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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e102683" ID-Pensoft-Pub="1864-8312-81-581" ID-Pensoft-UUID="E6F5D6150A225AFBB372E719EBBEDB68" ID-ZooBank="CA56E9F0B4264014B20BD7C455FD65A7" ModsDocID="1864-8312-81-581" checkinTime="1686864336769" checkinUser="pensoft" docAuthor="Wang, Ya-Lian, Peng, Zi-Qi, Jin, Feng-Liang, Chen, Hong-Wei &amp; Huang, Jia" docDate="2023" docId="EE341BB6C0975F41AE9F60D85CE95DFE" docLanguage="en" docName="ArthropodSystPhyl 81: 581-610" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-06-15" docSource="http://dx.doi.org/10.3897/asp.81.e102683" docTitle="Stegana (Orthostegana) macrostephana Peng &amp; Chen 2023, sp. nov." docType="treatment" docUuid="A88359E6-0151-4DD6-976C-8B008DB7A153" docUuidSource="ZooBank" docVersion="1" id="E6F5D6150A225AFBB372E719EBBEDB68" lastPageNumber="581" masterDocId="E6F5D6150A225AFBB372E719EBBEDB68" masterDocTitle="Integrative species delimitation in the morphologically conservative Stegana subgen. Orthostegana (Diptera, Drosophilidae)" masterLastPageNumber="610" masterPageNumber="581" pageNumber="581" updateTime="1686864336769" updateUser="pensoft">
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<mods:title id="20DA36772357DDAE820A790475BD6E6D">Integrative species delimitation in the morphologically conservative Stegana subgen. Orthostegana (Diptera, Drosophilidae)</mods:title>
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<mods:namePart id="02A11B2F030A6780675AF4B45253205A">Wang, Ya-Lian</mods:namePart>
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<mods:affiliation id="6A0163D8BEA6B8DA05E8F510C83CAA0F">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="1DB386F05A141FD4F3B4F382F4177851">Peng, Zi-Qi</mods:namePart>
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<mods:affiliation id="24B2F9F759E964EDB8F3CF87ED92469B">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="78B867A31988785022D8CAC101EC9702">Jin, Feng-Liang</mods:namePart>
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<mods:affiliation id="55907501715525CC7BA5EC8FD048A642">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="9A3568EC0BA3A72FB11381E4BCF13F06">Chen, Hong-Wei</mods:namePart>
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<mods:affiliation id="8AE1661014C9DC36C8266286286255F2">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:namePart id="E1703EF7E6075F6349C3A31E571D7D86">Huang, Jia</mods:namePart>
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<mods:affiliation id="447DFF609D218404320BF07D877F1573">Department of Entomology, South China Agricultural University, 483 Wushan-lu, Guangzhou 510642, Guangdong, P. R. China</mods:affiliation>
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<mods:date id="4E7FEA8A315503A092A97DA9451B311E">2023</mods:date>
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<subSubSection id="815968860DF243578E724B952570CD36" pageId="0" pageNumber="581" type="nomenclature">
<paragraph id="57E0FB41BCDCE7E6876C6A945589A1BE" pageId="0" pageNumber="581">
3.2.1.6. 
<taxonomicName id="A7D1A3E8F1954A9776000157579CE921" LSID="https://zoobank.org/A88359E6-0151-4DD6-976C-8B008DB7A153" authority="Peng &amp; Chen" authorityName="Peng &amp; Chen" authorityYear="2023" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegana (Orthostegana) macrostephana" order="Diptera" pageId="0" pageNumber="581" phylum="Arthropoda" rank="species" species="macrostephana" status="sp. nov." subGenus="Orthostegana">Stegana (Orthostegana) macrostephana Peng &amp; Chen</taxonomicName>
<taxonomicNameLabel id="1BF89092497DDCA88313C42288804BFE" pageId="0" pageNumber="581">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="D494EAD47C3435E587807C0F304A3468" pageId="0" pageNumber="581" type="description">
<paragraph id="8CB0883C334A092E63E508D659500A87" pageId="0" pageNumber="581">
<figureCitation id="9922ED0D35F5BAD72CD4436D176F60D0" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov. (male); C, D S. (O.) macrostephana Peng &amp; Chen sp. nov. (male); E, F S. (O.) macrostephana Peng &amp; Chen sp. nov. (female); G, H S. (O.) mohnihei Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure7" httpUri="https://binary.pensoft.net/fig/864474" pageId="0" pageNumber="581">Figs 7C-F</figureCitation>
<figureCitation id="D6E8D1BDF4F76BAC1050F4CE34662923" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Mesotarsus of males in ventral view. A Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov.; B S. (O.) macrostephana Peng &amp; Chen sp. nov.; C S. (O.) mohnihei Peng &amp; Chen sp. nov.; D S. (O.) obscurala Peng &amp; Chen sp. nov.; E S. (O.) pinguitia Peng &amp; Chen sp. nov." figureDoi="10.3897/asp.81.e102683.figure10" httpUri="https://binary.pensoft.net/fig/864477" pageId="0" pageNumber="581">, 10B</figureCitation>
<figureCitation id="E47BE5E412CFA50D696D64EE26481A7C" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">, 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9F2A8EEA52B9575397D6C9CFE24B165A" pageId="0" pageNumber="581" type="diagnosis">
<paragraph id="31786101BE7A1F90A51D85A9BC5526BB" pageId="0" pageNumber="581">Diagnosis.</paragraph>
<paragraph id="F219A5997B94D5F0EEC0E3F20EB1FA5C" pageId="0" pageNumber="581">
This species closely resembles 
<taxonomicName id="071258A9F8D13E566D5431CC9EE2699C" lsidName="S. (O.) pinguitia" pageId="0" pageNumber="581" rank="species" species="pinguitia" subGenus="O.">S. (O.) pinguitia</taxonomicName>
<emphasis id="B8DC4EDA6677E70D437D7BFB2FF8BD18" bold="true" pageId="0" pageNumber="581">sp. nov.</emphasis>
in the aedeagus medioventrally and distally with dense long hairs (Figs 
<figureCitation id="EDCB1832BD85924DB8F55BF768B23AB0" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17C</figureCitation>
, 
<figureCitation id="1C6707000AA4D135888CAC08C2610287" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17D</figureCitation>
, 
<figureCitation id="C1BC72A33E4122EBE748FA4A840D1B62" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Stegana (Orthostegana) pinguitia Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure20" httpUri="https://binary.pensoft.net/fig/864487" pageId="0" pageNumber="581">20C</figureCitation>
, 
<figureCitation id="C01D8CAF1A019A4F4C0F40A857EEE0FA" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Stegana (Orthostegana) pinguitia Peng &amp; Chen sp. nov., male terminalia. A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure20" httpUri="https://binary.pensoft.net/fig/864487" pageId="0" pageNumber="581">20D</figureCitation>
), but can be distinguished from the latter by having the mesotarsus ventrally with 2 rows of black, apically blunt, stout spines; metatarsus with 15/3+1 spines; 2nd to 4th segments with 6/1, 4/1, and 3/1 spines, respectively (Fig. 
<figureCitation id="A38E618CC8EF033FE7286443C62B4E30" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Mesotarsus of males in ventral view. A Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov.; B S. (O.) macrostephana Peng &amp; Chen sp. nov.; C S. (O.) mohnihei Peng &amp; Chen sp. nov.; D S. (O.) obscurala Peng &amp; Chen sp. nov.; E S. (O.) pinguitia Peng &amp; Chen sp. nov." figureDoi="10.3897/asp.81.e102683.figure10" httpUri="https://binary.pensoft.net/fig/864477" pageId="0" pageNumber="581">10B</figureCitation>
); pregonites slender, with 
<emphasis id="B0FFE28297DBB762F4C5A92F5DE3ED6C" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
2 or 3 sensilla medioventrally and some setae distally per side (Fig. 
<figureCitation id="12F67FD61299B503B23B3FBB1B505152" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17C, D</figureCitation>
).
</paragraph>
<caption id="AD2675D85E7E552E1067A8AAB61F9AEF" doi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581" start="Figure 17" startId="F17">
<paragraph id="922A60AFE13E83E85DB4CB99D43D4559" pageId="0" pageNumber="581">
<emphasis id="8E167B9F06CD949473B2BC382B7D9526" bold="true" pageId="0" pageNumber="581">Figure 17.</emphasis>
<taxonomicName id="522C7E698DE5D2C2499ED4EFD0807A30" authorityName="Peng &amp; Chen" authorityYear="2023" class="Insecta" family="Drosophilidae" genus="Stegana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stegana (Orthostegana) macrostephana" order="Diptera" pageId="0" pageNumber="581" phylum="Arthropoda" rank="species" species="macrostephana" subGenus="Orthostegana">Stegana (Orthostegana) macrostephana</taxonomicName>
Peng &amp; Chen 
<emphasis id="9595C368EEF0BB2047B8AFC64E82001A" bold="true" pageId="0" pageNumber="581">sp. nov.</emphasis>
, male (
<emphasis id="3204DBFD1865685A20DD874860A52D85" bold="true" pageId="0" pageNumber="581">A</emphasis>
- 
<emphasis id="4AF4BF7146A82B1C6077F2E0C8BFECA7" bold="true" pageId="0" pageNumber="581">D</emphasis>
) and female terminalia (
<emphasis id="992F17D7CABDA8A98FC114684085AEB1" bold="true" pageId="0" pageNumber="581">E</emphasis>
). 
<emphasis id="35FC554E132E4E53976ADA94F2C9AB43" bold="true" pageId="0" pageNumber="581">A</emphasis>
Epandrium, surstylus, and cercus in lateral view; 
<emphasis id="B15F962B0E0BE92221EA267CF6A48548" bold="true" pageId="0" pageNumber="581">B</emphasis>
Surstylus in ventral view; 
<emphasis id="562D8BEA8AB1173EBCC2F82FF45B5EA2" bold="true" pageId="0" pageNumber="581">C</emphasis>
, 
<emphasis id="1E23B840F5DBE87EEBEA284534B26BA1" bold="true" pageId="0" pageNumber="581">D</emphasis>
Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; 
<emphasis id="2C74BCD3D8C694AABB0647880D7A7C6B" bold="true" pageId="0" pageNumber="581">E</emphasis>
Eighth sternite in ventral view. Scale bars = 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="41827F5A1A5FA752783E7936588FBFE7" pageId="0" pageNumber="581" type="description">
<paragraph id="7EA9933D1FBE62562CB27F6F649738D5" pageId="0" pageNumber="581">Description.</paragraph>
<paragraph id="FD5EB0F28F79617116F7BCC59937FB17" pageId="0" pageNumber="581">
<emphasis id="310902AEA98EBA65E420A118DA8E4C58" bold="true" pageId="0" pageNumber="581">
Male and female. 
<emphasis id="C751B521DBD8EEA9A423D324921E309D" italics="true" pageId="0" pageNumber="581">Head</emphasis>
</emphasis>
: Ocellar triangle dark brown, with 2 small setae posterior to ocellar setae. Postocellar seta absent. Frons brown, with numerous minute interfrontal setulae. Pedicel yellow and first flagellomere brownish yellow to brown, with black pubescence. Face and gena brown. Clypeus dark brown medially and yellow laterally. Palpus brownish basally and dark brown distally. 
<emphasis id="BEC28A62796D04A17EB86B6F2590C069" bold="true" italics="true" pageId="0" pageNumber="581">Thorax</emphasis>
: Mesonotum yellow to brownish yellow (Fig. 
<figureCitation id="1A715A65AF0B00945C5D976BB3EEB055" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov. (male); C, D S. (O.) macrostephana Peng &amp; Chen sp. nov. (male); E, F S. (O.) macrostephana Peng &amp; Chen sp. nov. (female); G, H S. (O.) mohnihei Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure7" httpUri="https://binary.pensoft.net/fig/864474" pageId="0" pageNumber="581">7C, E</figureCitation>
). Pleura with a distinct brownish to brown longitudinal stripe above (Fig. 
<figureCitation id="7F6E389EE70A2D2FA3D20788316DBE0E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov. (male); C, D S. (O.) macrostephana Peng &amp; Chen sp. nov. (male); E, F S. (O.) macrostephana Peng &amp; Chen sp. nov. (female); G, H S. (O.) mohnihei Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure7" httpUri="https://binary.pensoft.net/fig/864474" pageId="0" pageNumber="581">7D, F</figureCitation>
). Scutellum mostly yellow to brownish yellow (Fig. 
<figureCitation id="FFB72F606C70E7749423024817C92B27" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov. (male); C, D S. (O.) macrostephana Peng &amp; Chen sp. nov. (male); E, F S. (O.) macrostephana Peng &amp; Chen sp. nov. (female); G, H S. (O.) mohnihei Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure7" httpUri="https://binary.pensoft.net/fig/864474" pageId="0" pageNumber="581">7C, E</figureCitation>
). Halter mostly brownish, with a yellow patch at knob (Fig. 
<figureCitation id="659599CAB7872B2936D93CC26875F95B" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Head, thorax, and abdomen in dorsal and lateral views. A, B Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov. (male); C, D S. (O.) macrostephana Peng &amp; Chen sp. nov. (male); E, F S. (O.) macrostephana Peng &amp; Chen sp. nov. (female); G, H S. (O.) mohnihei Peng &amp; Chen sp. nov. (male)." figureDoi="10.3897/asp.81.e102683.figure7" httpUri="https://binary.pensoft.net/fig/864474" pageId="0" pageNumber="581">7C-F</figureCitation>
). Legs yellowish (Fig. 
<figureCitation id="FEBFFA172F7C22E9BF482AC3FC9FA3A7" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Mesotarsus of males in ventral view. A Stegana (Orthostegana) latipalpula Peng &amp; Chen sp. nov.; B S. (O.) macrostephana Peng &amp; Chen sp. nov.; C S. (O.) mohnihei Peng &amp; Chen sp. nov.; D S. (O.) obscurala Peng &amp; Chen sp. nov.; E S. (O.) pinguitia Peng &amp; Chen sp. nov." figureDoi="10.3897/asp.81.e102683.figure10" httpUri="https://binary.pensoft.net/fig/864477" pageId="0" pageNumber="581">10B</figureCitation>
). 
<emphasis id="C32D358401E37A53D587D6E1B46080FE" bold="true" italics="true" pageId="0" pageNumber="581">Abdominal tergites</emphasis>
: brownish yellow to brownish. 
<emphasis id="98B51C5430E6F7D18AE781E83E8D20A2" bold="true" italics="true" pageId="0" pageNumber="581">Male terminalia</emphasis>
: Epandrium mostly pubescent except for anterior margin, with 
<emphasis id="2E85962027F2B3034A9079954F044810" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
28 setae on dorsal to posterior portions per side (Fig. 
<figureCitation id="7BCBCB2808A860A86B974BB77ECFA133" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17A</figureCitation>
). Surstylus separated from epandrium, with 
<emphasis id="E649E4061728CD903272B055088132D6" italics="true" pageId="0" pageNumber="581">ca.</emphasis>
22 long and 15 short peg-shaped prensisetae near anterior margin on inner surface (Fig. 
<figureCitation id="5853409596BAE6385EB7638B273A7F25" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17B</figureCitation>
). Pregonites fused basally (Fig. 
<figureCitation id="A7046464D7A8AAC30B60C79C8AF0807B" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17C, D</figureCitation>
). Aedeagus pubescent distally (Fig. 
<figureCitation id="739CDCFAFB321B5E886B2C126C50BFF7" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17C, D</figureCitation>
). Aedeagal sheath expanded and lobe-shaped distally (Fig. 
<figureCitation id="8C930E1920F8EB34415A1EBBDD418A4F" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17D</figureCitation>
). 
<emphasis id="EAD26CAB1C965E85E22C0DBD5ECC1712" bold="true" italics="true" pageId="0" pageNumber="581">Female terminalia</emphasis>
: Eighth sternite with 3 rows of orderly and nearly symmetrical peg-shaped prensisetae subposteromedially and posterolaterally, and with 3 sensilla distally per side (Fig. 
<figureCitation id="CD0F3049E9C288FC97DB35A2C8BF79C0" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Stegana (Orthostegana) macrostephana Peng &amp; Chen sp. nov., male (A - D) and female terminalia (E). A Epandrium, surstylus, and cercus in lateral view; B Surstylus in ventral view; C, D Hypandrium, phallapodeme, pregonite, aedeagus, and aedeagal sheath in ventral and lateral views; E Eighth sternite in ventral view. Scale bars = 0.1 mm." figureDoi="10.3897/asp.81.e102683.figure17" httpUri="https://binary.pensoft.net/fig/864484" pageId="0" pageNumber="581">17E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BBB5E15E7C9FEF27C49A1D37C73FB275" pageId="0" pageNumber="581" type="description">
<paragraph id="3B09D5E5348B605508D5DE4F964453C7" pageId="0" pageNumber="581">Measurements and indices.</paragraph>
<paragraph id="163BF12E2AB1B02A954E1C5064044A89" pageId="0" pageNumber="581">
<emphasis id="11C43AD2B907619D8051B89211175F18" bold="true" pageId="0" pageNumber="581">BL</emphasis>
= 3.50 mm in holotype (range in 5♂ and 5♀ paratypes: 3.30-3.50 in ♂, 2.67-3.33 in ♀), 
<emphasis id="FF231FC083D2AD1210E49586B24BBCF5" bold="true" pageId="0" pageNumber="581">ThL</emphasis>
= 1.50 mm (1.40-1.50 in ♂, 1.00-1.53 in ♀), 
<emphasis id="E94B8B2DC2770363FE03593C70120B9B" bold="true" pageId="0" pageNumber="581">WL</emphasis>
= 3.00 mm (2.77-2.95 in ♂, 2.27-3.23 in ♀), 
<emphasis id="D47901310E9A15C4D5B2EBBC38878416" bold="true" pageId="0" pageNumber="581">WW</emphasis>
= 1.25 mm (1.20-1.30 in ♂, 1.16-1.50 in ♀), 
<emphasis id="706A941D1ED2CB94A7FCD0CD7B88A402" bold="true" pageId="0" pageNumber="581">arb</emphasis>
= 8/4 (5-9/4-5), 
<emphasis id="4817E4D8786BD870C22C0825F278BBFD" bold="true" pageId="0" pageNumber="581">avd</emphasis>
= 0.63 (0.64-0.93), 
<emphasis id="6E705DC051FBD62DBED8D21A478C9CBC" bold="true" pageId="0" pageNumber="581">adf</emphasis>
= 1.60 (1.20-2.50), 
<emphasis id="63CB7EED96D8DF8C6F8C924C4AC81343" bold="true" pageId="0" pageNumber="581">flw</emphasis>
= 1.70 (1.63-2.00), 
<emphasis id="F3F161CE3995A9C02D270CE94514B9F5" bold="true" pageId="0" pageNumber="581">FW/HW</emphasis>
= 0.41 (0.35-0.41), 
<emphasis id="18FE916ECA2D0428EC3F6680E3FCE178" bold="true" pageId="0" pageNumber="581">ch/o</emphasis>
= 0.12 (0.08-0.14), 
<emphasis id="838DF3F7927B76CD0B1CA17D2B0AF9AA" bold="true" pageId="0" pageNumber="581">prorb</emphasis>
= 1.08 (0.90-1.39), 
<emphasis id="44D7C92A521B718B2F1ACFC3D1D7F01F" bold="true" pageId="0" pageNumber="581">rcorb</emphasis>
= 0.80 (0.53-0.83), 
<emphasis id="3BBDB0E810E31EEDAF87D21478346ED8" bold="true" pageId="0" pageNumber="581">vb</emphasis>
= 0.50 (0.24-0.56), 
<emphasis id="51F0440A6588772CFC504A1D2415C345" bold="true" pageId="0" pageNumber="581">dcl</emphasis>
= 0.50 (0.33-0.67), 
<emphasis id="4AA30913FC1DA0D4270957638F0D7AB2" bold="true" pageId="0" pageNumber="581">presctl</emphasis>
= 0.40 (0.33-0.57), 
<emphasis id="E082758F1E8305FF352BD272E3976EA8" bold="true" pageId="0" pageNumber="581">sctl</emphasis>
= 1.12 (1.06-1.29), 
<emphasis id="34D93C1D7EF8C6282A91FCFAAC7710FB" bold="true" pageId="0" pageNumber="581">sterno</emphasis>
= 0.77 (0.68-0.97), 
<emphasis id="1E1E5E798B5B560FCB8D08C2E2E74473" bold="true" pageId="0" pageNumber="581">orbito</emphasis>
= 1.67 (1.60-2.20), 
<emphasis id="B8515A081DCF3025EE206AA23A5CF465" bold="true" pageId="0" pageNumber="581">dcp</emphasis>
= 0.28 (0.23-0.41), 
<emphasis id="B5E64CB175DD7176D231A9E10326EA8D" bold="true" pageId="0" pageNumber="581">sctlp</emphasis>
= 1.00 (0.91-1.33), 
<emphasis id="54281F130335201E60C93FB13F271810" bold="true" pageId="0" pageNumber="581">C</emphasis>
= 2.33 (2.00-2.67), 
<emphasis id="8247985AB1B3A1A062B225A3CE61A37B" bold="true" pageId="0" pageNumber="581">4c</emphasis>
= 0.91 (0.85-1.00), 
<emphasis id="BE89BF45144898CBB6FDE54CEB2C9314" bold="true" pageId="0" pageNumber="581">4v</emphasis>
= 1.70 (1.56-0.95), 
<emphasis id="6941131D02E8FBE852555679883FD7DE" bold="true" pageId="0" pageNumber="581">5x</emphasis>
= 1.15 (1.20-2.11), 
<emphasis id="D193F6B3FDA6B5705816BCA384E9D326" bold="true" pageId="0" pageNumber="581">ac</emphasis>
= 10.00 (9.00-13.33), 
<emphasis id="0C261EA6ED908A85EE989F88161F2D78" bold="true" pageId="0" pageNumber="581">M</emphasis>
= 0.45 (0.38-0.48), 
<emphasis id="B83EF37CAA0753D688268E1C353AFB4C" bold="true" pageId="0" pageNumber="581">C3F</emphasis>
= 0.74 (0.65-0.81).
</paragraph>
</subSubSection>
<subSubSection id="33E99F5BFF95CECE3A85F62E5F96B8D0" pageId="0" pageNumber="581" type="distribution">
<paragraph id="6444EE6FD37F6E1567115E7874609BA4" pageId="0" pageNumber="581">Distribution.</paragraph>
<paragraph id="22E8403C65A699D265C99BFC084B1663" pageId="0" pageNumber="581">China (Yunnan).</paragraph>
</subSubSection>
<subSubSection id="96178771A05AC1381DDE0EE118F72F4A" pageId="0" pageNumber="581" type="type material">
<paragraph id="353EBD01DC4A7840768C8E29A441786F" pageId="0" pageNumber="581">Type material examined.</paragraph>
<paragraph id="FFCF3A8E7CA79DAD535B6CFE49CD0D41" pageId="0" pageNumber="581">
<emphasis id="789B27694E1D2EA75F12EF72C77DF64C" bold="true" pageId="0" pageNumber="581">Holotype</emphasis>
: ♂ (SCAU, No. 112090), China: Mengdong, Cangyun, Yunnan, 
<geoCoordinate id="A58287BF0DE083967D86C4FDD99E7E8C" degrees="23" direction="north" minutes="10" orientation="latitude" precision="15" seconds="08" value="23.168888">23°10′08″N</geoCoordinate>
, 99°13′52″ E, altitude 1323 m, 6.v.2016, swept from tussock, Y.Q. Liu. 
<emphasis id="249A5CB9B025030BF81DF1739216AA5E" bold="true" pageId="0" pageNumber="581">Paratypes</emphasis>
: 10♂, 5♀ (SCAU, Nos. 112091-98, 112110-16), altitudes 1300-1320 m, J. Huang, Y.Q. Liu, Y.L. Wang, L. Zhu, other data same as the holotype; 1♂, 9♀ (SCAU, Nos. 112099-108), Muyiji Park, Ximeng, Yunnan, 
<geoCoordinate id="ACB3B5705E5DEDD2A08FE3CACA593AAF" degrees="22" direction="north" minutes="37" orientation="latitude" precision="15" seconds="15" value="22.620834">22°37′15″N</geoCoordinate>
, 
<geoCoordinate id="32AA1ED88E802BB5D548B2B55BA26D7F" degrees="99" direction="east" minutes="35" orientation="longitude" precision="15" seconds="42" value="99.595">99°35′42″E</geoCoordinate>
, altitude 1203 m, 29.iv.2016, swept from tussocks, J. Huang, Y.Q. Liu, Y.L. Wang, L. Zhu; 1♂ (SCAU, No. 112109), Botanic Garden, Ruili, Yunnan, 
<geoCoordinate id="7E83615D039C3114E3ABDD56DAACC34B" degrees="24" direction="north" minutes="01" orientation="latitude" precision="15" seconds="12" value="24.02">24°01′12″N</geoCoordinate>
, 
<geoCoordinate id="B7481AF227BF44F4B12D4FFA1248D4D4" degrees="97" direction="east" minutes="51" orientation="longitude" precision="15" seconds="33" value="97.85916">97°51′33″E</geoCoordinate>
, altitude 1174 m, 22.v.2016, swept from tussock, J. Huang.
</paragraph>
</subSubSection>
<subSubSection id="008D31AD5AF9332CEEA3805B68679BFA" pageId="0" pageNumber="581" type="etymology">
<paragraph id="E4449147F23C23CF46EB449E510F2F8B" pageId="0" pageNumber="581">Etymology.</paragraph>
<paragraph id="C41AA5F135DB07AC3266E9A098811922" pageId="0" pageNumber="581">
A combination of the Greek words &quot; 
<emphasis id="AF25ED2529B2D836B2D693C34FDA622A" italics="true" pageId="0" pageNumber="581">macr</emphasis>
-&quot; (= large) and &quot; 
<emphasis id="7B439FE142EED0A1D14963C4629384F7" italics="true" pageId="0" pageNumber="581">stephan</emphasis>
-&quot; (= crown), referring to the apically floriform aedeagus.
</paragraph>
</subSubSection>
</treatment>
</document>