" pageNumber="721" type="Museum">NHMUK</collectionCode>
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<specimenCode id="612F652C9A69FFE40E66FB04FE4BFB0C" box="[361,472,1160,1183]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="31" pageNumber="721" type="Museum">EE 17688</specimenCode>
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) from the
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<location id="34569B8C9A69FFE40D5DFB04FC9BFB0C" LSID="urn:lsid:plazi:treatment:B9207C419A69FFE40E20FC79FB45FC34:34569B8C9A69FFE40D5DFB04FC9BFB0C" box="[594,776,1160,1183]" country="France" municipality="Zone of La Pouza" name="lower Bathonian" pageId="31" pageNumber="721">lower Bathonian</location>
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,
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<taxonomicName id="F689B6D49A69FFE40F90FB24FEEAFB2C" box="[159,377,1192,1215]" class="Cephalopoda" family="Perisphinctidae" genus="Zigzagiceras" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="31" pageNumber="721" phylum="Mollusca" rank="species" species="zigzag">
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<emphasis id="03FD11459A69FFE40F90FB24FEEAFB2C" box="[159,377,1192,1215]" italics="true" pageId="31" pageNumber="721">Zigzagiceras zigzag</emphasis>
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</taxonomicName>
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ammonite
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<collectingMunicipality id="D152572D9A69FFE40EF3FB24FD5CFB2C" box="[508,719,1192,1215]" pageId="31" pageNumber="721">Zone of La Pouza</collectingMunicipality>
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, near
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<collectorName id="9C7CA8819A69FFE40F90FB44FE15FB4C" box="[159,390,1224,1247]" pageId="31" pageNumber="721">La Voulte-sur-Rhone</collectorName>
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(Ard̀eche,
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<collectingCountry id="499E8DC79A69FFE40D0BFB44FDC1FB4C" box="[516,594,1224,1247]" name="France" pageId="31" pageNumber="721">France</collectingCountry>
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).
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</materialsCitation>
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<materialsCitation id="81E1C70A9A69FFE40D68FB44FE4DFAB3" collectionCode="NHMUK" collectorName="La Voulte-sur-Rhone" country="France" location="lower Bathonian" municipality="Zone of La Pouza" pageId="31" pageNumber="721" specimenCode="EE 17688, EE 17689-17695" specimenCount="2" typeStatus="holotype">
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A total of nine similar ossicles from the same locality are
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<typeStatus id="EE3273F59A69FFE40DA9FB64FC9FFB6C" box="[678,780,1256,1279]" pageId="31" pageNumber="721" type="paratype">paratypes</typeStatus>
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(
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<collectionCode id="579855929A69FFE40FA9FA85FE9EFAB3" box="[166,269,1289,1312]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="31" pageNumber="721" type="Museum">NHMUK</collectionCode>
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<specimenCode id="612F652C9A69FFE40E1AFA85FE45FAB3" box="[277,470,1288,1312]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="31" pageNumber="721" type="Museum">EE 17689–17695</specimenCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="79939EDC9A69FFE40F90FAB4FD6AF983" pageId="31" pageNumber="721" type="diagnosis">
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<paragraph id="3136CD579A69FFE40F90FAB4FE49FADC" blockId="31.[159,474,1336,1359]" box="[159,474,1336,1359]" pageId="31" pageNumber="721">
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<emphasis id="03FD11459A69FFE40F90FAB4FE80FADC" bold="true" box="[159,275,1336,1359]" pageId="31" pageNumber="721">Diagnosis.</emphasis>
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As for the genus.
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</paragraph>
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<paragraph id="3136CD579A69FFE40F90FAE5FDCDFA53" blockId="31.[159,781,1385,1472]" pageId="31" pageNumber="721">
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<emphasis id="03FD11459A69FFE40F90FAE5FE1AFA13" bold="true" box="[159,393,1385,1408]" pageId="31" pageNumber="721">Additional material.</emphasis>
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Two ossicles from the
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<emphasis id="03FD11459A69FFE40D9DFAE5FC9EFA13" box="[658,781,1385,1408]" italics="true" pageId="31" pageNumber="721">serpentinus</emphasis>
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ammonite Zone (Toarcian) of Fougerolles, near Caen,
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<collectingCountry id="499E8DC79A69FFE40F90FA25FF7BFA53" box="[159,232,1449,1472]" name="France" pageId="31" pageNumber="721">France</collectingCountry>
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(
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<collectionCode id="579855929A69FFE40FF8FA25FECCFA53" box="[247,351,1449,1472]" pageId="31" pageNumber="721">MHNLM</collectionCode>
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20201.2 and 20201.3).
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</paragraph>
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<paragraph id="3136CD579A69FFE40F90FA55FD6AF983" blockId="31.[159,781,1497,1552]" pageId="31" pageNumber="721">
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<emphasis id="03FD11459A69FFE40F90FA55FE16FA63" bold="true" box="[159,389,1497,1520]" pageId="31" pageNumber="721">Derivation of name.</emphasis>
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Latin for pockmarked (
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<emphasis id="03FD11459A69FFE40D84FA55FD2EFA63" box="[651,701,1497,1520]" italics="true" pageId="31" pageNumber="721">pock</emphasis>
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= scar,
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<emphasis id="03FD11459A69FFE40F90FA75FF70F983" box="[159,227,1529,1552]" italics="true" pageId="31" pageNumber="721">notata</emphasis>
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= marked), in reference to the surface sculpture.
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</paragraph>
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</subSubSection>
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<subSubSection id="79939EDC9A69FFE40F90F9A5FACBFC85" pageId="31" pageNumber="721" type="description">
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<paragraph id="3136CD579A69FFE40F90F9A5FACBFC85" blockId="31.[159,781,1577,1632]" lastBlockId="31.[844,1467,189,790]" pageId="31" pageNumber="721">
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<emphasis id="03FD11459A69FFE40F90F9A5FEB9F9D3" bold="true" box="[159,298,1577,1600]" pageId="31" pageNumber="721">Description.</emphasis>
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The abactinal plates are variable in shape, and hexagonal to oval in outline. All possess a
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<materialsCitation id="81E1C70A9A69FFE40C43FF31FAC6FC85" collectionCode="R" pageId="31" pageNumber="721" specimenCount="1">
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distinctive sculpture of irregular pitting, which has a weak radial alignment in some specimens (e.g.
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<figureCitation id="A9B2D1D29A69FFE40A53FF51FC3AFE87" captionStart="Figure 13" captionStartId="24.[159,226,1587,1608]" captionTargetId="figure-1@23.[159,1467,188,1921]" captionTargetPageId="23" captionText="Figure 13. A–H, P, Sphaeraster tabulatus (Goldfuss, 1833); A, actinal ossicle, to show crater-rimmed spine pits (NHMUK EE 17680); B, G, abactinal ossicles of small individuals which have smooth external faces (NHMUK EE 17682, 17683); C, D, abactinal ossicles of very small individuals (NHMUK EE 17684, 17685); E, lateral view of small abactinal plate (NHMUK EE 17686); F, abactinal ossicle with weak radial sculpture (NHMUK EE 17681); H, enlargement of sculpture of abactinal ossicle A; P, lateral view of abactinal plates of small individual, to show lack of papular pores and articulation structures (compare with Fig. 9M, N; NHMUK EE 17687). I–O, Q–S, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles; I–O, paratype abactinals (I, J, NHMUK EE 17689; K, NHMUK EE 17690; L, M, NHMUK EE 17691; N, NHMUK EE 17692; O, NHMUK EE 17693; R, S, NHMUK EE 17695). Q, holotype, abactinal (NHMUK EE 17688); S, enlargement of surface detail of K. Provenance: A–H, P are from the upper Oxfordian, Hypsilum ammonite Zone, Plettenberg, near Balingen, Germany. I–O, Q–S are from the lower Bathonian, Zigzagiceras zigzag ammonite Zone at La Pouza, near La-Voulte-sur-Rhone, Ard̀eche, France. Scale bars: A, F, O = 1 mm; B–E, G, I–N, P–R = 0.5mm; H, S = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.10955087" httpUri="https://zenodo.org/record/10955087/files/figure.png" pageId="31" pageNumber="721">
|
||
Fig. 13I, N, Q,
|
||
<collectionCode id="579855929A69FFE40C98FF71FC3AFE87" box="[919,937,253,276]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="721">R</collectionCode>
|
||
</figureCitation>
|
||
). The position of possible pr can be identified from comparison with that in other
|
||
<taxonomicName id="F689B6D49A69FFE40BD9FE91FAECFEA7" authorityName="Schondorf" authorityYear="1906" box="[1238,1407,285,308]" class="Asteroidea" family="Sphaerasteridae" kingdom="Animalia" order="Valvatida" pageId="31" pageNumber="721" phylum="Echinodermata" rank="family">Sphaerasteridae</taxonomicName>
|
||
(
|
||
<figureCitation id="A9B2D1D29A69FFE40A9FFE91FC0DFEC7" captionStart="Figure 13" captionStartId="24.[159,226,1587,1608]" captionTargetId="figure-1@23.[159,1467,188,1921]" captionTargetPageId="23" captionText="Figure 13. A–H, P, Sphaeraster tabulatus (Goldfuss, 1833); A, actinal ossicle, to show crater-rimmed spine pits (NHMUK EE 17680); B, G, abactinal ossicles of small individuals which have smooth external faces (NHMUK EE 17682, 17683); C, D, abactinal ossicles of very small individuals (NHMUK EE 17684, 17685); E, lateral view of small abactinal plate (NHMUK EE 17686); F, abactinal ossicle with weak radial sculpture (NHMUK EE 17681); H, enlargement of sculpture of abactinal ossicle A; P, lateral view of abactinal plates of small individual, to show lack of papular pores and articulation structures (compare with Fig. 9M, N; NHMUK EE 17687). I–O, Q–S, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles; I–O, paratype abactinals (I, J, NHMUK EE 17689; K, NHMUK EE 17690; L, M, NHMUK EE 17691; N, NHMUK EE 17692; O, NHMUK EE 17693; R, S, NHMUK EE 17695). Q, holotype, abactinal (NHMUK EE 17688); S, enlargement of surface detail of K. Provenance: A–H, P are from the upper Oxfordian, Hypsilum ammonite Zone, Plettenberg, near Balingen, Germany. I–O, Q–S are from the lower Bathonian, Zigzagiceras zigzag ammonite Zone at La Pouza, near La-Voulte-sur-Rhone, Ard̀eche, France. Scale bars: A, F, O = 1 mm; B–E, G, I–N, P–R = 0.5mm; H, S = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.10955087" httpUri="https://zenodo.org/record/10955087/files/figure.png" pageId="31" pageNumber="721">
|
||
Fig. 13O,
|
||
<collectionCode id="579855929A69FFE40C83FEB1FC0DFEC7" box="[908,926,317,340]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="31" pageNumber="721">R</collectionCode>
|
||
</figureCitation>
|
||
). They are hexagonal and symmetical about the vertical axis, and the articular facet contacting the successive radial is broad. The external face is gently convex, and appears to be smooth in
|
||
<specimenCount id="278F06DE9A69FFE40BEEFE12FAE9FE26" box="[1249,1402,414,437]" count="1" pageId="31" pageNumber="721" type="generic">one specimen</specimenCount>
|
||
(
|
||
<figureCitation id="A9B2D1D29A69FFE40A9FFE12FCE9FE46" captionStart="Figure 13" captionStartId="24.[159,226,1587,1608]" captionTargetId="figure-1@23.[159,1467,188,1921]" captionTargetPageId="23" captionText="Figure 13. A–H, P, Sphaeraster tabulatus (Goldfuss, 1833); A, actinal ossicle, to show crater-rimmed spine pits (NHMUK EE 17680); B, G, abactinal ossicles of small individuals which have smooth external faces (NHMUK EE 17682, 17683); C, D, abactinal ossicles of very small individuals (NHMUK EE 17684, 17685); E, lateral view of small abactinal plate (NHMUK EE 17686); F, abactinal ossicle with weak radial sculpture (NHMUK EE 17681); H, enlargement of sculpture of abactinal ossicle A; P, lateral view of abactinal plates of small individual, to show lack of papular pores and articulation structures (compare with Fig. 9M, N; NHMUK EE 17687). I–O, Q–S, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles; I–O, paratype abactinals (I, J, NHMUK EE 17689; K, NHMUK EE 17690; L, M, NHMUK EE 17691; N, NHMUK EE 17692; O, NHMUK EE 17693; R, S, NHMUK EE 17695). Q, holotype, abactinal (NHMUK EE 17688); S, enlargement of surface detail of K. Provenance: A–H, P are from the upper Oxfordian, Hypsilum ammonite Zone, Plettenberg, near Balingen, Germany. I–O, Q–S are from the lower Bathonian, Zigzagiceras zigzag ammonite Zone at La Pouza, near La-Voulte-sur-Rhone, Ard̀eche, France. Scale bars: A, F, O = 1 mm; B–E, G, I–N, P–R = 0.5mm; H, S = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.10955087" httpUri="https://zenodo.org/record/10955087/files/figure.png" pageId="31" pageNumber="721">Fig. 13O</figureCitation>
|
||
); however, under low-oblique lighting it displays radiating ridges around the margin of the plate. The ossicles are half as thick as broad, and each side has a well-defined rounded rectangular articular surface which bears a scatter of fine pits. A single poorly defined papular notch is present on each corner (
|
||
<figureCitation id="A9B2D1D29A69FFE40A4DFDD3FA38FDE5" box="[1346,1451,607,630]" captionStart="Figure 19" captionStartId="33.[159,226,1341,1362]" captionTargetBox="[190,1447,219,1287]" captionTargetId="figure-177@33.[159,1467,188,1319]" captionTargetPageId="33" captionText="Figure 19. A, C, interpretative drawings of Eosphaeraster amellagensis gen. et sp. nov. A, lateral view of large abactinal plate to show candelabra-like, outward-branching papular pores on rim of plate; C, reconstruction of part of abactinal surface; the large central ossicle is the primary interradial. Note concentration of papular pores towards corners of plates. B, Bolbosphaeraster valettei gen. et sp. nov. Note concentration of few papular pores at corners of plates. D, Pouzaster pocknotata gen. et sp. nov., to show single papular pores at plate junctions." figureDoi="http://doi.org/10.5281/zenodo.10949945" httpUri="https://zenodo.org/record/10949945/files/figure.png" pageId="31" pageNumber="721">Fig. 19D</figureCitation>
|
||
). Other abactinals with a similar sculpture include rather tall, narrow, hexagonal to lozenge-shaped plates (e.g.
|
||
<figureCitation id="A9B2D1D29A69FFE40C43FD33FC5DFD45" box="[844,974,703,727]" captionStart="Figure 13" captionStartId="24.[159,226,1587,1608]" captionTargetId="figure-1@23.[159,1467,188,1921]" captionTargetPageId="23" captionText="Figure 13. A–H, P, Sphaeraster tabulatus (Goldfuss, 1833); A, actinal ossicle, to show crater-rimmed spine pits (NHMUK EE 17680); B, G, abactinal ossicles of small individuals which have smooth external faces (NHMUK EE 17682, 17683); C, D, abactinal ossicles of very small individuals (NHMUK EE 17684, 17685); E, lateral view of small abactinal plate (NHMUK EE 17686); F, abactinal ossicle with weak radial sculpture (NHMUK EE 17681); H, enlargement of sculpture of abactinal ossicle A; P, lateral view of abactinal plates of small individual, to show lack of papular pores and articulation structures (compare with Fig. 9M, N; NHMUK EE 17687). I–O, Q–S, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles; I–O, paratype abactinals (I, J, NHMUK EE 17689; K, NHMUK EE 17690; L, M, NHMUK EE 17691; N, NHMUK EE 17692; O, NHMUK EE 17693; R, S, NHMUK EE 17695). Q, holotype, abactinal (NHMUK EE 17688); S, enlargement of surface detail of K. Provenance: A–H, P are from the upper Oxfordian, Hypsilum ammonite Zone, Plettenberg, near Balingen, Germany. I–O, Q–S are from the lower Bathonian, Zigzagiceras zigzag ammonite Zone at La Pouza, near La-Voulte-sur-Rhone, Ard̀eche, France. Scale bars: A, F, O = 1 mm; B–E, G, I–N, P–R = 0.5mm; H, S = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.10955087" httpUri="https://zenodo.org/record/10955087/files/figure.png" pageId="31" pageNumber="721">Figs 13J–N</figureCitation>
|
||
,
|
||
<figureCitation id="A9B2D1D29A69FFE40CD2FD33FBB0FD45" box="[989,1059,703,726]" captionStart="Figure 22" captionStartId="39.[159,226,1644,1665]" captionTargetId="figure-1@38.[159,1466,188,1920]" captionTargetPageId="38" captionText="Figure 22. A–D, G, J, Valettaster digitatus (Quenstedt, 1858), abactinal ossicles, in external view (A, C, D, G, J) and lateral view (B) (GPIT/A5/50). E, Valettaster ocellatus (Forbes, 1848), abactinal ossicle, in external view (NHMUK EE 17696). F, I, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles in external view (MHNML 2020.1.2, 1.3). H, K, Valettaster granulatus (Brunnich Nielsen, 1943), abactinal ossicles, external view (SNSB-BSGP 2020 XLV1, 2). L, Valettaster argus (Spencer, 1907), articulated group of abactinal ossicles (NHMUK EE 17637f). M, N, Valettaster thuyi sp. nov., holotype, abactinal ossicle in external and lateral views, respectively (MHNML 2020.1.1). Provenance: A–D, G, J are from the Oxfordian of Oerlingen, south Germany. B is from the Santonian of East Kent, UK. F, I are from the serpentinus ammonite Zone, Toarcian at Fougerolles, France. H, K are from the lower Maastrichtian of R¨ugen, Germany. M, N are from the bifrons ammonite Zone, Toarcian, Fougerolles, France. L is from the coranguinum Zone, Santonian, Broadstairs, Kent, UK. Scale bars: A–E, G, H, J, K = 5 mm; F, I, L = 0.5mm; M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10955101" httpUri="https://zenodo.org/record/10955101/files/figure.png" pageId="31" pageNumber="721">22F, I</figureCitation>
|
||
) which were probably unpaired and occupied an interradial position, and broad hexagonal plates (
|
||
<figureCitation id="A9B2D1D29A69FFE40C93FD73FB91FC85" box="[924,1026,767,790]" captionStart="Figure 13" captionStartId="24.[159,226,1587,1608]" captionTargetId="figure-1@23.[159,1467,188,1921]" captionTargetPageId="23" captionText="Figure 13. A–H, P, Sphaeraster tabulatus (Goldfuss, 1833); A, actinal ossicle, to show crater-rimmed spine pits (NHMUK EE 17680); B, G, abactinal ossicles of small individuals which have smooth external faces (NHMUK EE 17682, 17683); C, D, abactinal ossicles of very small individuals (NHMUK EE 17684, 17685); E, lateral view of small abactinal plate (NHMUK EE 17686); F, abactinal ossicle with weak radial sculpture (NHMUK EE 17681); H, enlargement of sculpture of abactinal ossicle A; P, lateral view of abactinal plates of small individual, to show lack of papular pores and articulation structures (compare with Fig. 9M, N; NHMUK EE 17687). I–O, Q–S, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles; I–O, paratype abactinals (I, J, NHMUK EE 17689; K, NHMUK EE 17690; L, M, NHMUK EE 17691; N, NHMUK EE 17692; O, NHMUK EE 17693; R, S, NHMUK EE 17695). Q, holotype, abactinal (NHMUK EE 17688); S, enlargement of surface detail of K. Provenance: A–H, P are from the upper Oxfordian, Hypsilum ammonite Zone, Plettenberg, near Balingen, Germany. I–O, Q–S are from the lower Bathonian, Zigzagiceras zigzag ammonite Zone at La Pouza, near La-Voulte-sur-Rhone, Ard̀eche, France. Scale bars: A, F, O = 1 mm; B–E, G, I–N, P–R = 0.5mm; H, S = 0.2mm." figureDoi="http://doi.org/10.5281/zenodo.10955087" httpUri="https://zenodo.org/record/10955087/files/figure.png" pageId="31" pageNumber="721">Fig. 13M</figureCitation>
|
||
) which might be distal radials
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="79939EDC9A69FFE40C43FCBCFB45FC34" pageId="31" pageNumber="721" type="discussion">
|
||
<paragraph id="3136CD579A69FFE40C43FCBCFB45FC34" blockId="31.[844,1467,816,935]" pageId="31" pageNumber="721">
|
||
<emphasis id="03FD11459A69FFE40C43FCBCFC28FCD4" bold="true" box="[844,955,816,839]" pageId="31" pageNumber="721">Remarks.</emphasis>
|
||
The distinctive sculpture of the plates, irregular pitting with a weak radial element towards the margins, distinguish this species from all others. Reconstruction is not possible without further material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |