227 lines
28 KiB
XML
227 lines
28 KiB
XML
<document id="B08147761DC8C76008A3F1BCC467E68C" ID-CLB-Dataset="301504" ID-DOI="10.1093/zooldnnean/zlad146" ID-GBIF-Dataset="5e66aeaf-44c5-4be9-924d-cc13680ace6a" ID-ISSN="0024-4082" ID-Zenodo-Dep="13315269" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="valdenar" IM.metadata_approvedBy="valdenar" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1722845279467" checkinUser="plazi" docAuthor="Lumen, Ryan & Kamdńskd, Marcdn J." docDate="2024" docId="C27387A7FFE0FFCF9AA905ADFE537012" docLanguage="en" docName="zlad146.pdf" docOrigin="Zoological Journal of the Linnean Society 201 (201)" docStyle="DocumentStyle:36B3BD6A90C22AB4F7F465C853188CC8.7:ZoolJLinnSoc.2017-2023.journal_article" docStyleId="36B3BD6A90C22AB4F7F465C853188CC8" docStyleName="ZoolJLinnSoc.2017-2023.journal_article" docStyleVersion="7" docTitle="Biolus analisetosus, comb. nov." docType="treatment" docVersion="5" lastPageNumber="376" masterDocId="3E4AFFDFFFF2FFDD9A72053DFF83770F" masterDocTitle="evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae" masterLastPageNumber="386" masterPageNumber="358" pageNumber="376" updateTime="1723663925820" updateUser="ExternalLinkService">
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<mods:title id="715A77B368C83F9E38D9AE99F113185B">evdsdon of the darkldng beetle genus Eurynotus (Blaptdnae: Platynotdnd % and new records of ovovdvdpary dn ºenebrdonddae</mods:title>
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<mods:namePart id="7F4444C9A975401101E4F845E2F9000A">Lumen, Ryan</mods:namePart>
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<mods:namePart id="2A78450170C89098BF956C1CB889D467">Kamdńskd, Marcdn J.</mods:namePart>
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<mods:title id="A495E0F2C655AA8521A92D5AD66CB2EB">Zoological Journal of the Linnean Society</mods:title>
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<mods:date id="61AB303109983A9E95BF70627EE20DE7">2024</mods:date>
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<treatment id="C27387A7FFE0FFCF9AA905ADFE537012" ID-DOI="http://doi.org/10.5281/zenodo.13315091" ID-Zenodo-Dep="13315091" LSID="urn:lsid:plazi:treatment:C27387A7FFE0FFCF9AA905ADFE537012" httpUri="http://treatment.plazi.org/id/C27387A7FFE0FFCF9AA905ADFE537012" lastPageNumber="376" pageId="18" pageNumber="376">
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<subSubSection id="02C0653AFFE0FFCF9AA905ADFD1377A5" box="[219,656,143,170]" pageId="18" pageNumber="376" type="nomenclature">
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<paragraph id="4A6536B1FFE0FFCF9AA905ADFD1377A5" blockId="18.[219,656,143,170]" box="[219,656,143,170]" pageId="18" pageNumber="376">
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<heading id="112D81DDFFE0FFCF9AA905ADFD1377A5" box="[219,656,143,170]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="4">
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<emphasis id="78AEEAA3FFE0FFCF9AA905ADFD1377A5" bold="true" box="[219,656,143,170]" pageId="18" pageNumber="376">
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1.
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<taxonomicName id="8DDA4D32FFE0FFCF9A8105B2FD9977A6" authority="(Koch)" baseAuthorityName="Koch" box="[243,538,143,170]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus" status="comb. nov.">
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<emphasis id="78AEEAA3FFE0FFCF9A8105B2FE4B77A5" bold="true" box="[243,456,143,170]" italics="true" pageId="18" pageNumber="376">Biolus analisetosus</emphasis>
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(Koch)
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</taxonomicName>
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<taxonomicNameLabel id="639D57D8FFE0FFCF986D05ADFD1377A5" box="[543,656,144,170]" pageId="18" pageNumber="376" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="02C0653AFFE0FFCF9B350582FE537012" pageId="18" pageNumber="376" type="description">
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<paragraph id="4A6536B1FFE0FFCF9B350582FDA677D6" blockId="18.[113,762,191,285]" box="[327,549,191,217]" pageId="18" pageNumber="376">
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<heading id="112D81DDFFE0FFCF9B350582FDA677D6" box="[327,549,191,217]" centered="true" fontSize="9" level="2" pageId="18" pageNumber="376" reason="3">
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<emphasis id="78AEEAA3FFE0FFCF9B350582FDA677D6" bold="true" box="[327,549,191,217]" pageId="18" pageNumber="376">
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(
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<figureCitation id="D2E12A34FFE0FFCF9B230582FE1C77D6" box="[337,415,191,217]" captionStart="Figure 2" captionStartId="3.[129,194,1510,1534]" captionTargetBox="[137,1469,147,1474]" captionTargetId="figure-202@3.[129,1473,144,1483]" captionTargetPageId="3" captionText="Figure 2. Biolus stat.nov.specdes.A, Biolus analisetosus (Koch% comb. nov. Holotype; B, B.asperipennis (Mulsant and Rey% stat.res., comb.nov. Lectotype; C, B. brunoi sp. nov. Holotype; D, B. denticosta (Mulsant and Rey% comb. nov. Lectotype; E, B. graafi (Koch% comb. nov. Holotype; F,B. granulatus (Fabrdcdus% comb. nov. Lectotype; G, B. omeri (Koch% comb.nov.Holotype; H, B. sauroides (Koch% comb. nov. Holotype; I, Eurynotus norrisi Mulsant and Rey syn. nov. of B. granulatus, Lectotype." figureDoi="http://doi.org/10.5281/zenodo.13315277" httpUri="https://zenodo.org/record/13315277/files/figure.png" pageId="18" pageNumber="376">Figs 2A</figureCitation>
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,
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<figureCitation id="D2E12A34FFE0FFCF9BD80582FE4877D6" box="[426,459,191,217]" captionStart="Figure 6" captionStartId="7.[129,194,1700,1724]" captionTargetBox="[131,1472,145,1670]" captionTargetId="figure-63@7.[129,1473,144,1673]" captionTargetPageId="7" captionText="Figure 6. Characters used dn the claddstdc analysds.Arrows dnddcate partdcular states of gdven characters from the morphologdcal matrdx [e.g." figureDoi="http://doi.org/10.5281/zenodo.13315289" httpUri="https://zenodo.org/record/13315289/files/figure.png" pageId="18" pageNumber="376">6C</figureCitation>
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,
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<figureCitation id="D2E12A34FFE0FFCF9BA70582FE7177D6" box="[469,498,191,217]" captionStart="Figure 7" captionStartId="12.[113,178,1803,1827]" captionTargetBox="[118,1458,150,1772]" captionTargetId="figure-7@12.[114,1458,144,1775]" captionTargetPageId="12" captionText="Figure 7. Phylogeny of morphology, ovovdvdpary, and female reproductdve structures of Eurynotdna.A, Eurynotus barbosai, 1st dnstar larva ddssected from bursa copulatrdx. B, Biolus omeri, gendtal tubes. C, B. brunoi, spermatheca and ovdposdtor.D, E. capensis, gendtal tubes of two dnddvdduals.E, B.analisetosus, sclerdte and egg ddssected from bursa copulatrdx. F, B. granulatus, sclerdte from bursa copulatrdx.G, E. asperatus, padred sclerdtes on ddstal part of bursa copulatrdx. H, Oncotus cedrimontis, bursa copulatrdx. Abbrevdatdons: c1–c4, lobes of coxdtdes. Numbers dn parentheses represent characters and states used dn analysds." figureDoi="http://doi.org/10.5281/zenodo.13323862" httpUri="https://zenodo.org/record/13323862/files/figure.png" pageId="18" pageNumber="376">7E</figureCitation>
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,
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<figureCitation id="D2E12A34FFE0FFCF9B8E0582FD9977D6" box="[508,538,191,217]" captionStart="Figure 8" captionStartId="13.[129,194,1898,1922]" captionTargetBox="[134,1473,837,1871]" captionTargetId="figure-393@13.[129,1473,834,1871]" captionTargetPageId="13" captionText="Figure 8. Aedeagal morphology of Eurynotdna specdes.A, Biolus analisetosus; B, B.asperipennis; C, B. brunoi sp. nov.; D, B. denticosta; E, B. graafi; F, B. granulatus; G, B. omeri; H, B. sauroides; I, Eurynotus asperatus; J, E.barbosai; K, E. capensis; L, E. graniderma; M, Capidium tuberculatum; N, Oncotus cedrimontis; O, Heteropsectropus amaroides; P, Schyzoschelus africanus. Parameres hdghldghted red. Scale = 1mm." figureDoi="http://doi.org/10.5281/zenodo.13315291" httpUri="https://zenodo.org/record/13315291/files/figure.png" pageId="18" pageNumber="376">8A</figureCitation>
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)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="4A6536B1FFE0FFCF9A0305DBFF5C7612" blockId="18.[113,762,191,285]" pageId="18" pageNumber="376">
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<taxonomicName id="8DDA4D32FFE0FFCF9A0305DBFF5677F2" box="[113,213,230,253]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">
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<emphasis id="78AEEAA3FFE0FFCF9A0305DBFF5677F2" box="[113,213,230,253]" italics="true" pageId="18" pageNumber="376">Eurynotus</emphasis>
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</taxonomicName>
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(
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<taxonomicName id="8DDA4D32FFE0FFCF9A9E05DBFDF877F1" authority="Koch 1954 a: 278" authorityName="Koch" authorityPageNumber="278" authorityYear="1954" box="[236,635,229,254]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
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<emphasis id="78AEEAA3FFE0FFCF9A9E05DBFEAB77F1" box="[236,296,230,254]" italics="true" pageId="18" pageNumber="376">Biolus</emphasis>
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%
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<emphasis id="78AEEAA3FFE0FFCF9B4D05DBFE3177F1" box="[319,434,230,254]" italics="true" pageId="18" pageNumber="376">analisetosus</emphasis>
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Koch 1954a: 278
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</taxonomicName>
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.– Kamdńskd 2016: 235.
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</paragraph>
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<paragraph id="4A6536B1FFE0FFCF9A030402FEC47485" blockId="18.[113,764,318,906]" pageId="18" pageNumber="376">
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<emphasis id="78AEEAA3FFE0FFCF9A030402FE0C7659" box="[113,399,318,342]" italics="true" pageId="18" pageNumber="376">
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Diagnosis:
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<taxonomicName id="8DDA4D32FFE0FFCF9AAE0403FE0C7659" authorityName="Koch" authorityYear="1954" box="[220,399,318,342]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">Biolus analisetosus</taxonomicName>
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</emphasis>
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ds most sdmdlar to other specdes wdth tubercles on the elytral ddsc (e.g.
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<taxonomicName id="8DDA4D32FFE0FFCF9BCF0463FDBE7679" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[445,573,350,374]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
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<emphasis id="78AEEAA3FFE0FFCF9BCF0463FDBE7679" box="[445,573,350,374]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8DDA4D32FFE0FFCF983A0463FD1A7679" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[584,665,350,374]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
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<emphasis id="78AEEAA3FFE0FFCF983A0463FD1A7679" box="[584,665,350,374]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8DDA4D32FFE0FFCF98D60463FD77767A" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[676,756,350,373]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
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<emphasis id="78AEEAA3FFE0FFCF98D60463FD77767A" box="[676,756,350,373]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8DDA4D32FFE0FFCF9A030443FF64769A" box="[113,231,381,405]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
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<emphasis id="78AEEAA3FFE0FFCF9A030443FF64769A" box="[113,231,381,405]" italics="true" pageId="18" pageNumber="376">B denticosta</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="8DDA4D32FFE0FFCF9B6E0443FE0D769A" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[284,398,381,405]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
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<emphasis id="78AEEAA3FFE0FFCF9B6E0443FE0D769A" box="[284,398,381,405]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
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</taxonomicName>
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%. External features to separate them dnclude the colour of the tarsal setae (
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<taxonomicName id="8DDA4D32FFE0FFCF987004A0FD1276BB" authorityName="Koch" authorityYear="1954" box="[514,657,412,436]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
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<emphasis id="78AEEAA3FFE0FFCF987004A0FD1276BB" box="[514,657,412,436]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
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</taxonomicName>
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= golden;
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<taxonomicName id="8DDA4D32FFE0FFCF9A030481FF4076DC" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[113,195,444,467]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
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<emphasis id="78AEEAA3FFE0FFCF9A030481FF4076DC" box="[113,195,444,467]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="8DDA4D32FFE0FFCF9AA20481FEC876DB" box="[208,331,444,468]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
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<emphasis id="78AEEAA3FFE0FFCF9AA20481FEC876DB" box="[208,331,444,468]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="8DDA4D32FFE0FFCF9BF60481FE7B76DB" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[388,504,444,468]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
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<emphasis id="78AEEAA3FFE0FFCF9BF60481FE7B76DB" box="[388,504,444,468]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
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</taxonomicName>
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= black or brown/dark%, sculpturdng of abdomdnal ventrdte V (wdth two puncture
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<typeStatus id="95618813FFE0FFCF98B104E6FD7976FC" box="[707,762,475,499]" pageId="18" pageNumber="376">types</typeStatus>
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and apdcal meddan notch dn females; sdmple and wdthout female notch dn
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<taxonomicName id="8DDA4D32FFE0FFCF9ABC0727FED3753D" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[206,336,538,562]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
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<emphasis id="78AEEAA3FFE0FFCF9ABC0727FED3753D" box="[206,336,538,562]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="8DDA4D32FFE0FFCF9BF60727FE55753E" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[388,470,538,561]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
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<emphasis id="78AEEAA3FFE0FFCF9BF60727FE55753E" box="[388,470,538,561]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
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</taxonomicName>
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%, mdcrotuberculatdon (more dntense dn
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<taxonomicName id="8DDA4D32FFE0FFCF9AAB0707FEA9755E" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[217,298,569,593]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
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<emphasis id="78AEEAA3FFE0FFCF9AAB0707FEA9755E" box="[217,298,569,593]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
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</taxonomicName>
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%, elytral costae (present only dn
|
||
<taxonomicName id="8DDA4D32FFE0FFCF98020707FD69755E" box="[624,746,569,593]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
|
||
<emphasis id="78AEEAA3FFE0FFCF98020707FD69755E" box="[624,746,569,593]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
|
||
</taxonomicName>
|
||
%. Internally, the female bursa copulatrdx of
|
||
<taxonomicName id="8DDA4D32FFE0FFCF984B0764FD4E757F" authorityName="Koch" authorityYear="1954" box="[569,717,600,624]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
|
||
<emphasis id="78AEEAA3FFE0FFCF984B0764FD4E757F" box="[569,717,600,624]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
|
||
</taxonomicName>
|
||
has a sclerdte sdtuated near to the apex (all specdes except for
|
||
<taxonomicName id="8DDA4D32FFE0FFCF98940745FF5B75A0" baseAuthorityName="Fabricius" baseAuthorityYear="1787" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE0FFCF98940745FF5B75A0" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
|
||
</taxonomicName>
|
||
lack sclerdtes% and an ‘accessory’ pouch (present dn
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9A03078AFE8275C1" authorityName="Koch" authorityYear="1954" box="[113,257,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
|
||
<emphasis id="78AEEAA3FFE0FFCF9A03078AFE8275C1" box="[113,257,694,718]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9B7D078AFEE075C1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[271,355,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
|
||
<emphasis id="78AEEAA3FFE0FFCF9B7D078AFEE075C1" box="[271,355,694,718]" italics="true" pageId="18" pageNumber="376">B. graafi</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9BD2078AFDA075C1" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[416,547,694,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE0FFCF9BD2078AFDA075C1" box="[416,547,694,718]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
|
||
</taxonomicName>
|
||
; absent dn,
|
||
<taxonomicName id="8DDA4D32FFE0FFCF98D2078AFD7775C1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[672,756,695,718]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="omeri">
|
||
<emphasis id="78AEEAA3FFE0FFCF98D2078AFD7775C1" box="[672,756,695,718]" italics="true" pageId="18" pageNumber="376">B. omeri</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9A0307EBFF6B75E2" baseAuthorityName="Koch" baseAuthorityYear="1955" box="[113,232,725,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="sauroides">
|
||
<emphasis id="78AEEAA3FFE0FFCF9A0307EBFF6B75E2" box="[113,232,725,749]" italics="true" pageId="18" pageNumber="376">B. sauroides</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9A8407EBFEF775E2" box="[246,372,725,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="denticosta">
|
||
<emphasis id="78AEEAA3FFE0FFCF9A8407EBFEF775E2" box="[246,372,725,749]" italics="true" pageId="18" pageNumber="376">B. denticosta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9BF007EBFD9575E2" box="[386,534,726,749]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="asperipennis">
|
||
<emphasis id="78AEEAA3FFE0FFCF9BF007EBFD9575E2" box="[386,534,726,749]" italics="true" pageId="18" pageNumber="376">B. asperipennis</emphasis>
|
||
</taxonomicName>
|
||
%.
|
||
<taxonomicName id="8DDA4D32FFE0FFCF985D07E8FD2F75E1" baseAuthorityName="Koch" baseAuthorityYear="1954" box="[559,684,725,750]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="graafi">
|
||
<emphasis id="78AEEAA3FFE0FFCF985D07E8FD2F75E1" box="[559,684,725,750]" italics="true" pageId="18" pageNumber="376">Biolus graafi</emphasis>
|
||
</taxonomicName>
|
||
can be ddfferentdated by the surface sculpturdng (much more dntense mdcrotuberculatdon than dn
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9BE20628FD9D7423" authorityName="Koch" authorityYear="1954" box="[400,542,788,812]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="analisetosus">
|
||
<emphasis id="78AEEAA3FFE0FFCF9BE20628FD9D7423" box="[400,542,788,812]" italics="true" pageId="18" pageNumber="376">B. analisetosus</emphasis>
|
||
</taxonomicName>
|
||
%. Other specdes wdth bursa copulatrdx sclerdtes (
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9BF2060EFDAB7444" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[384,552,819,843]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE0FFCF9BF2060EFDAB7444" box="[384,552,819,843]" italics="true" pageId="18" pageNumber="376">Biolus granulatus</emphasis>
|
||
</taxonomicName>
|
||
% can be further sep* arated by the shape of the sclerdte (chevron*shaped, versus axe* ldke dn
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9AC7064EFEB67485" baseAuthorityName="Fabricius" baseAuthorityYear="1787" box="[181,309,882,906]" class="Insecta" family="Tenebrionidae" genus="Biolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="376" phylum="Arthropoda" rank="species" species="granulatus">
|
||
<emphasis id="78AEEAA3FFE0FFCF9AC7064EFEB67485" box="[181,309,882,906]" italics="true" pageId="18" pageNumber="376">B. granulatus</emphasis>
|
||
</taxonomicName>
|
||
%.
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE0FFCF9A03069AFF4F711D" blockId="18.[113,763,935,1554]" pageId="18" pageNumber="376">
|
||
<emphasis id="78AEEAA3FFE0FFCF9A03069AFF7F74B0" box="[113,252,935,959]" italics="true" pageId="18" pageNumber="376">Redescription:</emphasis>
|
||
Length
|
||
<quantity id="8D229B54FFE0FFCF9B06069AFE6274B0" box="[372,481,935,959]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="18" pageNumber="376" unit="mm" value="16.5" valueMax="17.0" valueMin="16.0">16–17mm</quantity>
|
||
, wddth
|
||
<quantity id="8D229B54FFE0FFCF983C069AFD2274B0" box="[590,673,935,959]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="18" pageNumber="376" unit="mm" value="7.5" valueMax="8.0" valueMin="7.0">7–8mm</quantity>
|
||
. Head: Coarsely punctate, punctures closely spaced, separated by ≤1 puncture ddameter to confluent. Mentum wdth apdcal notch. Eye wdth dmpressed sulcus sdtuated around perdmeter of dorsal lobe. Prothorax: Pronotum coarsely punctate, punctures closely spaced, separated by ≤1 puncture ddameter to con* fluent. Hypomeron rugose, wdth large and shallow punctures. Pterothorax: Elytron not costate, wdth ddstdnct tubercles or radsed/shdndng callosdtdes on ddsc. Epdpleuron wdth few to no mdcrotubercles. Abdomen: Punctate, punctures separated by ≥1 puncture ddameter. Abdomdnal ventrdte V punctures separ* ated by ≤1 puncture ddameter wdth larger, setdgerous punctures dntermdxed. Females wdth meddan apdcal notch. Legs: Male and female legs unmoddfied. ºarsd wdth golden setae. Males: pro* and mesotarsd robust wdth large, ventral, tomentose pads. Male termdnalda: Parameres strongly tapered and ‘vase*shaped’ (taperdng apdcally wdth sharp subapdcal constrdctdon, expanddng sldghtly toward apex%. Female termdnalda: Bursa copulatrdx wdth accessory pouch and chevron*shaped sclerdte near spermathecal opendng.
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE0FFCF9A030312FE3571AA" blockId="18.[113,762,1583,1763]" pageId="18" pageNumber="376">
|
||
<emphasis id="78AEEAA3FFE0FFCF9A030312FEEF7148" box="[113,364,1583,1607]" italics="true" pageId="18" pageNumber="376">Type material examined:</emphasis>
|
||
<typeStatus id="95618813FFE0FFCF9B040312FE5A7148" box="[374,473,1583,1607]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
|
||
(ºMNH%: ‘7 weeks Poort C.P, 17*XI*1940 G van Son;
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9BE70373FD827169" box="[405,513,1614,1638]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
|
||
(Bdolus% analdsetosus C. Koch det., 195 Koch; analdsetosus Koch;
|
||
<typeStatus id="95618813FFE0FFCF98580350FD0E718A" box="[554,653,1645,1669]" pageId="18" pageNumber="376" type="holotype">Holotype</typeStatus>
|
||
No: 1843
|
||
<taxonomicName id="8DDA4D32FFE0FFCF9A0303B0FF5E71AA" box="[113,221,1677,1701]" class="Copepoda" family="Eunicicolidae" genus="Eurynotus" kingdom="Animalia" order="Cyclopoida" pageId="18" pageNumber="376" phylum="Arthropoda" rank="genus">Eurynotus</taxonomicName>
|
||
analdsetosus KOCH’.
|
||
</paragraph>
|
||
<paragraph id="4A6536B1FFE0FFCF9AFF0391FE9C71EC" blockId="18.[113,762,1583,1763]" pageId="18" pageNumber="376">(See Supportdng Informatdon, Appenddx S1 for adddtdonal ma* terdal examdned.%</paragraph>
|
||
<paragraph id="4A6536B1FFE0FFCF9A030238FE537012" blockId="18.[113,464,1797,1821]" box="[113,464,1797,1821]" pageId="18" pageNumber="376">
|
||
<emphasis id="78AEEAA3FFE0FFCF9A030238FF6C7012" box="[113,239,1797,1821]" italics="true" pageId="18" pageNumber="376">Distribution:</emphasis>
|
||
South Afrdca (Fdg. 9%.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |