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<document id="ADA620CE095FACC6BBCEBCB76CF78809" ID-CLB-Dataset="21174" ID-DOI="10.11646/zootaxa.4629.3.1" ID-GBIF-Dataset="9419bc9b-dbe4-426c-a34e-f0034848ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995508" ID-ZooBank="1D47B452-0A40-4BC5-9BBA-C06DEB298C9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598059654494" checkinUser="felipe" docAuthor="Silva, Fernando A. B. &amp; Valois, Marcely" docDate="2019" docId="9D12E5573A5EFF9695F1FE6AFBCDC70D" docLanguage="en" docName="zootaxa.4629.3.1.pdf" docOrigin="Zootaxa 4629 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scybalocanthon acrianus Silva &amp; Valois 2019, new species" docType="treatment" docVersion="8" lastPageNumber="309" masterDocId="612B9D2F3A59FF9E9566F840FFF1C516" masterDocTitle="A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="341" masterPageNumber="301" pageNumber="308" updateTime="1698845243763" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C39679066971AF04148460F4460E2448">A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:role id="41565B404608A0D00DD5A82AE5E2A8D8">
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<mods:namePart id="96063576A691D24AEF0014FAB4245B91">Silva, Fernando A. B.</mods:namePart>
<mods:affiliation id="FDF1E619CEF4E51C77818EEF26A2372A">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil. E-mail: fernandoabsilva @ yahoo. com. br</mods:affiliation>
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<mods:namePart id="9BF98DDE710F62B588B736ED461B7256">Valois, Marcely</mods:namePart>
<mods:affiliation id="4A149C05BDF3AEA3B7FF7A81D04D63F1">Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<mods:title id="1469C5CE119465FBA3E35EE0732A4F56">Zootaxa</mods:title>
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<mods:date id="2DC94999F1DB1A070BAC8907E3E5710D">2019</mods:date>
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<mods:number id="3BFB219E32ACE1C6B40641501E077766">2019-07-08</mods:number>
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<mods:number id="8220B2BE752D039678A83718CC4E51FD">4629</mods:number>
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<mods:classification id="67F27F49BF989623E2C92E6D27FEC0D7">journal article</mods:classification>
<mods:identifier id="C3771264B5C9B71F1EA9400D38215AD5" type="CLB-Dataset">21174</mods:identifier>
<mods:identifier id="F998E4B42CC0D21C37D425A074C04AF7" type="DOI">10.11646/zootaxa.4629.3.1</mods:identifier>
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<treatment id="9D12E5573A5EFF9695F1FE6AFBCDC70D" ID-DOI="http://doi.org/10.5281/zenodo.4335169" ID-GBIF-Taxon="167293214" ID-Zenodo-Dep="4335169" LSID="urn:lsid:plazi:treatment:9D12E5573A5EFF9695F1FE6AFBCDC70D" httpUri="http://treatment.plazi.org/id/9D12E5573A5EFF9695F1FE6AFBCDC70D" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
<subSubSection id="5DA107CA3A5EFF9995F1FE6AFCD1C352" box="[151,800,1578,1605]" pageId="7" pageNumber="308" type="nomenclature">
<paragraph id="150454413A5EFF9995F1FE6AFCD1C352" blockId="7.[151,800,1578,1641]" box="[151,800,1578,1605]" pageId="7" pageNumber="308">
<heading id="4E4CE32D3A5EFF9995F1FE6AFCD1C352" bold="true" box="[151,800,1578,1605]" fontSize="11" level="1" pageId="7" pageNumber="308" reason="1">
<emphasis id="27CF88533A5EFF9995F1FE6AFCD1C352" bold="true" box="[151,800,1578,1605]" pageId="7" pageNumber="308">
<taxonomicName id="D2BB2FC23A5EFF9995F1FE6AFD72C352" ID-CoL="4W3L6" authority="Silva &amp; Valois" authorityName="Silva &amp; Valois" authorityYear="2019" box="[151,643,1578,1605]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="acrianus" status="sp. nov.">
<emphasis id="27CF88533A5EFF9995F1FE6AFE3BC352" bold="true" box="[151,458,1578,1605]" italics="true" pageId="7" pageNumber="308">Scybalocanthon acrianus</emphasis>
Silva &amp; Valois
</taxonomicName>
,
<taxonomicNameLabel id="3CFC35283A5EFF9997F6FE6AFCD1C352" box="[656,800,1578,1604]" pageId="7" pageNumber="308" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A5EFF9995F1FE0EFE39C37F" box="[151,456,1613,1641]" pageId="7" pageNumber="308" type="description">
<paragraph id="150454413A5EFF9995F1FE0EFE39C37F" blockId="7.[151,800,1578,1641]" box="[151,456,1613,1641]" pageId="7" pageNumber="308">
(
<figureCitation id="8D8048C43A5EFF9995F9FE0EFEF2C37E" box="[159,259,1614,1640]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="7" pageNumber="308">Figs. 1A</figureCitation>
,
<figureCitation id="8D8048C43A5EFF999469FE0EFEC3C37E" box="[271,306,1614,1640]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="7" pageNumber="308">3B</figureCitation>
,
<figureCitation id="8D8048C43A5EFF999458FE0EFE93C37E" box="[318,354,1614,1640]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="7" pageNumber="308">4A</figureCitation>
,
<figureCitation id="8D8048C43A5EFF999408FE0DFE63C37E" box="[366,402,1613,1640]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="7" pageNumber="308">5A</figureCitation>
,
<figureCitation id="8D8048C43A5EFF9994F8FE0EFE31C37F" box="[414,448,1614,1641]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="7" pageNumber="308">6C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A5EFF9995F1FED6FEB5C3C2" pageId="7" pageNumber="308" type="etymology">
<paragraph id="150454413A5EFF9995F1FED6FEB5C3C2" blockId="7.[151,1437,1686,2037]" pageId="7" pageNumber="308">
<emphasis id="27CF88533A5EFF9995F1FED6FEEFC3A6" bold="true" box="[151,286,1686,1712]" pageId="7" pageNumber="308">Etymology.</emphasis>
The specific name is a Latinization of the demonym for the Brazilian state of
<collectingRegion id="D77F9AA33A5EFF999113FED6FB5AC3A6" box="[1141,1195,1686,1712]" country="Brazil" name="Acre" pageId="7" pageNumber="308">Acre</collectingRegion>
and should be treated as an adjective.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A5EFF9995A1FEA0FA81C272" pageId="7" pageNumber="308" type="diagnosis">
<paragraph id="150454413A5EFF9995A1FEA0FA81C272" blockId="7.[151,1437,1686,2037]" pageId="7" pageNumber="308">
<emphasis id="27CF88533A5EFF9995A1FEA0FEB1C3E1" box="[199,320,1760,1784]" inLineHeading="true" pageId="7" pageNumber="308" reason="1">
<emphasis id="27CF88533A5EFF9995A1FEA0FEC8C3EE" bold="true" box="[199,313,1760,1784]" pageId="7" pageNumber="308">Diagnosis</emphasis>
.
</emphasis>
Specimens of
<taxonomicName id="D2BB2FC23A5EFF999482FEA0FD91C3EE" authorityName="Silva &amp; Valois" authorityYear="2019" box="[484,608,1760,1784]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="acrianus">
<emphasis id="27CF88533A5EFF999482FEA0FD91C3EE" box="[484,608,1760,1784]" italics="true" pageId="7" pageNumber="308">S. acrianus</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="D2BB2FC23A5EFF999639FEA0FC1FC3EE" authority="Silva &amp; Valois, 2019" authorityName="Silva &amp; Valois" authorityYear="2019" box="[863,1006,1760,1784]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="chamorroi" status="sp. nov.">
<emphasis id="27CF88533A5EFF999639FEA0FC1FC3EE" box="[863,1006,1760,1784]" italics="true" pageId="7" pageNumber="308">S. chamorroi</emphasis>
</taxonomicName>
<emphasis id="27CF88533A5EFF999693FEA0FB75C3EE" bold="true" box="[1013,1156,1760,1784]" pageId="7" pageNumber="308">
<taxonomicNameLabel id="3CFC35283A5EFF999693FEA0FB8EC3EE" box="[1013,1151,1760,1784]" pageId="7" pageNumber="308" rank="species">new species</taxonomicNameLabel>
.
</emphasis>
in body coloration (
<figureCitation id="8D8048C43A5EFF999002FEA0FF28C20A" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="7" pageNumber="308">Figs. 1A, F</figureCitation>
); eighth elytral stria lacking carina at the anterior portion; sigmoid shaped of the FLP sclerite (
<figureCitation id="8D8048C43A5EFF999186FF44FA91C20A" box="[1248,1376,1796,1820]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="7" pageNumber="308">Figs. 5A, E</figureCitation>
); and endophallus with microbristles right beside the FLP sclerite (
<figureCitation id="8D8048C43A5EFF999651FF68FC49C256" box="[823,952,1832,1856]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="7" pageNumber="308">Figs. 5A, E</figureCitation>
). However,
<taxonomicName id="D2BB2FC23A5EFF999158FF68FB4BC256" authorityName="Silva &amp; Valois" authorityYear="2019" box="[1086,1210,1832,1856]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="acrianus">
<emphasis id="27CF88533A5EFF999158FF68FB4BC256" box="[1086,1210,1832,1856]" italics="true" pageId="7" pageNumber="308">S. acrianus</emphasis>
</taxonomicName>
can be easily distinguished from
<taxonomicName id="D2BB2FC23A5EFF999457FF0CFE30C272" authorityName="Silva &amp; Valois" authorityYear="2019" box="[305,449,1868,1892]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="chamorroi">
<emphasis id="27CF88533A5EFF999457FF0CFE30C272" box="[305,449,1868,1892]" italics="true" pageId="7" pageNumber="308">S. chamorroi</emphasis>
</taxonomicName>
by the opaque body surface (
<figureCitation id="8D8048C43A5EFF99966EFF0CFCAEC272" box="[776,863,1868,1892]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="7" pageNumber="308">Fig. 1A</figureCitation>
), and weakly impressed elytral striae (
<figureCitation id="8D8048C43A5EFF99906DFF0CFA93C272" box="[1291,1378,1868,1892]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="7" pageNumber="308">Fig. 3B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A5EFF9695A1FF2DFCA7C455" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308" type="description">
<paragraph id="150454413A5EFF9695A1FF2DFCA7C455" blockId="7.[151,1437,1686,2037]" lastBlockId="8.[151,1437,152,539]" lastPageId="8" lastPageNumber="309" pageId="7" pageNumber="308">
<emphasis id="27CF88533A5EFF9995A1FF2DFE53C29E" bold="true" box="[199,418,1901,1928]" pageId="7" pageNumber="308">Description. Body.</emphasis>
Oval, lateral edges rounded. Surface opaque, completely microgranulate.
<emphasis id="27CF88533A5EFF9991B0FF2DFAEBC29E" bold="true" box="[1238,1306,1901,1928]" pageId="7" pageNumber="308">Color</emphasis>
. Pronotum, lateral portion of hypomera, and legs yellow or light brown. Other parts dark brown.
<emphasis id="27CF88533A5EFF999121FFD2FB52C2BA" bold="true" box="[1095,1187,1938,1964]" pageId="7" pageNumber="308">Length.</emphasis>
6.0–7.0 mm.
<emphasis id="27CF88533A5EFF99905BFFD2FA67C2BA" bold="true" box="[1341,1430,1938,1964]" pageId="7" pageNumber="308">Thorax</emphasis>
. Anterior angles of pronotum approximately 85°. Lateral margin regularly curved outward, not forming an angle at the middle portion.
<emphasis id="27CF88533A5EFF999417FF9AFE4CC2E2" bold="true" box="[369,445,2010,2036]" pageId="7" pageNumber="308">Elytra</emphasis>
. Striae thin and shiny, weakly impressed (
<figureCitation id="8D8048C43A5EFF9996F7FF9AFC16C2E2" box="[913,999,2010,2036]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="7" pageNumber="308">Fig. 3B</figureCitation>
), punctures inconspicuous. Eighth stria effaced at the basal third, lacking carina at the anterior portion.
<emphasis id="27CF88533A51FF969639F8D8FC2BC5A5" bold="true" box="[863,986,152,179]" pageId="8" pageNumber="309">Aedeagus.</emphasis>
Parameres symmetrical, obliquely truncate apically (
<figureCitation id="8D8048C43A51FF969453F8FDFE61C5C1" box="[309,400,189,215]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="8" pageNumber="309">Fig. 4A</figureCitation>
). Dorsal margin of parameres curved inward. Ventral margin of parameres curved inward at the basal portion and substraight from the medial to apical portions. SRP circular, with curved handle-shaped extension (
<figureCitation id="8D8048C43A51FF9695B2F945FEDFC409" box="[212,302,260,287]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="8" pageNumber="309">Fig. 5A</figureCitation>
). FLP sigmoid shaped, with a set of microbristles right beside it (
<figureCitation id="8D8048C43A51FF969699F945FBA9C409" box="[1023,1112,260,287]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="8" pageNumber="309">Fig. 5A</figureCitation>
). A+SA with two superposed and elongate sclerites (
<figureCitation id="8D8048C43A51FF9694FEF969FE03C455" box="[408,498,296,323]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="8" pageNumber="309">Fig. 5A</figureCitation>
).
<collectingCountry id="6DAC14D13A51FF969762F969FDD9C455" box="[516,552,297,323]" name="American Samoa" pageId="8" pageNumber="309">AS</collectingCountry>
sigmoid-shaped (
<figureCitation id="8D8048C43A51FF969789F969FCBBC455" box="[751,842,296,323]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="8" pageNumber="309">Fig. 5A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A51FF9695A1F90DFCD1C4C5" pageId="8" pageNumber="309" type="materials_examined">
<paragraph id="150454413A51FF9695A1F90DFCD1C4C5" blockId="8.[151,1437,152,539]" pageId="8" pageNumber="309">
<emphasis id="27CF88533A51FF9695A1F90DFE87C471" bold="true" box="[199,374,332,359]" pageId="8" pageNumber="309">Type material.</emphasis>
<materialsCitation id="A5D35E1C3A51FF969419F90DFD38C49D" ID-GBIF-Occurrence="2847261304" collectingDate="1997-02" collectionCode="CEMT" country="Brazil" location="Rio Branco" pageId="8" pageNumber="309" specimenCount="1" specimenCount-male="1" stateProvince="Acre" typeStatus="holotype">
<emphasis id="27CF88533A51FF969419F90DFD81C471" bold="true" box="[383,624,332,359]" pageId="8" pageNumber="309">
<typeStatus id="CA00EAE33A51FF969419F90DFE1FC471" box="[383,494,333,359]" pageId="8" pageNumber="309" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="6DAC14D13A51FF96949DF90CFD9AC471" box="[507,619,332,359]" name="Brazil" pageId="8" pageNumber="309">BRAZIL</collectingCountry>
:
</emphasis>
<collectingRegion id="D77F9AA33A51FF96971EF90DFD36C471" box="[632,711,333,359]" country="Brazil" name="Acre" pageId="8" pageNumber="309">ACRE</collectingRegion>
:
<location id="1064029A3A51FF9697B5F90DFCA9C471" LSID="urn:lsid:plazi:treatment:9D12E5573A5EFF9695F1FE6AFBCDC70D:1064029A3A51FF9697B5F90DFCA9C471" box="[723,856,333,359]" country="Brazil" name="Rio Branco" pageId="8" pageNumber="309" stateProvince="Acre">Rio Branco</location>
, UFAC [Universidade Federal do
<collectingRegion id="D77F9AA33A51FF9691BAF90CFAE9C471" box="[1244,1304,332,359]" country="Brazil" name="Acre" pageId="8" pageNumber="309">Acre</collectingRegion>
], secondary forest,
<date id="610572813A51FF96958FF931FECBC49D" box="[233,314,369,395]" pageId="8" pageNumber="309" value="1997-02">
<collectingDate id="71418B693A51FF96958FF931FECBC49D" box="[233,314,369,395]" pageId="8" pageNumber="309" value="1997-02">ii.1997</collectingDate>
</date>
, F.Z. Vaz-de-Mello—
<specimenCount id="03BD9FC83A51FF969759F931FDABC49D" box="[575,602,369,395]" pageId="8" pageNumber="309" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A51FF96970BF931FD30C49D" box="[621,705,369,395]" pageId="8" pageNumber="309">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A5D35E1C3A51FF9697B1F931FBCBC4B9" ID-GBIF-Occurrence="2847261475" collectingDate="1997-02" collectionCode="CEMT" country="Brazil" location="Rio Branco" pageId="8" pageNumber="309" specimenCount="3" specimenCount-female="3" stateProvince="Acre" typeStatus="paratype">
<emphasis id="27CF88533A51FF9697B1F931FC27C49D" bold="true" box="[727,982,368,395]" pageId="8" pageNumber="309">
<typeStatus id="CA00EAE33A51FF9697B1F931FCA0C49D" box="[727,849,369,395]" pageId="8" pageNumber="309" type="paratype">Paratypes</typeStatus>
.
<collectingCountry id="6DAC14D13A51FF969607F930FC20C49D" box="[865,977,368,395]" name="Brazil" pageId="8" pageNumber="309">BRAZIL</collectingCountry>
:
</emphasis>
<collectingRegion id="D77F9AA33A51FF969687F931FBC1C49D" box="[993,1072,369,395]" country="Brazil" name="Acre" pageId="8" pageNumber="309">ACRE</collectingRegion>
:
<location id="1064029A3A51FF969159F931FB37C49D" LSID="urn:lsid:plazi:treatment:9D12E5573A5EFF9695F1FE6AFBCDC70D:1064029A3A51FF969159F931FB37C49D" box="[1087,1222,369,395]" country="Brazil" name="Rio Branco" pageId="8" pageNumber="309" stateProvince="Acre">Rio Branco</location>
, UFAC [Universidade Federal do
<collectingRegion id="D77F9AA33A51FF969436F9D4FE7DC4B9" box="[336,396,404,431]" country="Brazil" name="Acre" pageId="8" pageNumber="309">Acre</collectingRegion>
], secondary forest,
<date id="610572813A51FF969703F9D5FD47C4B9" box="[613,694,405,431]" pageId="8" pageNumber="309" value="1997-02">
<collectingDate id="71418B693A51FF969703F9D5FD47C4B9" box="[613,694,405,431]" pageId="8" pageNumber="309" value="1997-02">ii.1997</collectingDate>
</date>
, F.Z. Vaz-de-Mello—
<specimenCount id="03BD9FC83A51FF9696D0F9D5FC3EC4B9" box="[950,975,405,431]" pageId="8" pageNumber="309" type="female">3♀</specimenCount>
(
<collectionCode id="73AACC843A51FF9696B8F9D5FBC3C4B9" box="[990,1074,405,431]" pageId="8" pageNumber="309">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A5D35E1C3A51FF969123F9D4FCECC4C5" ID-GBIF-Occurrence="2847261432" collectingDate="1992-11" collectionCode="CMNC" country="Bolivia" latitude="-12.516666" location="Guanay" longLatPrecision="925" longitude="-66.825" pageId="8" pageNumber="309" specimenCount="1" specimenCount-male="1" stateProvince="Beni" typeStatus="paratype">
<emphasis id="27CF88533A51FF969123F9D4FB4FC4B9" bold="true" box="[1093,1214,404,431]" pageId="8" pageNumber="309">
<collectingCountry id="6DAC14D13A51FF969123F9D4FB4FC4B9" box="[1093,1214,404,431]" name="Bolivia" pageId="8" pageNumber="309">BOLIVIA</collectingCountry>
</emphasis>
: EL
<collectingRegion id="D77F9AA33A51FF969190F9D5FACBC4B9" box="[1270,1338,405,431]" country="Bolivia" name="Beni" pageId="8" pageNumber="309">BENI</collectingRegion>
,
<location id="1064029A3A51FF969023F9D5FA6DC4B9" LSID="urn:lsid:plazi:treatment:9D12E5573A5EFF9695F1FE6AFBCDC70D:1064029A3A51FF969023F9D5FA6DC4B9" box="[1349,1436,405,431]" country="Bolivia" latitude="-12.516666" longLatPrecision="925" longitude="-66.825" name="Guanay" pageId="8" pageNumber="309" stateProvince="Beni">Guanay</location>
(
<geoCoordinate id="708F32863A51FF9695C6F9F9FF0FC4C5" box="[160,254,441,467]" degrees="12" direction="south" minutes="31" orientation="latitude" pageId="8" pageNumber="309" precision="925" value="-12.516666">12°31’S</geoCoordinate>
,
<geoCoordinate id="708F32863A51FF96946DF9F9FE50C4C2" box="[267,417,441,468]" degrees="66" direction="west" minutes="49" orientation="longitude" pageId="8" pageNumber="309" precision="15" seconds="30" value="-66.825">66°49’30’’W</geoCoordinate>
),
<date id="610572813A51FF9694D3F9F9FDFDC4C5" box="[437,524,441,467]" pageId="8" pageNumber="309" value="1992-11">
<collectingDate id="71418B693A51FF9694D3F9F9FDFDC4C5" box="[437,524,441,467]" pageId="8" pageNumber="309" value="1992-11">xi.1992</collectingDate>
</date>
, L. Peña—
<specimenCount id="03BD9FC83A51FF9697F7F9F9FD5CC4C5" box="[657,685,441,467]" pageId="8" pageNumber="309" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A51FF9697DAF9F9FCE4C4C5" box="[700,789,441,467]" pageId="8" pageNumber="309">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A51FF9695A1F99CFBCDC70D" pageId="8" pageNumber="309" type="distribution">
<paragraph id="150454413A51FF9695A1F99CFBCDC70D" blockId="8.[151,1437,152,539]" pageId="8" pageNumber="309">
<emphasis id="27CF88533A51FF9695A1F99CFE90C4E1" bold="true" box="[199,353,476,503]" pageId="8" pageNumber="309">Distribution.</emphasis>
Known from
<collectingCountry id="6DAC14D13A51FF969762F99DFDB9C4E1" box="[516,584,477,503]" name="Brazil" pageId="8" pageNumber="309">Brazil</collectingCountry>
(
<collectingRegion id="D77F9AA33A51FF96973FF99DFD62C4E1" box="[601,659,477,503]" country="Brazil" name="Acre" pageId="8" pageNumber="309">Acre</collectingRegion>
) and
<collectingCountry id="6DAC14D13A51FF9697B2F99DFCD7C4E1" box="[724,806,477,503]" name="Bolivia" pageId="8" pageNumber="309">Bolivia</collectingCountry>
(El
<collectingRegion id="D77F9AA33A51FF96963CF99DFC61C4EE" box="[858,912,477,504]" country="Bolivia" name="Beni" pageId="8" pageNumber="309">Beni</collectingRegion>
) (
<figureCitation id="8D8048C43A51FF9696CFF99DFBF5C4EE" box="[937,1028,477,504]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="8" pageNumber="309">Fig. 6C</figureCitation>
).
<emphasis id="27CF88533A51FF96917FF99CFB10C4E1" bold="true" box="[1049,1249,476,503]" pageId="8" pageNumber="309">Endemism areas</emphasis>
:
<emphasis id="27CF88533A51FF969194F99CFF12C70D" bold="true" pageId="8" pageNumber="309">Brazilian subregion</emphasis>
: South Brazilian dominion: Rondônia province (see
<bibRefCitation id="712A29B03A51FF969655FA41FC29C70D" author="Morrone, J. J." box="[819,984,513,539]" pageId="8" pageNumber="309" pagination="1 - 110" refId="ref47051" refString="Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
; fig. 12).
</paragraph>
</subSubSection>
</treatment>
</document>
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