296 lines
49 KiB
XML
296 lines
49 KiB
XML
<document id="AF700677AB0BF6587DCA5FA56D5107A2" ID-CLB-Dataset="21174" ID-DOI="10.11646/zootaxa.4629.3.1" ID-GBIF-Dataset="9419bc9b-dbe4-426c-a34e-f0034848ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995508" ID-ZooBank="1D47B452-0A40-4BC5-9BBA-C06DEB298C9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598059654494" checkinUser="felipe" docAuthor="Silva, Fernando A. B. & Valois, Marcely" docDate="2019" docId="9D12E5573A4DFF8A95F1F969FD0AC339" docLanguage="en" docName="zootaxa.4629.3.1.pdf" docOrigin="Zootaxa 4629 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scybalocanthon federicoescobari Silva & Valois 2019, new species" docType="treatment" docVersion="8" lastPageNumber="321" masterDocId="612B9D2F3A59FF9E9566F840FFF1C516" masterDocTitle="A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="341" masterPageNumber="301" pageNumber="321" updateTime="1698845243763" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B2B03DC540C09122A9BD2A4E56619C42">
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<mods:title id="52FCBB724DE58FC1773BE3FD207F3CFC">A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:name id="A25DE5D3038A6700E4C0F549204C533F" type="personal">
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<mods:roleTerm id="E6155CC51AC291B043CC3987455CA13E">Author</mods:roleTerm>
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<mods:namePart id="9EC9F395089B18C4465118733E831CD2">Silva, Fernando A. B.</mods:namePart>
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<mods:affiliation id="784DF44620E7BF111A6CBC84E86B917F">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil. E-mail: fernandoabsilva @ yahoo. com. br</mods:affiliation>
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<mods:namePart id="0D87D11F06515F15CBAF6F454D707B51">Valois, Marcely</mods:namePart>
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<mods:affiliation id="1E20C93B3E96C6247CA821A44AEE654F">Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<mods:title id="3A8E85855E39C7CCA7F289E78EA6944D">Zootaxa</mods:title>
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<mods:part id="6AAE230753F09AA0A39B89AD22EB4042">
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<mods:date id="3F6642144749F1DA05ED57529E8F2423">2019</mods:date>
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<mods:number id="5891623A5CDA279253DBDA9432AD7F64">2019-07-08</mods:number>
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<mods:number id="6DEA7FABAA1F4897E27946CC7B589CEB">4629</mods:number>
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<mods:identifier id="5BE3193D7F1AA57C561F625AC5B8D53C" type="CLB-Dataset">21174</mods:identifier>
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<mods:identifier id="D7C59CF5D14A3DA8BA64AAFA06703CDB" type="DOI">10.11646/zootaxa.4629.3.1</mods:identifier>
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<treatment id="9D12E5573A4DFF8A95F1F969FD0AC339" ID-DOI="http://doi.org/10.5281/zenodo.4335180" ID-GBIF-Taxon="167293225" ID-Zenodo-Dep="4335180" LSID="urn:lsid:plazi:treatment:9D12E5573A4DFF8A95F1F969FD0AC339" httpUri="http://treatment.plazi.org/id/9D12E5573A4DFF8A95F1F969FD0AC339" lastPageNumber="321" pageId="20" pageNumber="321">
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<subSubSection id="5DA107CA3A4DFF8A95F1F969FC8CC455" box="[151,893,297,324]" pageId="20" pageNumber="321" type="nomenclature">
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<paragraph id="150454413A4DFF8A95F1F969FC8CC455" blockId="20.[151,893,297,360]" box="[151,893,297,324]" pageId="20" pageNumber="321">
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<heading id="4E4CE32D3A4DFF8A95F1F969FC8CC455" bold="true" box="[151,893,297,324]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
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<emphasis id="27CF88533A4DFF8A95F1F969FC8CC455" bold="true" box="[151,893,297,324]" pageId="20" pageNumber="321">
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<taxonomicName id="D2BB2FC23A4DFF8A95F1F969FD11C455" ID-CoL="4W3LH" authority="Silva & Valois" authorityName="Silva & Valois" authorityYear="2019" box="[151,736,297,324]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="federicoescobari" status="sp. nov.">
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<emphasis id="27CF88533A4DFF8A95F1F969FDD6C452" bold="true" box="[151,551,297,324]" italics="true" pageId="20" pageNumber="321">Scybalocanthon federicoescobari</emphasis>
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Silva & Valois
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</taxonomicName>
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,
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<taxonomicNameLabel id="3CFC35283A4DFF8A978BF969FC8CC455" box="[749,893,297,323]" pageId="20" pageNumber="321" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="5DA107CA3A4DFF8A95F1F90DFE74C47E" box="[151,389,332,360]" pageId="20" pageNumber="321" type="description">
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<paragraph id="150454413A4DFF8A95F1F90DFE74C47E" blockId="20.[151,893,297,360]" box="[151,389,332,360]" pageId="20" pageNumber="321">
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(
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<figureCitation id="8D8048C43A4DFF8A95F9F90DFF09C471" box="[159,248,333,359]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="20" pageNumber="321">Figs. 1I</figureCitation>
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,
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<figureCitation id="8D8048C43A4DFF8A9462F90DFEEFC471" box="[260,286,333,359]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">4I</figureCitation>
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,
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<figureCitation id="8D8048C43A4DFF8A944FF90CFEBFC471" box="[297,334,332,359]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="20" pageNumber="321">5H</figureCitation>
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,
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<figureCitation id="8D8048C43A4DFF8A943CF90DFE8CC47E" box="[346,381,333,360]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="20" pageNumber="321">6D</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="5DA107CA3A4DFF8A95F1F9D5FEE5C4C5" pageId="20" pageNumber="321" type="etymology">
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<paragraph id="150454413A4DFF8A95F1F9D5FEE5C4C5" blockId="20.[151,1437,405,1583]" pageId="20" pageNumber="321">
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<emphasis id="27CF88533A4DFF8A95F1F9D5FEEFC4B9" bold="true" box="[151,286,405,431]" pageId="20" pageNumber="321">Etymology.</emphasis>
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Named in honor of Federico Escobar, a scarab ecologist who was responsible for collecting the
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<typeStatus id="CA00EAE33A4DFF8A900AF9D5FA6DC4B9" box="[1388,1436,405,431]" pageId="20" pageNumber="321">type</typeStatus>
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specimens.
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</paragraph>
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</subSubSection>
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<subSubSection id="5DA107CA3A4DFF8A95A1F99CFBCAC6B1" pageId="20" pageNumber="321" type="diagnosis">
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<paragraph id="150454413A4DFF8A95A1F99CFBCAC6B1" blockId="20.[151,1437,405,1583]" pageId="20" pageNumber="321">
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<emphasis id="27CF88533A4DFF8A95A1F99CFECBC4E1" bold="true" box="[199,314,476,503]" pageId="20" pageNumber="321">Diagnosis</emphasis>
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. Specimens of
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<taxonomicName id="D2BB2FC23A4DFF8A948BF99DFD36C4E1" authorityName="Silva & Valois" authorityYear="2019" box="[493,711,477,503]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="federicoescobari">
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<emphasis id="27CF88533A4DFF8A948BF99DFD36C4E1" box="[493,711,477,503]" italics="true" pageId="20" pageNumber="321">S. federicoescobari</emphasis>
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</taxonomicName>
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are similar to those of
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<taxonomicName id="D2BB2FC23A4DFF8A96B3F99DFBBAC4E1" authorityName="Silva & Valois" authorityYear="2019" box="[981,1099,477,503]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="arnaudi">
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<emphasis id="27CF88533A4DFF8A96B3F99DFBBAC4E1" box="[981,1099,477,503]" italics="true" pageId="20" pageNumber="321">S. arnaudi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="D2BB2FC23A4DFF8A913DF99DFB12C4E1" authorityName="Silva & Valois" authorityYear="2019" box="[1115,1251,477,503]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="martinezi">
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<emphasis id="27CF88533A4DFF8A913DF99DFB12C4E1" box="[1115,1251,477,503]" italics="true" pageId="20" pageNumber="321">S. martinezi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="D2BB2FC23A4DFF8A9195F99DFA89C4E1" authorityName="Silva & Valois" authorityYear="2019" box="[1267,1400,477,503]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="papaxibe">
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<emphasis id="27CF88533A4DFF8A9195F99DFA89C4E1" box="[1267,1400,477,503]" italics="true" pageId="20" pageNumber="321">S. papaxibe</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="D2BB2FC23A4DFF8A90E1F99DFEFEC70D" baseAuthorityName="Kirsch" baseAuthorityYear="1873" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pinopterus">
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<emphasis id="27CF88533A4DFF8A90E1F99DFEFEC70D" italics="true" pageId="20" pageNumber="321">S. pinopterus</emphasis>
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</taxonomicName>
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, and
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<taxonomicName id="D2BB2FC23A4DFF8A942DFA41FE02C70D" authorityName="Silva & Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[331,499,513,539]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="uniplagiatus">
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<emphasis id="27CF88533A4DFF8A942DFA41FE02C70D" box="[331,499,513,539]" italics="true" pageId="20" pageNumber="321">S. uniplagiatus</emphasis>
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</taxonomicName>
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by having the pronotum uniformly colored, or with one elliptical spot on the central portion (
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<figureCitation id="8D8048C43A4DFF8A9591FA65FEBFC729" box="[247,334,549,575]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="20" pageNumber="321">Figs. 1I</figureCitation>
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,
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<figureCitation id="8D8048C43A4DFF8A943CFA65FE8FC729" box="[346,382,549,575]" captionStart="FIGURE 2" captionStartId="9.[151,250,1178,1203]" captionTargetBox="[165,1424,188,1136]" captionTargetId="figure@9.[151,1436,181,1152]" captionTargetPageId="9" captionText="FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995514" httpUri="https://zenodo.org/record/3995514/files/figure.png" pageId="20" pageNumber="321">2A</figureCitation>
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,
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<figureCitation id="8D8048C43A4DFF8A94ECFA65FE56C729" box="[394,423,549,575]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="20" pageNumber="321">1P</figureCitation>
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,
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<figureCitation id="8D8048C43A4DFF8A94D7FA65FE27C729" box="[433,470,549,575]" captionStart="FIGURE 2" captionStartId="9.[151,250,1178,1203]" captionTargetBox="[165,1424,188,1136]" captionTargetId="figure@9.[151,1436,181,1152]" captionTargetPageId="9" captionText="FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995514" httpUri="https://zenodo.org/record/3995514/files/figure.png" pageId="20" pageNumber="321">2K</figureCitation>
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,
|
||
<figureCitation id="8D8048C43A4DFF8A9487FA65FDC0C729" box="[481,561,549,575]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="20" pageNumber="321">1D, 1T</figureCitation>
|
||
); femora almost completely yellow or brown, with black spots only on the tips (
|
||
<figureCitation id="8D8048C43A4DFF8A95C6FA09FF0AC775" box="[160,251,585,611]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="20" pageNumber="321">Fig. 3G</figureCitation>
|
||
); eighth elytral stria with thin carina on anterior portion (
|
||
<figureCitation id="8D8048C43A4DFF8A96E3FA09FC11C775" box="[901,992,585,611]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="20" pageNumber="321">Fig. 3D</figureCitation>
|
||
); endophallus with bristles right beside the FLP sclerite (
|
||
<figureCitation id="8D8048C43A4DFF8A9431FA2DFD99C791" box="[343,616,620,647]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="20" pageNumber="321">Figs. 5H, R, N, W, D, Q</figureCitation>
|
||
); and additional sclerite (
|
||
<collectingCountry id="6DAC14D13A4DFF8A96E2FA2DFC5BC791" box="[900,938,621,647]" name="American Samoa" pageId="20" pageNumber="321">AS</collectingCountry>
|
||
) absent (
|
||
<figureCitation id="8D8048C43A4DFF8A9176FA2DFAD3C791" box="[1040,1314,620,647]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="20" pageNumber="321">Figs. 5H, R, N, W, D, Q</figureCitation>
|
||
). They can be distinguished from those of
|
||
<taxonomicName id="D2BB2FC23A4DFF8A949DFAD1FD61C7BD" baseAuthorityName="Kirsch" baseAuthorityYear="1873" box="[507,656,657,683]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pinopterus">
|
||
<emphasis id="27CF88533A4DFF8A949DFAD1FD61C7BD" box="[507,656,657,683]" italics="true" pageId="20" pageNumber="321">S. pinopterus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="D2BB2FC23A4DFF8A97ACFAD1FCB1C7BD" authorityName="Silva & Valois" authorityYear="2019" box="[714,832,657,683]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="arnaudi">
|
||
<emphasis id="27CF88533A4DFF8A97ACFAD1FCB1C7BD" box="[714,832,657,683]" italics="true" pageId="20" pageNumber="321">S. arnaudi</emphasis>
|
||
</taxonomicName>
|
||
by the strongly asymmetrical parameres; left paramere with acute projection on dorsal margin and bilobate excavation on ventral margin (
|
||
<figureCitation id="8D8048C43A4DFF8A910FFAF5FB46C7D9" box="[1129,1207,693,719]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">Fig. 4I</figureCitation>
|
||
) (other species with slightly asymmetrical parameres, lacking acute projection and bilobate excavation on dorsal and ventral margins, respectively (
|
||
<figureCitation id="8D8048C43A4DFF8A944BFABDFE49C601" box="[301,440,765,791]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">Figs. 4E, 4S</figureCitation>
|
||
)); from those of
|
||
<taxonomicName id="D2BB2FC23A4DFF8A9713FABDFD0AC601" authorityName="Silva & Valois" authorityYear="2019" box="[629,763,765,791]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="27CF88533A4DFF8A9713FABDFD0AC601" box="[629,763,765,791]" italics="true" pageId="20" pageNumber="321">S. martinezi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="D2BB2FC23A4DFF8A966EFABDFC7AC601" authorityName="Silva & Valois" authorityYear="2019" box="[776,907,765,791]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="papaxibe">
|
||
<emphasis id="27CF88533A4DFF8A966EFABDFC7AC601" box="[776,907,765,791]" italics="true" pageId="20" pageNumber="321">S. papaxibe</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="D2BB2FC23A4DFF8A96AEFABDFB81C601" authorityName="Silva & Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[968,1136,765,791]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="uniplagiatus">
|
||
<emphasis id="27CF88533A4DFF8A96AEFABDFB81C601" box="[968,1136,765,791]" italics="true" pageId="20" pageNumber="321">S. uniplagiatus</emphasis>
|
||
</taxonomicName>
|
||
by the bilobate excavation of the ventral margin of the left paramere deeper than wide in lateral view, extending along one-fourth of the paramere margin (
|
||
<figureCitation id="8D8048C43A4DFF8A9455FB05FE71C649" box="[307,384,837,863]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">Fig. 4I</figureCitation>
|
||
) (in
|
||
<taxonomicName id="D2BB2FC23A4DFF8A94D2FB05FDCBC649" authorityName="Silva & Valois" authorityYear="2019" box="[436,570,837,863]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="27CF88533A4DFF8A94D2FB05FDCBC649" box="[436,570,837,863]" italics="true" pageId="20" pageNumber="321">S. martinezi</emphasis>
|
||
</taxonomicName>
|
||
the bilobate excavation is wider and deeper, extending at least one-half of the paramere margin (
|
||
<figureCitation id="8D8048C43A4DFF8A9405FB29FE4DC695" box="[355,444,873,899]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">Fig. 4O</figureCitation>
|
||
); in
|
||
<taxonomicName id="D2BB2FC23A4DFF8A948DFB29FD63C695" authorityName="Silva & Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[491,658,873,899]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="uniplagiatus">
|
||
<emphasis id="27CF88533A4DFF8A948DFB29FD63C695" box="[491,658,873,899]" italics="true" pageId="20" pageNumber="321">S. uniplagiatus</emphasis>
|
||
</taxonomicName>
|
||
its excavation extending along one-third of the paramere margin (
|
||
<figureCitation id="8D8048C43A4DFF8A9009FB29FF35C6B1" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">Fig. 4W</figureCitation>
|
||
); in
|
||
<taxonomicName id="D2BB2FC23A4DFF8A9592FBCDFE86C6B1" authorityName="Silva & Valois" authorityYear="2019" box="[244,375,909,935]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="papaxibe">
|
||
<emphasis id="27CF88533A4DFF8A9592FBCDFE86C6B1" box="[244,375,909,935]" italics="true" pageId="20" pageNumber="321">S. papaxibe</emphasis>
|
||
</taxonomicName>
|
||
its excavation is wider than deep in the lateral view (
|
||
<figureCitation id="8D8048C43A4DFF8A96AAFBCDFBD5C6B1" box="[972,1060,909,935]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">Fig. 4R</figureCitation>
|
||
)).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5DA107CA3A4DFF8A95A1FBF0FAC7C041" pageId="20" pageNumber="321" type="description">
|
||
<paragraph id="150454413A4DFF8A95A1FBF0FAC7C041" blockId="20.[151,1437,405,1583]" pageId="20" pageNumber="321">
|
||
<emphasis id="27CF88533A4DFF8A95A1FBF0FE59C6DD" bold="true" box="[199,424,944,971]" pageId="20" pageNumber="321">Description. Body.</emphasis>
|
||
Oval-elongate. Surface opaque, completely microgranulate.
|
||
<emphasis id="27CF88533A4DFF8A910CFBF0FB5FC6DD" bold="true" box="[1130,1198,944,971]" pageId="20" pageNumber="321">Color</emphasis>
|
||
. Most of pronotum, hypomera, metaepisternae, metaventrite, abdominal ventrites, pygidium, and femora yellow or light brown. Head, elytra, prosternum, mesoventrite, mesoepisternae, trochanters, and tibiae dark brown.
|
||
<emphasis id="27CF88533A4DFF8A9101FBB9FB32C105" bold="true" box="[1127,1219,1017,1043]" pageId="20" pageNumber="321">Length.</emphasis>
|
||
<quantity id="D243F9A43A4DFF8A91AAFBB8FAA6C105" box="[1228,1367,1016,1043]" metricMagnitude="-3" metricUnit="m" metricValue="8.65" metricValueMax="9.5" metricValueMin="7.8" pageId="20" pageNumber="321" unit="mm" value="8.65" valueMax="9.5" valueMin="7.8">7.8–9.5 mm</quantity>
|
||
.
|
||
<emphasis id="27CF88533A4DFF8A9005FBB9FF4EC121" bold="true" pageId="20" pageNumber="321">Thorax</emphasis>
|
||
. Pronotum with one brown rounded spot at the anterocentral portion. Spot elliptical in some specimens.Anterior angles of pronotum approximately 85°. Lateral margin regularly curved outward, not forming an angle at the middle portion.
|
||
<emphasis id="27CF88533A4DFF8A9591FC25FEB2C169" bold="true" box="[247,323,1125,1151]" pageId="20" pageNumber="321">Elytra</emphasis>
|
||
. Striae thin and shiny, punctures inconspicuous. Eighth stria with a thin carina at the anterior portion.
|
||
<emphasis id="27CF88533A4DFF8A95B4FCC8FEBDC1B5" bold="true" box="[210,332,1160,1187]" pageId="20" pageNumber="321">Aedeagus.</emphasis>
|
||
Parameres strongly asymmetrical (
|
||
<figureCitation id="8D8048C43A4DFF8A97BDFCC9FCDAC1B5" box="[731,811,1161,1187]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="20" pageNumber="321">Fig. 4I</figureCitation>
|
||
). Dorsal margin of right paramere curved inward, apex obliquely truncate. Ventral margin of right paramere substraight, with a rounded excavation at the basal portion. Dorsal margin of left paramere curved inward from the basal to medial portions, apical portion with a short and pointed projection. Ventral margin of left paramere substraight, with a bilobate excavation at the basal portion, apical portion pointed. SRP circular, with rounded handle-shaped extension (
|
||
<figureCitation id="8D8048C43A4DFF8A96AAFD59FBD7C025" box="[972,1062,1304,1331]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="20" pageNumber="321">Fig. 5H</figureCitation>
|
||
). FLP short, comma-shaped, with three sets of bristles right beside it (
|
||
<figureCitation id="8D8048C43A4DFF8A9741FD7DFD70C041" box="[551,641,1340,1367]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="20" pageNumber="321">Fig. 5H</figureCitation>
|
||
). A+SA with two superposed and elongate sclerites (
|
||
<figureCitation id="8D8048C43A4DFF8A91A9FD7DFADBC041" box="[1231,1322,1340,1367]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="20" pageNumber="321">Fig. 5H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5DA107CA3A4DFF8A95A1FD21FE98C0F1" pageId="20" pageNumber="321" type="materials_examined">
|
||
<paragraph id="150454413A4DFF8A95A1FD21FE98C0F1" blockId="20.[151,1437,405,1583]" pageId="20" pageNumber="321">
|
||
<emphasis id="27CF88533A4DFF8A95A1FD21FE8EC06D" bold="true" box="[199,383,1376,1403]" pageId="20" pageNumber="321">Type material.</emphasis>
|
||
<materialsCitation id="A5D35E1C3A4DFF8A94E9FD21FBF3C089" ID-GBIF-Occurrence="2847261485" collectingDate="1996-02" collectionCode="CEMT" country="Colombia" elevation="250" latitude="2.176389" location="Cerro Moyano" longLatPrecision="21" longitude="-71.18278" municipality="Reserve Nukak" pageId="20" pageNumber="321" specimenCount="1" specimenCount-male="1" stateProvince="Guaviare" typeStatus="holotype">
|
||
<emphasis id="27CF88533A4DFF8A94E9FD21FD4DC06D" bold="true" box="[399,700,1376,1403]" pageId="20" pageNumber="321">
|
||
<typeStatus id="CA00EAE33A4DFF8A94E9FD21FE0FC06D" box="[399,510,1377,1403]" pageId="20" pageNumber="321" type="holotype">Holotype</typeStatus>
|
||
.
|
||
<collectingCountry id="6DAC14D13A4DFF8A9772FD20FD46C06D" box="[532,695,1376,1403]" name="Colombia" pageId="20" pageNumber="321">COLOMBIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
<collectingRegion id="D77F9AA33A4DFF8A97ABFD21FCAEC06D" box="[717,863,1377,1403]" country="Colombia" name="Guaviare" pageId="20" pageNumber="321">GUAVIARE</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="F560CE3B3A4DFF8A9615FD21FBDDC06D" box="[883,1068,1377,1403]" pageId="20" pageNumber="321">Reserve Nukak</collectingMunicipality>
|
||
,
|
||
<location id="1064029A3A4DFF8A9127FD21FB1FC06D" LSID="urn:lsid:plazi:treatment:9D12E5573A4DFF8A95F1F969FD0AC339:1064029A3A4DFF8A9127FD21FB1FC06D" box="[1089,1262,1377,1403]" country="Colombia" latitude="2.176389" longLatPrecision="21" longitude="-71.18278" municipality="Reserve Nukak" name="Cerro Moyano" pageId="20" pageNumber="321" stateProvince="Guaviare">Cerro Moyano</location>
|
||
(
|
||
<geoCoordinate id="708F32863A4DFF8A906EFD20FA69C06D" box="[1288,1432,1376,1403]" degrees="71" direction="west" minutes="10" orientation="longitude" pageId="20" pageNumber="321" precision="15" seconds="58" value="-71.18278">71°10’58”W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="708F32863A4DFF8A95F1FDC4FED2C089" box="[151,291,1412,1439]" degrees="02" direction="north" minutes="10" orientation="latitude" pageId="20" pageNumber="321" precision="15" seconds="35" value="2.176389">02°10’35”N</geoCoordinate>
|
||
),
|
||
<date id="610572813A4DFF8A945CFDC5FE7FC089" box="[314,398,1413,1439]" pageId="20" pageNumber="321" value="1996-02">
|
||
<collectingDate id="71418B693A4DFF8A945CFDC5FE7FC089" box="[314,398,1413,1439]" pageId="20" pageNumber="321" value="1996-02">II.1996</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="CCFA2C563A4DFF8A94FDFDC5FE2FC089" box="[411,478,1413,1439]" pageId="20" pageNumber="321">pitfall</collectingMethod>
|
||
human feces,
|
||
<quantity id="D243F9A43A4DFF8A97E0FDC4FD20C089" box="[646,721,1412,1439]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="321" unit="m" value="250.0">
|
||
<elevation id="9E96B3723A4DFF8A97E0FDC4FD20C089" box="[646,721,1412,1439]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="321" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
, F. Escobar—
|
||
<specimenCount id="03BD9FC83A4DFF8A961CFDC5FC64C089" box="[890,917,1413,1439]" pageId="20" pageNumber="321" type="male">1♂</specimenCount>
|
||
(
|
||
<collectionCode id="73AACC843A4DFF8A96C0FDC5FC0BC089" box="[934,1018,1413,1439]" pageId="20" pageNumber="321">CEMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="A5D35E1C3A4DFF8A9168FDC5FE94C0F1" ID-GBIF-Occurrence="2847261335" collectingDate="1996-02" collectionCode="CEMT" country="Colombia" elevation="250" latitude="2.176389" location="Cerro Moyano" longLatPrecision="21" longitude="-71.18278" municipality="Reserve Nukak" pageId="20" pageNumber="321" specimenCount="1" specimenCount-female="1" stateProvince="ARE" typeStatus="paratype">
|
||
<emphasis id="27CF88533A4DFF8A9168FDC5FB8AC089" bold="true" box="[1038,1147,1413,1439]" pageId="20" pageNumber="321">
|
||
<typeStatus id="CA00EAE33A4DFF8A9168FDC5FB8AC089" box="[1038,1147,1413,1439]" pageId="20" pageNumber="321" type="paratype">Paratype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="27CF88533A4DFF8A91EDFDC4FAC2C089" bold="true" box="[1163,1331,1412,1439]" pageId="20" pageNumber="321">
|
||
<collectingCountry id="6DAC14D13A4DFF8A91EDFDC4FADFC089" box="[1163,1326,1412,1439]" name="Colombia" pageId="20" pageNumber="321">COLOMBIA</collectingCountry>
|
||
:
|
||
</emphasis>
|
||
GUAVI-
|
||
<collectingRegion id="D77F9AA33A4DFF8A95F1FDE9FF23C0D5" box="[151,210,1449,1475]" country="Colombia" name="ARE" pageId="20" pageNumber="321">ARE</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="F560CE3B3A4DFF8A9586FDE9FE60C0D5" box="[224,401,1449,1475]" pageId="20" pageNumber="321">Reserve Nukak</collectingMunicipality>
|
||
,
|
||
<location id="1064029A3A4DFF8A94F8FDE9FDB5C0D5" LSID="urn:lsid:plazi:treatment:9D12E5573A4DFF8A95F1F969FD0AC339:1064029A3A4DFF8A94F8FDE9FDB5C0D5" box="[414,580,1449,1475]" country="Colombia" latitude="2.176389" longLatPrecision="21" longitude="-71.18278" municipality="Reserve Nukak" name="Cerro Moyano" pageId="20" pageNumber="321" stateProvince="ARE">Cerro Moyano</location>
|
||
(
|
||
<geoCoordinate id="708F32863A4DFF8A9731FDE8FD16C0D5" box="[599,743,1448,1475]" degrees="71" direction="west" minutes="10" orientation="longitude" pageId="20" pageNumber="321" precision="15" seconds="58" value="-71.18278">71°10’58”W</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="708F32863A4DFF8A9793FDE8FC71C0D5" box="[757,896,1448,1475]" degrees="02" direction="north" minutes="10" orientation="latitude" pageId="20" pageNumber="321" precision="15" seconds="35" value="2.176389">02°10’35”N</geoCoordinate>
|
||
),
|
||
<date id="610572813A4DFF8A96F1FDE9FC1AC0D5" box="[919,1003,1449,1475]" pageId="20" pageNumber="321" value="1996-02">
|
||
<collectingDate id="71418B693A4DFF8A96F1FDE9FC1AC0D5" box="[919,1003,1449,1475]" pageId="20" pageNumber="321" value="1996-02">II.1996</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="CCFA2C563A4DFF8A969FFDE9FBCDC0D5" box="[1017,1084,1449,1475]" pageId="20" pageNumber="321">pitfall</collectingMethod>
|
||
human feces,
|
||
<quantity id="D243F9A43A4DFF8A9183FDE8FAC0C0D5" box="[1253,1329,1448,1475]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="321" unit="m" value="250.0">
|
||
<elevation id="9E96B3723A4DFF8A9183FDE8FAC0C0D5" box="[1253,1329,1448,1475]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="321" unit="m" value="250.0">250 m</elevation>
|
||
</quantity>
|
||
, F. Esco-bar—
|
||
<specimenCount id="03BD9FC83A4DFF8A9584FD8DFF0BC0F1" box="[226,250,1485,1511]" pageId="20" pageNumber="321" type="female">1♀</specimenCount>
|
||
(
|
||
<collectionCode id="73AACC843A4DFF8A946FFD8DFEACC0F1" box="[265,349,1485,1511]" pageId="20" pageNumber="321">CEMT</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5DA107CA3A4DFF8A95A1FDB0FD0AC339" pageId="20" pageNumber="321" type="distribution">
|
||
<paragraph id="150454413A4DFF8A95A1FDB0FD0AC339" blockId="20.[151,1437,405,1583]" pageId="20" pageNumber="321">
|
||
<emphasis id="27CF88533A4DFF8A95A1FDB0FE90C31D" bold="true" box="[199,353,1520,1547]" pageId="20" pageNumber="321">Distribution.</emphasis>
|
||
Known from
|
||
<collectingCountry id="6DAC14D13A4DFF8A976EFDB1FD87C31D" box="[520,630,1521,1547]" name="Colombia" pageId="20" pageNumber="321">Colombia</collectingCountry>
|
||
(
|
||
<figureCitation id="8D8048C43A4DFF8A97EEFDB1FD17C31A" box="[648,742,1521,1548]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="20" pageNumber="321">Fig. 6D</figureCitation>
|
||
).
|
||
<emphasis id="27CF88533A4DFF8A979BFDB0FC34C31D" bold="true" box="[765,965,1520,1547]" pageId="20" pageNumber="321">Endemism areas</emphasis>
|
||
:
|
||
<emphasis id="27CF88533A4DFF8A96B1FDB0FB3EC31D" bold="true" box="[983,1231,1520,1547]" pageId="20" pageNumber="321">Brazilian sub-region</emphasis>
|
||
: Boreal Brazilian dominion: Imeri province (see
|
||
<bibRefCitation id="712A29B03A4DFF8A9494FE55FD66C339" author="Morrone, J. J." box="[498,663,1557,1583]" pageId="20" pageNumber="321" pagination="1 - 110" refId="ref47051" refString="Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
|
||
; fig. 12).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |