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<document id="E261959642E3E9AFBCF460D5776A8900" ID-CLB-Dataset="293303" ID-DOI="10.1080/14772019.2021.1960911" ID-GBIF-Dataset="53749253-12aa-41cc-9eec-e28c0b075738" ID-ISSN="1478-0941" ID-Zenodo-Dep="10949919" ID-ZooBank="F8991F09-B5FB-40EF-B4CC-474D925085B8" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1711484962835" checkinUser="felipe" docAuthor="Gale, Andrew Scott" docDate="2021" docId="B9207C419A5AFFD70CD3FC9EFB56FA8B" docLanguage="en" docName="JSystPalaeontol.19.10.691-741.pdf" docOrigin="Journal of Systematic Palaeontology 19 (10)" docSource="http://dx.doi.org/10.1080/14772019.2021.1960911" docStyle="DocumentStyle:6400638016C1F4AE0D0B6502E032D571.1:JSystPalaeontol.2016-.journal_article.closed" docStyleId="6400638016C1F4AE0D0B6502E032D571" docStyleName="JSystPalaeontol.2016-.journal_article.closed" docStyleVersion="1" docTitle="Valettaster stipes Villier 2010" docType="treatment" docVersion="5" lastPageNumber="734" masterDocId="451904399A76FFFB0F0FFF8CFF93FF93" masterDocTitle="Taxonomy and phylogeny of the football stars (Asteroidea, Sphaerasteridae)" masterLastPageNumber="741" masterPageNumber="691" pageNumber="734" updateTime="1712755325394" updateUser="ExternalLinkService">
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<mods:title id="13C0DEC0C89C3901CBCB1D425DC3956F">Taxonomy and phylogeny of the football stars (Asteroidea, Sphaerasteridae)</mods:title>
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<subSubSection id="79939EDC9A5AFFD70CD3FC9EFAB8FCBA" box="[988,1323,786,809]" pageId="44" pageNumber="734" type="nomenclature">
<paragraph id="3136CD579A5AFFD70CD3FC9EFAB8FCBA" blockId="44.[988,1323,786,809]" box="[988,1323,786,809]" pageId="44" pageNumber="734">
<heading id="6A7E7A3B9A5AFFD70CD3FC9EFAB8FCBA" box="[988,1323,786,809]" centered="true" fontSize="10" level="2" pageId="44" pageNumber="734" reason="8">
<taxonomicName id="F689B6D49A5AFFD70CD3FC9EFAB8FCBA" ID-CoL="84F2R" authority="Villier, 2010" authorityName="Villier" authorityYear="2010" box="[988,1323,786,809]" class="Asteroidea" family="Sphaerasteridae" genus="Valettaster" kingdom="Animalia" order="Valvatida" pageId="44" pageNumber="734" phylum="Echinodermata" rank="species" species="stipes">
<emphasis id="03FD11459A5AFFD70CD3FC9EFB0BFCBA" bold="true" box="[988,1176,786,809]" italics="true" pageId="44" pageNumber="734">Valettaster stipes</emphasis>
<bibRefCitation id="5518B0A69A5AFFD70BAEFC9EFAB8FCBA" author="Villier, L." box="[1185,1323,786,809]" pageId="44" pageNumber="734" pagination="701 - 732" refId="ref29738" refString="Villier, L. 2010. Asteroids from the Barremian calciturbidites of the Serre de Bleyton (Drome, SE France). Annalen des Naturhistorischen Museum in Wein, A 112, 701 - 732." type="journal article" year="2010">Villier, 2010</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="79939EDC9A5AFFD70C45FCDEFA13FCFA" box="[842,1408,849,873]" pageId="44" pageNumber="734" type="reference_group">
<paragraph id="3136CD579A5AFFD70C45FCDEFA13FCFA" blockId="44.[842,1408,849,873]" box="[842,1408,849,873]" pageId="44" pageNumber="734">
<bibRefCitation id="5518B0A69A5AFFD70C45FCDEFCECFCFA" author="Villier, L." box="[842,895,850,873]" pageId="44" pageNumber="734" pagination="701 - 732" refId="ref29738" refString="Villier, L. 2010. Asteroids from the Barremian calciturbidites of the Serre de Bleyton (Drome, SE France). Annalen des Naturhistorischen Museum in Wein, A 112, 701 - 732." type="journal article" year="2010">2010</bibRefCitation>
<taxonomicName id="F689B6D49A5AFFD70C86FCDEFB05FCFA" authority="Villier" authorityName="Villier" authorityYear="2010" box="[905,1174,850,873]" class="Asteroidea" family="Sphaerasteridae" genus="Valettaster" kingdom="Animalia" order="Valvatida" pageId="44" pageNumber="734" phylum="Echinodermata" rank="species" species="stipes">
<emphasis id="03FD11459A5AFFD70C86FCDEFBD0FCFA" box="[905,1091,850,873]" italics="true" pageId="44" pageNumber="734">Valettaster stipes</emphasis>
Villier
</taxonomicName>
: 714, fig. 7, pl. 1pr.
</paragraph>
</subSubSection>
<subSubSection id="79939EDC9A5AFFD70C43FC0FFA3CFC2F" pageId="44" pageNumber="734" type="diagnosis">
<paragraph id="3136CD579A5AFFD70C43FC0FFA3CFC2F" blockId="44.[844,1467,899,956]" pageId="44" pageNumber="734">
<emphasis id="03FD11459A5AFFD70C43FC0FFC52FC09" bold="true" box="[844,961,899,922]" pageId="44" pageNumber="734">Diagnosis.</emphasis>
Primary abactinal ossicles low, with irregular digitate outline. External surface flat, lacking ornament.
</paragraph>
</subSubSection>
<subSubSection id="79939EDC9A5AFFD70C43FC5AFA39FB9D" pageId="44" pageNumber="734" type="materials_examined">
<paragraph id="3136CD579A5AFFD70C43FC5AFA39FB9D" blockId="44.[844,1466,982,1038]" pageId="44" pageNumber="734">
<emphasis id="03FD11459A5AFFD70C43FC5AFC0BFC7E" bold="true" box="[844,920,982,1005]" pageId="44" pageNumber="734">Types.</emphasis>
The
<typeStatus id="EE3273F59A5AFFD70CE6FC5AFBD2FC7E" box="[1001,1089,982,1005]" pageId="44" pageNumber="734" type="holotype">holotype</typeStatus>
is an abactinal ossicle, MPUP VILL026.
<typeStatus id="EE3273F59A5AFFD70CB4FC7BFB8CFB9D" box="[955,1055,1015,1038]" pageId="44" pageNumber="734" type="paratype">Paratypes</typeStatus>
, MPUP VILL027-029, VILL049-051.
</paragraph>
</subSubSection>
<subSubSection id="79939EDC9A5AFFD70C43FBA5FB37FBF2" pageId="44" pageNumber="734" type="distribution">
<paragraph id="3136CD579A5AFFD70C43FBA5FB37FBF2" blockId="44.[844,1466,1065,1121]" pageId="44" pageNumber="734">
<emphasis id="03FD11459A5AFFD70C43FBA5FC4AFBD3" bold="true" box="[844,985,1065,1088]" pageId="44" pageNumber="734">Occurrence.</emphasis>
All are from the Barremian of Serre de Blayton, near Arnayon,
<collectingCountry id="499E8DC79A5AFFD70B5BFBC6FB33FBF2" box="[1108,1184,1098,1121]" name="France" pageId="44" pageNumber="734">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="79939EDC9A5AFFD70C43FBF7FB56FA8B" pageId="44" pageNumber="734" type="discussion">
<paragraph id="3136CD579A5AFFD70C43FBF7FB56FA8B" blockId="44.[844,1467,1147,1304]" pageId="44" pageNumber="734">
<emphasis id="03FD11459A5AFFD70C43FBF7FC28FB01" bold="true" box="[844,955,1147,1170]" pageId="44" pageNumber="734">Remarks.</emphasis>
<taxonomicName id="F689B6D49A5AFFD70CC5FBF7FB18FB01" authorityName="Villier" authorityYear="2010" box="[970,1163,1147,1170]" class="Asteroidea" family="Sphaerasteridae" genus="Valettaster" kingdom="Animalia" order="Valvatida" pageId="44" pageNumber="734" phylum="Echinodermata" rank="species" species="stipes">
<emphasis id="03FD11459A5AFFD70CC5FBF7FB18FB01" box="[970,1163,1147,1170]" italics="true" pageId="44" pageNumber="734">Valettaster stipes</emphasis>
</taxonomicName>
resembles
<taxonomicName id="F689B6D49A5AFFD70A1CFBF7FA05FB01" box="[1299,1430,1147,1170]" class="Asteroidea" family="Sphaerasteridae" genus="Valettaster" kingdom="Animalia" order="Valvatida" pageId="44" pageNumber="734" phylum="Echinodermata" rank="species" species="digitatus">
<emphasis id="03FD11459A5AFFD70A1CFBF7FA05FB01" box="[1299,1430,1147,1170]" italics="true" pageId="44" pageNumber="734">V. digitatus</emphasis>
</taxonomicName>
in the flattened form of the abactinal ossicles and the digitate margin, but differs in the smooth external surface, which in
<taxonomicName id="F689B6D49A5AFFD70CB8FB53FBA6FB65" box="[951,1077,1247,1270]" class="Asteroidea" family="Sphaerasteridae" genus="Valettaster" kingdom="Animalia" order="Valvatida" pageId="44" pageNumber="734" phylum="Echinodermata" rank="species" species="digitatus">
<emphasis id="03FD11459A5AFFD70CB8FB53FBA6FB65" box="[951,1077,1247,1270]" italics="true" pageId="44" pageNumber="734">V. digitatus</emphasis>
</taxonomicName>
carries fine, closely spaced crater-like spine pits (
<figureCitation id="A9B2D1D29A5AFFD70CF8FA8DFB2BFA8B" box="[1015,1208,1280,1304]" captionStart="Figure 22" captionStartId="39.[159,226,1644,1665]" captionTargetId="figure-1@38.[159,1466,188,1920]" captionTargetPageId="38" captionText="Figure 22. AD, G, J, Valettaster digitatus (Quenstedt, 1858), abactinal ossicles, in external view (A, C, D, G, J) and lateral view (B) (GPIT/A5/50). E, Valettaster ocellatus (Forbes, 1848), abactinal ossicle, in external view (NHMUK EE 17696). F, I, Pouzaster pocknotata gen. et sp. nov., abactinal ossicles in external view (MHNML 2020.1.2, 1.3). H, K, Valettaster granulatus (Brunnich Nielsen, 1943), abactinal ossicles, external view (SNSB-BSGP 2020 XLV1, 2). L, Valettaster argus (Spencer, 1907), articulated group of abactinal ossicles (NHMUK EE 17637f). M, N, Valettaster thuyi sp. nov., holotype, abactinal ossicle in external and lateral views, respectively (MHNML 2020.1.1). Provenance: AD, G, J are from the Oxfordian of Oerlingen, south Germany. B is from the Santonian of East Kent, UK. F, I are from the serpentinus ammonite Zone, Toarcian at Fougerolles, France. H, K are from the lower Maastrichtian of R¨ugen, Germany. M, N are from the bifrons ammonite Zone, Toarcian, Fougerolles, France. L is from the coranguinum Zone, Santonian, Broadstairs, Kent, UK. Scale bars: AE, G, H, J, K = 5 mm; F, I, L = 0.5mm; M, N = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.10955101" httpUri="https://zenodo.org/record/10955101/files/figure.png" pageId="44" pageNumber="734">Fig. 22AD, G, J</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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