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<mods:title id="D148457BCCA535C9DEF9C4F6EF254595">A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California-Oregon</mods:title>
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<mods:namePart id="87425FB40F0627D8FB07BCE51B2D71A1">Liu, Hsiu-Ping</mods:namePart>
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<treatment id="ACD2E4CBD6C514A0E13E04C72040EF0F" ID-GBIF-Taxon="154125939" LSID="urn:lsid:plazi:treatment:ACD2E4CBD6C514A0E13E04C72040EF0F" httpUri="http://treatment.plazi.org/id/ACD2E4CBD6C514A0E13E04C72040EF0F" lastPageId="9" lastPageNumber="56" pageId="8" pageNumber="55">
<subSubSection id="540661E85D599091B234B99FDC057BA8" pageId="8" pageNumber="55" type="nomenclature">
<paragraph id="17F2BE4D33CD067685F176D3959A7E7A" pageId="8" pageNumber="55">
<taxonomicName id="1B269F131F57B0BFE02C83B9D92E0427" ID-CoL="6JB2L" authority="Hershler, Liu, Frest &amp; Hubbart, 2017" authorityName="Hershler, Liu, Frest &amp; Hubbart" authorityYear="2017" class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Fluminicola fresti" order="Littorinimorpha" pageId="8" pageNumber="55" phylum="Mollusca" rank="species" species="fresti">Fluminicola fresti Hershler, Liu, Frest &amp; Hubbart, 2017</taxonomicName>
Figure 4H
</paragraph>
</subSubSection>
<subSubSection id="9F29F8565E5315B625CE314CA0A98C98" pageId="9" pageNumber="56" type="reference_group">
<paragraph id="C52406D4E19A2EAB7F2799D2B3FA4E1C" pageId="9" pageNumber="56">
<taxonomicName id="60AEC64D9FE54C326330C9D77734E167" class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Fluminicola fresti" order="Littorinimorpha" pageId="9" pageNumber="56" phylum="Mollusca" rank="species" species="fresti">
<pageBreakToken id="129DA989671C7A90420AC6A8B491CE71" pageId="9" pageNumber="56" start="start">Fluminicola</pageBreakToken>
fresti
</taxonomicName>
-
<bibRefCitation id="016522F8590F3EAA38C93DD327A78E7A" author="Hershler, R" journalOrPublisher="ZooKeys" pageId="15" pageNumber="62" pagination="1 - 20" title="Two new species of Fluminicola (Caenogastropoda, Lithoglyphidae) from southwest Oregon, USA, and a range extension for F. multifarius." url="https://doi.org/10.3897/zookeys.679.13472" volume="679" year="2017">Hershler et al. 2017</bibRefCitation>
: 10-14, figs 4
<normalizedToken id="DED06EB822215FFA7B4E81467C1A8AB0" originalValue="EG">E-G</normalizedToken>
, 5 (Diversion from Big Butte Springs through Butte Falls Hatchery, just south of Butte
<normalizedToken id="98FF5159D08880BD49D9200B635EF231" originalValue="FallsFish">Falls-Fish</normalizedToken>
Lake Road (Jackson County 321) and 0.16 km west of Butte
<normalizedToken id="A0A5D9E361EB897FFE83E1380FC1B0B1" originalValue="FallsProspect">Falls-Prospect</normalizedToken>
Road (Jackson County 922), Jackson County, Oregon,
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,
<geoCoordinate id="C816784057BC931475F5486D16247A47" direction="west" orientation="longitude" precision="5" value="-122.5551">122.5551W</geoCoordinate>
).
</paragraph>
</subSubSection>
<subSubSection id="1FB4D839AB9EF4412F92938F6AA61DE2" pageId="9" pageNumber="56" type="distribution">
<paragraph id="13DE3D8276F35BA93FC54277DCE5B5D1" pageId="9" pageNumber="56">Distribution.</paragraph>
<paragraph id="23B827B094D4383043C6BE821CA83219" pageId="9" pageNumber="56">
North Fork Umpqua River drainage and Rogue River basin north of Little Butte Creek, Oregon (
<bibRefCitation id="11FD82E5D08A98ED7B80A706DB8655F6" author="Hershler, R" journalOrPublisher="ZooKeys" pageId="15" pageNumber="62" pagination="1 - 20" title="Two new species of Fluminicola (Caenogastropoda, Lithoglyphidae) from southwest Oregon, USA, and a range extension for F. multifarius." url="https://doi.org/10.3897/zookeys.679.13472" volume="679" year="2017">Hershler et al. 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6D632D9C134AEC9D747F1D189457D20F" pageId="9" pageNumber="56" type="materials_examined">
<paragraph id="01D046067E5703D4CB124F26C266E70B" pageId="9" pageNumber="56">Referred material.</paragraph>
<paragraph id="35ABBA218F873E2DB8145BD38B8B0FA4" pageId="9" pageNumber="56">
OREGON. Klamath County. USNM 1144900, USNM 1469091, Harriman Spring, outflow of main spring (
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,
<geoCoordinate id="6D49DA0713AEEEEFB8A0DDA8BECFE537" direction="west" orientation="longitude" precision="5" value="-122.1009">122.1009W</geoCoordinate>
),
</paragraph>
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<subSubSection id="86CEB862D6632A35FA9EEC9B43EE621F" pageId="9" pageNumber="56" type="remarks">
<paragraph id="BB5D192F04C0CB509179D27BEAD9FBD6" pageId="9" pageNumber="56">Remarks.</paragraph>
<paragraph id="7D91868DEC6A05E985748F15A49D219C" pageId="9" pageNumber="56">
The Harriman Springs pebblesnails have small (shell height, about 4.0 mm), narrowly conical shells with convex whorls that well conform to
<taxonomicName id="1F3525DAEF8D825C4D20814ADE5024E2" class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" kingdom="Animalia" lsidName="F. fresti" order="Littorinimorpha" pageId="9" pageNumber="56" phylum="Mollusca" rank="species" species="fresti">F. fresti</taxonomicName>
; they also closely resemble this species in details of radula morphology. Radula: central teeth with 3-4 lateral cusps, 1 basal cusp; lateral teeth with 3-4 cusps on outer side, 2 cusps on inner side; inner marginal teeth with 23-28 cusps; outer marginal teeth with 28-36 cusps(USNM 1144900).
</paragraph>
<paragraph id="B24BF21191F4CAC5B7BC3537B76977E8" pageId="9" pageNumber="56">
We sequenced only a single specimen of
<taxonomicName id="E0E55258F8DABB534B4F88183C912E48" class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" kingdom="Animalia" lsidName="F. fresti" order="Littorinimorpha" pageId="9" pageNumber="56" phylum="Mollusca" rank="species" species="fresti">F. fresti</taxonomicName>
from Harriman Springs, which was collected during our first visit to this locality. Our subsequent collections from this site that were preserved in 90% ethanol for mtDNA analysis did not contain this species although a few specimens were found in one of the subsamples that had been prepared for anatomical study.
</paragraph>
<paragraph id="D3884CE0C60E6481171AC90A96F1D613" pageId="9" pageNumber="56">
This new record extends the range of
<taxonomicName id="609313665AC2603BD2ACDC61AFF3D825" class="Gastropoda" family="Lithoglyphidae" genus="Fluminicola" kingdom="Animalia" lsidName="F. fresti" order="Littorinimorpha" pageId="9" pageNumber="56" phylum="Mollusca" rank="species" species="fresti">F. fresti</taxonomicName>
about 26 km eastward from the Rogue River headwaters.
</paragraph>
</subSubSection>
</treatment>
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