161 lines
23 KiB
XML
161 lines
23 KiB
XML
<document id="F467C33B6D3933A12A29F35A055F9580" ENCODING="UTF-8" ID-GBIF-Dataset="2515ee23-bd5e-4eb7-b560-e395a6353603" ModsDocID="z01410p001" checkinTime="1247236958323" checkinUser="thomas" docAuthor="Wilson J. E. M. Costa" docDate="2007" docId="AC7345DA5C3F8AD235E66175F43E0CEF" docLanguage="en" docName="2007_Costa_gg1.xml" docOrigin="Zootaxa 1410" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:A1E8EDF5-B267-4CB6-9206-9F014134DFF2" docTitle="Plesiolebiasini Costa" docType="treatment" docVersion="12" lastPageNumber="3" masterDocId="7C34FA08AA4B6CA35B8B5C94E53E7F90" masterDocTitle="Taxonomy of the plesiolebiasine killifish genera Pituna, Plesiolebias and Maratecoara (Teleostei: Cyprinodontiformes: Rivulidae), with descriptions of nine new species." masterLastPageNumber="41" masterPageNumber="1" pageNumber="2" updateTime="1701314555653" updateUser="plazi">
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<mods:title id="CA74425A066E9683CA3192DF15371DA3">Taxonomy of the plesiolebiasine killifish genera Pituna, Plesiolebias and Maratecoara (Teleostei: Cyprinodontiformes: Rivulidae), with descriptions of nine new species.</mods:title>
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<mods:namePart id="91EE1EB2FF3DF9FA63FFD83266E5EECB">Wilson J. E. M. Costa</mods:namePart>
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<mods:date id="C41372F8CFCD7596FC0D4D9CA105446A">2007</mods:date>
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<treatment id="AC7345DA5C3F8AD235E66175F43E0CEF" ID-DOI="http://doi.org/10.5281/zenodo.6237649" ID-GBIF-Taxon="100121467" ID-Zenodo-Dep="6237649" LSID="urn:lsid:plazi:treatment:AC7345DA5C3F8AD235E66175F43E0CEF" httpUri="http://treatment.plazi.org/id/AC7345DA5C3F8AD235E66175F43E0CEF" lastPageNumber="3" pageNumber="2">
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<subSubSection id="E488FDAB265A59C3108ABEBA2A3DF2E3" type="nomenclature">
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<paragraph id="6BE4C4909F95D68235E7A202ABBE2877" pageNumber="2">
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[[ Tribe
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<taxonomicName id="516F1D987B8749ACAC8A5DED5CB1E591" pageNumber="2" rank="tribe" tribe="Plesiolebiasini">Plesiolebiasini Costa</taxonomicName>
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]]
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</paragraph>
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</subSubSection>
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<subSubSection id="9BCFA8C5C9D6166626C8C97292DE427D" type="discussion">
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<paragraph id="C61B882CC335BDCDBF545962065E8C67" pageNumber="2">
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The
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<taxonomicName id="1D8B9131129BDBB84141F48E10C8E9A1" pageNumber="2" rank="tribe" tribe="Plesiolebiasini">Plesiolebiasini</taxonomicName>
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(sens. Costa, 2004) is a well corroborated clade of Neotropical seasonal killifishes, whose recognition is supported both by morphological (Costa, 1998a) and genetic data (Murphy et al, 1999; Hrbek & Larson, 1999). It includes the following genera (number of valid species according to Costa, 2003, appear in parentheses):
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<taxonomicName id="251D76CA0FA219CDC0844F77500FEACE" LSID-ZBK="urn:lsid:zoobank.org:act:A083017D-552D-4D60-AAC9-76E17E602498" class="Actinopterygii" family="Rivulidae" genus="Maratecoara" kingdom="Animalia" lsidName-ZBK="Maratecoara Costa 1995:68" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Maratecoara Costa</taxonomicName>
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(2),
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<taxonomicName id="642259E19F2855F087DDDC98021AE5CD" LSID-ZBK="urn:lsid:zoobank.org:act:45418B12-BD99-4AA8-932B-AA7D4E5E4E82" class="Actinopterygii" family="Rivulidae" genus="Papiliolebias" kingdom="Animalia" lsidName-ZBK="Papiliolebias Plesiolebias Costa 1998:319" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Papiliolebias Costa</taxonomicName>
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(1),
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<taxonomicName id="44D06CC4AF1A78FDA1419E41AE083788" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Pituna Costa</taxonomicName>
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(2),
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<taxonomicName id="E7BA96AE8276BADF7DAA6707F031C98F" LSID-ZBK="urn:lsid:zoobank.org:act:81D9E5F6-D246-4FCD-BC27-6C3DE9A12A63" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias Costa 1989:193" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Plesiolebias Costa</taxonomicName>
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(4), and
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<taxonomicName id="1141445C428350F2816755C77DD07888" LSID-ZBK="urn:lsid:zoobank.org:act:C4B36824-2379-45D5-B023-5869C055E393" class="Actinopterygii" family="Rivulidae" genus="Stenolebias" kingdom="Animalia" lsidName-ZBK="Stenolebias Costa 1995:70" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Stenolebias Costa</taxonomicName>
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(2). Species of
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<taxonomicName id="95FED2BC8A3AC3C479DB9A40B5713876" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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,
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<taxonomicName id="EB1740B3E0E6691E41AFCF3B50FD13CF" LSID-ZBK="urn:lsid:zoobank.org:act:81D9E5F6-D246-4FCD-BC27-6C3DE9A12A63" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias Costa 1989:193" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Plesiolebias</taxonomicName>
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and
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<taxonomicName id="BF8E14EE58F3BF6B187A77500E91626A" LSID-ZBK="urn:lsid:zoobank.org:act:A083017D-552D-4D60-AAC9-76E17E602498" class="Actinopterygii" family="Rivulidae" genus="Maratecoara" kingdom="Animalia" lsidName-ZBK="Maratecoara Costa 1995:68" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Maratecoara</taxonomicName>
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are often found in the same biotopes in the Araguaia and Tocantins river basins, but
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<taxonomicName id="C381A4CEC7F7A76C72DB693564FAD543" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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also occurs in the rio
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<normalizedToken id="026129644CEFB945480B38A898E73409" originalValue="Paranaíba">Paranaiba</normalizedToken>
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basin and
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<taxonomicName id="751A4F8719A531DA21DEE8ECC7D5E98C" LSID-ZBK="urn:lsid:zoobank.org:act:81D9E5F6-D246-4FCD-BC27-6C3DE9A12A63" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias Costa 1989:193" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Plesiolebias</taxonomicName>
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is in the rio Paraguay basin.
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<taxonomicName id="B1332FF4B45A3581743B75D4BDABBBFA" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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and
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<taxonomicName id="D9DCDC8D269B650F0108006FAA084DEC" LSID-ZBK="urn:lsid:zoobank.org:act:81D9E5F6-D246-4FCD-BC27-6C3DE9A12A63" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias Costa 1989:193" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Plesiolebias</taxonomicName>
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are also present in the rio Xingu basin (e. g., Costa, 1998a, 1998b, 2003; present paper). The remaining two plesiolebiasine genera,
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<taxonomicName id="37F2EFA3CE4702294D89AC360A494550" LSID-ZBK="urn:lsid:zoobank.org:act:45418B12-BD99-4AA8-932B-AA7D4E5E4E82" class="Actinopterygii" family="Rivulidae" genus="Papiliolebias" kingdom="Animalia" lsidName-ZBK="Papiliolebias Plesiolebias Costa 1998:319" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Papiliolebias</taxonomicName>
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and
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<taxonomicName id="1DFFCA5BC5D21FFB6E3179E25D41351B" LSID-ZBK="urn:lsid:zoobank.org:act:C4B36824-2379-45D5-B023-5869C055E393" class="Actinopterygii" family="Rivulidae" genus="Stenolebias" kingdom="Animalia" lsidName-ZBK="Stenolebias Costa 1995:70" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Stenolebias</taxonomicName>
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, are endemic to the rio Paraguay basin, in the Chaco (Paraguay, Bolivia and Argentina) and the Brazilian Pantanal, respectively (e. g., Costa, 2003).
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</paragraph>
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<paragraph id="FD216F2118B69D38ADEDACC175A6A6DC" pageNumber="2">
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Plesiolebiasines are seasonal fishes, inhabiting temporary freshwater biotopes formed during the rainy season (October-April) and which are completely dry during part of the year (June-September). In southern Amazonian tributaries, plesiolebiasines exhibit a noteworthy evolutionary radiation, involving both morphological and ecological features. Species of
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<taxonomicName id="CFED25333918E46ED5DD87B5802FE11C" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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are slender, usually found only in the peripheral shallowest parts of pools, about 100 mm deep or less.
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<taxonomicName id="404652406BA4AFB61A856EB3126B478E" LSID-ZBK="urn:lsid:zoobank.org:act:81D9E5F6-D246-4FCD-BC27-6C3DE9A12A63" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias Costa 1989:193" order="Cyprinodontiformes" pageNumber="2" phylum="Chordata" rank="genus">Plesiolebias</taxonomicName>
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comprises small species, in which the maximum adult size often does not surpass 20 mm SL, and typically lives in mid-water, usually close to aquatic vegetation.
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</paragraph>
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<paragraph id="2D2537171E9A4CB0F36EB9E1D748721E" pageNumber="3">
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<taxonomicName id="75577C7161101DD56240C74BAE5057C9" LSID-ZBK="urn:lsid:zoobank.org:act:A083017D-552D-4D60-AAC9-76E17E602498" class="Actinopterygii" family="Rivulidae" genus="Maratecoara" kingdom="Animalia" lsidName-ZBK="Maratecoara Costa 1995:68" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">
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<pageStartToken id="2C2BBCC6EF6C561F6E49182052992A51" pageNumber="3">Maratecoara</pageStartToken>
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</taxonomicName>
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includes deep-bodied species, in which males have long unpaired fins and anterodorsal trunk profile strongly convex. It is collected near the bottom of pools (Costa, 1998a).
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</paragraph>
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<paragraph id="9D1C783E8A082C05D983868CDE67BFC0" pageNumber="3">
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<taxonomicName id="C36A1FB28B1600C75401E125C089A6C3" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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was first erected to include a single species,
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<taxonomicName id="1AA323C1F0B03912B2D978ECF86242D1" LSID-ZBK="urn:lsid:zoobank.org:act:D03646D6-94E9-4B78-9DB1-193B232214C4" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna poranga Costa 1989:226" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="poranga">P. poranga Costa</taxonomicName>
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, from the rio Araguaia basin, central Brazil (Costa, 1989a). Subsequently, Costa (1991a) examined the type specimens of
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<taxonomicName id="ED2734FC501EC3FAEF806EDBADA98D36" LSID-ZBK="urn:lsid:zoobank.org:act:E26AA8C0-D309-45EB-BE05-FB8D9F3CFC85" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" lsidName-ZBK="Rivulus compactus Myers 1927:120" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="compactus">Rivulus compactus Myers</taxonomicName>
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, from the rio Tocantins basin (also in central Brazil), and reported that they were similar to
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<taxonomicName id="859F222167508D3EF499CE1678E057DC" LSID-ZBK="urn:lsid:zoobank.org:act:D03646D6-94E9-4B78-9DB1-193B232214C4" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna poranga Costa 1989:226" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="poranga">P. poranga</taxonomicName>
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. He consequently transferred
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<taxonomicName id="E864016E419057EE7D35BF9CF17A2EFF" LSID-ZBK="urn:lsid:zoobank.org:act:E26AA8C0-D309-45EB-BE05-FB8D9F3CFC85" class="Actinopterygii" family="Rivulidae" genus="Rivulus" kingdom="Animalia" lsidName-ZBK="Rivulus compactus Myers 1927:120" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="compactus">R. compactus</taxonomicName>
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to
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<taxonomicName id="F9DC9209245DBE598B9064A156B836A0" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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. However, Costa (1991a) did not provide characters for distinguishing
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<taxonomicName id="36CE0275B98854553C39246A6839A0DD" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="compacta">P. compacta</taxonomicName>
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and
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<taxonomicName id="63A489262B7E28DCFAB07AC9C33F9A11" LSID-ZBK="urn:lsid:zoobank.org:act:D03646D6-94E9-4B78-9DB1-193B232214C4" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna poranga Costa 1989:226" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="poranga">P. poranga</taxonomicName>
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, due to the poor condition of the three small female specimens in the type series of
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<taxonomicName id="D7A67C1B28C023F75C7C942A59A8C7D2" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="compacta">P. compacta</taxonomicName>
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, but noted that females of
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<taxonomicName id="4927D49D039DFE0427B1442ED797B7B8" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="compacta">P. compacta</taxonomicName>
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have isolated spots on the flanks, while females of
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<taxonomicName id="3FA94C016314CD4B49ED23E10CD9DB01" LSID-ZBK="urn:lsid:zoobank.org:act:D03646D6-94E9-4B78-9DB1-193B232214C4" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna poranga Costa 1989:226" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="poranga">P. poranga</taxonomicName>
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have flank spots fused to form a distinct color pattern.
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</paragraph>
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<paragraph id="90EC7E8A39EE66EAC4C5E485AEF5EC07" pageNumber="3">
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Additional collections of
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<taxonomicName id="8DFDC158F90F1013D4A823EA08F7DA85" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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were made from the Araguaia-Tocantins River basin, making possible a first revision of the genus, which included, for the first time, data on osteology and frontal squamation pattern (Costa, 1998b). However, based upon similarities of most morphological features (except for significant variation of female color patterns among different populations), Costa (1998b) considered
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<taxonomicName id="C532987231F73C9FD7394F5C6DEC9BB7" LSID-ZBK="urn:lsid:zoobank.org:act:D03646D6-94E9-4B78-9DB1-193B232214C4" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna poranga Costa 1989:226" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="poranga">P. poranga</taxonomicName>
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to be a synonym of
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<taxonomicName id="7E1AF9DC1E7857F6B61A622F83117B49" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="compacta">P. compacta</taxonomicName>
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, and
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<taxonomicName id="5463CAC6C29FD546DB288CCDB6B9CA1F" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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thus a monotypic genus.
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</paragraph>
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<paragraph id="96B343CE9DD50416F331116978E307E9" pageNumber="3">
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<taxonomicName id="4E76295720306629041BE33A0A8AE492" LSID-ZBK="urn:lsid:zoobank.org:act:81D9E5F6-D246-4FCD-BC27-6C3DE9A12A63" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias Costa 1989:193" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Plesiolebias Costa</taxonomicName>
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includes four valid species:
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<taxonomicName id="15A004EF58FB83B19B81D554F8A1B683" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="aruana">P. aruana (Lazara)</taxonomicName>
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, from the rio Araguaia basin;
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<taxonomicName id="1F60CCD1936C7ADC1ABD660997CB42B5" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="glaucopterus">P. glaucopterus (Costa & Lacerda)</taxonomicName>
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, from the rio Paraguay basin;
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<taxonomicName id="18CBA676D697C7B4657173EC15C0F399" LSID-ZBK="urn:lsid:zoobank.org:act:C369282A-8BDD-44FD-9057-EDD9185ED9B3" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias lacerdai Costa 1989:196" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="lacerdai">P. lacerdai Costa</taxonomicName>
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, from the rio das Mortes floodplains; and
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<taxonomicName id="C679B1397D8C0150777B695BF56CDBDA" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="xavantei">P. xavantei (Costa, Lacerda & Tanizaki)</taxonomicName>
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, from the rio Tocantins basin (Costa, 1998c). All species were described after 1986, and the genus was recently revised (Costa, 1998c). In this revision, the geographic distributions of some species were expanded, based on samples containing small, poorly preserved specimens. For example,
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<taxonomicName id="841606105CB4895AA764ABA70AC021E6" LSID-ZBK="urn:lsid:zoobank.org:act:C369282A-8BDD-44FD-9057-EDD9185ED9B3" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias lacerdai Costa 1989:196" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="lacerdai">P. lacerdai</taxonomicName>
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, first known only from the rio das Mortes floodplains, was recorded from an area of the middle rio Araguaia basin near ilha do Bananal; and
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<taxonomicName id="0AE7595F29536AEE9A395AC84B31B15C" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="aruana">P. aruana</taxonomicName>
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, previously known from localities along the rio Araguaia floodplains, was recorded from the middle rio Tocantins basin.
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</paragraph>
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<paragraph id="2446EBD5184BE5C32DE5AEAD07F74F3B" pageNumber="3">
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<taxonomicName id="2D0AB762E90F62B4DEF27F1866F56DF6" LSID-ZBK="urn:lsid:zoobank.org:act:A083017D-552D-4D60-AAC9-76E17E602498" class="Actinopterygii" family="Rivulidae" genus="Maratecoara" kingdom="Animalia" lsidName-ZBK="Maratecoara Costa 1995:68" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Maratecoara</taxonomicName>
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comprises
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<taxonomicName id="8EEF0544D1C8CE906ECCC3142F5B5E54" class="Actinopterygii" family="Rivulidae" genus="Maratecoara" kingdom="Animalia" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="lacortei">M. lacortei (Lazara)</taxonomicName>
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, from the middle rio Araguaia basin, and
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<taxonomicName id="94E5DA2201AD984BCF6814FD3EF9BA21" LSID-ZBK="urn:lsid:zoobank.org:act:765C1627-EF34-4EA6-BCF0-D7EF6466D7E9" class="Actinopterygii" family="Rivulidae" genus="Maratecoara" kingdom="Animalia" lsidName-ZBK="Maratecoara formosa Costa & Brasil 1995:69" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="species" species="formosa">M. formosa Costa & Brasil</taxonomicName>
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, from the rio Tocantins basin (Costa, 1995a). Both species were known from few specimens and available descriptions are limited (Lazara, 1991; Costa, 1995a, 1995b).
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</paragraph>
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<paragraph id="3F5DE53DAE96CF7A996D881A3D4A25BA" pageNumber="3">
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Recent additional collections from poorly sampled areas, examination of live and preserved fully-developed adult specimens (not available in previous studies), and the study of new morphological characters provide an opportunity to further clarify the systematics of the genera
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<taxonomicName id="FD86DFBCE2BB202591CB5CCE612DC569" LSID-ZBK="urn:lsid:zoobank.org:act:9B0B7242-C581-472B-9727-F6C16CCCFD8A" class="Actinopterygii" family="Rivulidae" genus="Pituna" kingdom="Animalia" lsidName-ZBK="Pituna Costa 1989:225" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Pituna</taxonomicName>
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,
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<taxonomicName id="707D88FC7FABA8F103C2605C5E570397" LSID-ZBK="urn:lsid:zoobank.org:act:81D9E5F6-D246-4FCD-BC27-6C3DE9A12A63" class="Actinopterygii" family="Rivulidae" genus="Plesiolebias" kingdom="Animalia" lsidName-ZBK="Plesiolebias Costa 1989:193" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Plesiolebias</taxonomicName>
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and
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<taxonomicName id="9D73AD062B77E895F4650540CF7884CC" LSID-ZBK="urn:lsid:zoobank.org:act:A083017D-552D-4D60-AAC9-76E17E602498" class="Actinopterygii" family="Rivulidae" genus="Maratecoara" kingdom="Animalia" lsidName-ZBK="Maratecoara Costa 1995:68" order="Cyprinodontiformes" pageNumber="3" phylum="Chordata" rank="genus">Maratecoara</taxonomicName>
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, as presented here.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |