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<document id="2CA58DB1298FB8C1CEA60B51DABF9006" ID-CLB-Dataset="21174" ID-DOI="10.11646/zootaxa.4629.3.1" ID-GBIF-Dataset="9419bc9b-dbe4-426c-a34e-f0034848ff99" ID-ISSN="1175-5326" ID-Zenodo-Dep="3995508" ID-ZooBank="1D47B452-0A40-4BC5-9BBA-C06DEB298C9A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598059654494" checkinUser="felipe" docAuthor="Silva, Fernando A. B. &amp; Valois, Marcely" docDate="2019" docId="9D12E5573A40FF8495F1FBC2FC16C49D" docLanguage="en" docName="zootaxa.4629.3.1.pdf" docOrigin="Zootaxa 4629 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scybalocanthon martinezi Silva &amp; Valois 2019, new species" docType="treatment" docVersion="8" lastPageNumber="327" masterDocId="612B9D2F3A59FF9E9566F840FFF1C516" masterDocTitle="A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)" masterLastPageNumber="341" masterPageNumber="301" pageNumber="326" updateTime="1698845243763" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="3E61D733473FB50E2ED54B4A9863A026">A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)</mods:title>
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<mods:name id="E5C19A1DACB25726B35F20BF96113D66" type="personal">
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<mods:namePart id="B38C3429B2906A5BA3604181EFC21C0E">Silva, Fernando A. B.</mods:namePart>
<mods:affiliation id="7DE681424D2284EF3C5087AD9D904AE9">Universidade Federal do Pará, Instituto de Ciências Biológicas, setor de Zoologia, Campus Belém. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil. E-mail: fernandoabsilva @ yahoo. com. br</mods:affiliation>
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<mods:namePart id="E5C230EEFE19039A85A2385E83116FEB">Valois, Marcely</mods:namePart>
<mods:affiliation id="A06279B08856C4650C09F138CAD2B740">Programa de Pós-graduação em Zoologia, Universidade Federal do Pará / Museu Paraense Emílio Goeldi. Rua Augusto Corrêa, 01. Guamá. Belém PA 66075 - 110, Brazil.</mods:affiliation>
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<mods:title id="BF1059DCB419892A2D4DBE7E1BC3159B">Zootaxa</mods:title>
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<mods:date id="5BE39E7B82FBAEBD66FC891C3E48D994">2019</mods:date>
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<mods:number id="96B645824815C590A24DBD24572BA2D6">2019-07-08</mods:number>
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<treatment id="9D12E5573A40FF8495F1FBC2FC16C49D" ID-DOI="http://doi.org/10.5281/zenodo.4335194" ID-GBIF-Taxon="167293234" ID-Zenodo-Dep="4335194" LSID="urn:lsid:plazi:treatment:9D12E5573A40FF8495F1FBC2FC16C49D" httpUri="http://treatment.plazi.org/id/9D12E5573A40FF8495F1FBC2FC16C49D" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
<subSubSection id="5DA107CA3A40FF8795F1FBC2FCD9C68A" box="[151,808,898,925]" pageId="25" pageNumber="326" type="nomenclature">
<paragraph id="150454413A40FF8795F1FBC2FCD9C68A" blockId="25.[151,808,898,961]" box="[151,808,898,925]" pageId="25" pageNumber="326">
<heading id="4E4CE32D3A40FF8795F1FBC2FCD9C68A" bold="true" box="[151,808,898,925]" fontSize="11" level="1" pageId="25" pageNumber="326" reason="1">
<emphasis id="27CF88533A40FF8795F1FBC2FCD9C68A" bold="true" box="[151,808,898,925]" pageId="25" pageNumber="326">
<taxonomicName id="D2BB2FC23A40FF8795F1FBC2FD7AC68A" ID-CoL="4W3LQ" authority="Silva &amp; Valois" authorityName="Silva &amp; Valois" authorityYear="2019" box="[151,651,898,925]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="martinezi" status="sp. nov.">
<emphasis id="27CF88533A40FF8795F1FBC2FE23C68A" bold="true" box="[151,466,898,925]" italics="true" pageId="25" pageNumber="326">Scybalocanthon martinezi</emphasis>
Silva &amp; Valois
</taxonomicName>
,
<taxonomicNameLabel id="3CFC35283A40FF8797FFFBC3FCD9C68A" box="[665,808,898,925]" pageId="25" pageNumber="326" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="5DA107CA3A40FF8795F1FBE6FE62C6D7" box="[151,403,933,961]" pageId="25" pageNumber="326" type="description">
<paragraph id="150454413A40FF8795F1FBE6FE62C6D7" blockId="25.[151,808,898,961]" box="[151,403,933,961]" pageId="25" pageNumber="326">
(
<figureCitation id="8D8048C43A40FF8795F9FBE6FF0DC6D6" box="[159,252,934,960]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="25" pageNumber="326">Figs. 1P</figureCitation>
,
<figureCitation id="8D8048C43A40FF879461FBE6FEDAC6D6" box="[263,299,934,960]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="25" pageNumber="326">4O</figureCitation>
,
<figureCitation id="8D8048C43A40FF879451FBE5FEADC6D6" box="[311,348,933,960]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="25" pageNumber="326">5N</figureCitation>
,
<figureCitation id="8D8048C43A40FF87940EFBE6FE7AC6D7" box="[360,395,934,961]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="25" pageNumber="326">6D</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A40FF8795F1FBA9FD94C131" pageId="25" pageNumber="326" type="etymology">
<paragraph id="150454413A40FF8795F1FBA9FD94C131" blockId="25.[151,1437,1001,2035]" pageId="25" pageNumber="326">
<emphasis id="27CF88533A40FF8795F1FBA9FEEFC115" bold="true" box="[151,286,1001,1027]" pageId="25" pageNumber="326">Etymology.</emphasis>
Named in honour of Antonio Martínez, for his description of
<taxonomicName id="D2BB2FC23A40FF8796A2FBA9FB86C115" authorityName="Martinez" authorityYear="1948" box="[964,1143,1001,1027]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="genus">
<emphasis id="27CF88533A40FF8796A2FBA9FB86C115" box="[964,1143,1001,1027]" italics="true" pageId="25" pageNumber="326">Scybalocanthon</emphasis>
</taxonomicName>
and his valuable contributions towards the taxonomy of the genus.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A40FF8795A1FC70FDD7C0C1" pageId="25" pageNumber="326" type="diagnosis">
<paragraph id="150454413A40FF8795A1FC70FDD7C0C1" blockId="25.[151,1437,1001,2035]" pageId="25" pageNumber="326">
<emphasis id="27CF88533A40FF8795A1FC70FECBC15D" bold="true" box="[199,314,1072,1099]" pageId="25" pageNumber="326">Diagnosis</emphasis>
. Specimens of
<taxonomicName id="D2BB2FC23A40FF87948BFC71FD84C15D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[493,629,1073,1099]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="martinezi">
<emphasis id="27CF88533A40FF87948BFC71FD84C15D" box="[493,629,1073,1099]" italics="true" pageId="25" pageNumber="326">S. martinezi</emphasis>
</taxonomicName>
are similar to those of
<taxonomicName id="D2BB2FC23A40FF8796E4FC71FC08C15D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[898,1017,1073,1099]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="27CF88533A40FF8796E4FC71FC08C15D" box="[898,1017,1073,1099]" italics="true" pageId="25" pageNumber="326">S. arnaudi</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A40FF87916FFC71FB12C15D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[1033,1251,1073,1099]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="federicoescobari">
<emphasis id="27CF88533A40FF87916FFC71FB12C15D" box="[1033,1251,1073,1099]" italics="true" pageId="25" pageNumber="326">S. federicoescobari</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A40FF879195FC71FA89C15D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[1267,1400,1073,1099]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="papaxibe">
<emphasis id="27CF88533A40FF879195FC71FA89C15D" box="[1267,1400,1073,1099]" italics="true" pageId="25" pageNumber="326">S. papaxibe</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A40FF8790E1FC71FEFEC179" baseAuthorityName="Kirsch" baseAuthorityYear="1873" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="pinopterus">
<emphasis id="27CF88533A40FF8790E1FC71FEFEC179" italics="true" pageId="25" pageNumber="326">S. pinopterus</emphasis>
</taxonomicName>
, and
<taxonomicName id="D2BB2FC23A40FF87942DFC15FE02C179" authorityName="Silva &amp; Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[331,499,1109,1135]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="uniplagiatus">
<emphasis id="27CF88533A40FF87942DFC15FE02C179" box="[331,499,1109,1135]" italics="true" pageId="25" pageNumber="326">S. uniplagiatus</emphasis>
</taxonomicName>
by having the pronotum uniformly colored, or with one elliptical spot on the central portion (
<figureCitation id="8D8048C43A40FF87959FFC39FEA6C185" box="[249,343,1145,1171]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="25" pageNumber="326">Figs. 1P</figureCitation>
,
<figureCitation id="8D8048C43A40FF879403FC39FE78C185" box="[357,393,1145,1171]" captionStart="FIGURE 2" captionStartId="9.[151,250,1178,1203]" captionTargetBox="[165,1424,188,1136]" captionTargetId="figure@9.[151,1436,181,1152]" captionTargetPageId="9" captionText="FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995514" httpUri="https://zenodo.org/record/3995514/files/figure.png" pageId="25" pageNumber="326">2A</figureCitation>
,
<figureCitation id="8D8048C43A40FF8794F1FC39FE41C185" box="[407,432,1145,1171]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="25" pageNumber="326">1I</figureCitation>
,
<figureCitation id="8D8048C43A40FF8794D8FC39FE13C185" box="[446,482,1145,1171]" captionStart="FIGURE 2" captionStartId="9.[151,250,1178,1203]" captionTargetBox="[165,1424,188,1136]" captionTargetId="figure@9.[151,1436,181,1152]" captionTargetPageId="9" captionText="FIGURE 2. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon pinopterus; B, S. pygidialis; C–G, S. sexpilotus; H–J, S. trimaculatus; K, S. uniplagiatus. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995514" httpUri="https://zenodo.org/record/3995514/files/figure.png" pageId="25" pageNumber="326">2K</figureCitation>
,
<figureCitation id="8D8048C43A40FF879496FC39FDB2C185" box="[496,579,1145,1171]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="25" pageNumber="326">1D, 1T</figureCitation>
); femora almost completely yellow or brown, with black spots only on the tips (
<figureCitation id="8D8048C43A40FF8795B6FCDDFEDAC1A1" box="[208,299,1181,1207]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="25" pageNumber="326">Fig. 3G</figureCitation>
); eighth elytral stria with thin carina at the anterior portion (
<figureCitation id="8D8048C43A40FF8796B3FCDDFBC1C1A1" box="[981,1072,1181,1207]" captionStart="FIGURE 3" captionStartId="12.[151,250,1773,1798]" captionTargetBox="[159,1427,925,1741]" captionTargetId="figure@12.[151,1436,921,1749]" captionTargetPageId="12" captionText="FIGURE 3. Details of the external morphology in Scybalocanthon. A, Elytral surface in S. chamorroi new species; B, elytral surface in S. acrianus new species; C, pygidium in S. sexpilotus; D, detail of the eighth elytral stria in S. pygidialis (carina point- ed); E, mesofemur in S. kaestneri; F, mesofemur in S. nigriceps; G, mesotarsus in S. moniliatus; H, metatarsus in S. moniliatus; I, mesotibia in S. aereus; J, mesotibia of Canthon arcabuquensis." figureDoi="http://doi.org/10.5281/zenodo.3995516" httpUri="https://zenodo.org/record/3995516/files/figure.png" pageId="25" pageNumber="326">Fig. 3D</figureCitation>
); endophallus with bristles right beside the FLP sclerite (
<figureCitation id="8D8048C43A40FF8794CDFC81FD34C1CD" box="[427,709,1216,1243]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="25" pageNumber="326">Figs. 5N, R, H, W, D, Q</figureCitation>
); and additional sclerite (
<collectingCountry id="6DAC14D13A40FF879680FC81FBFDC1CD" box="[998,1036,1217,1243]" name="American Samoa" pageId="25" pageNumber="326">AS</collectingCountry>
) absent (
<figureCitation id="8D8048C43A40FF879113FC81FA61C1CD" box="[1141,1424,1216,1243]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="25" pageNumber="326">Figs. 5N, R, H, W, D, Q</figureCitation>
). They can be distinguished from those of
<taxonomicName id="D2BB2FC23A40FF879704FCA5FD07C1E9" baseAuthorityName="Kirsch" baseAuthorityYear="1873" box="[610,758,1253,1279]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="pinopterus">
<emphasis id="27CF88533A40FF879704FCA5FD07C1E9" box="[610,758,1253,1279]" italics="true" pageId="25" pageNumber="326">S. pinopterus</emphasis>
</taxonomicName>
and
<taxonomicName id="D2BB2FC23A40FF87964AFCA5FC50C1E9" authorityName="Silva &amp; Valois" authorityYear="2019" box="[812,929,1253,1279]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="arnaudi">
<emphasis id="27CF88533A40FF87964AFCA5FC50C1E9" box="[812,929,1253,1279]" italics="true" pageId="25" pageNumber="326">S. arnaudi</emphasis>
</taxonomicName>
by the strongly asymmetrical parameres; left paramere with acute projection on dorsal margin and bilobate excavation on ventral margin (
<figureCitation id="8D8048C43A40FF8791FAFD49FB05C035" box="[1180,1268,1289,1315]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="25" pageNumber="326">Fig. 4O</figureCitation>
) (other species with slightly asymmetrical parameres, lacking acute projection and bilobate excavation on dorsal and ventral margins, respectively (
<figureCitation id="8D8048C43A40FF87940CFD11FE16C07D" box="[362,487,1361,1387]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="25" pageNumber="326">Figs. 4S, E</figureCitation>
)); from those of
<taxonomicName id="D2BB2FC23A40FF8797C5FD11FC8AC07D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[675,891,1361,1387]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="federicoescobari">
<emphasis id="27CF88533A40FF8797C5FD11FC8AC07D" box="[675,891,1361,1387]" italics="true" pageId="25" pageNumber="326">S. federicoescobari</emphasis>
</taxonomicName>
,
<taxonomicName id="D2BB2FC23A40FF8796EFFD11FBFDC07D" authorityName="Silva &amp; Valois" authorityYear="2019" box="[905,1036,1361,1387]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="papaxibe">
<emphasis id="27CF88533A40FF8796EFFD11FBFDC07D" box="[905,1036,1361,1387]" italics="true" pageId="25" pageNumber="326">S. papaxibe</emphasis>
</taxonomicName>
, and
<taxonomicName id="D2BB2FC23A40FF87912EFD11FB01C07D" authorityName="Silva &amp; Valois" authorityYear="2019" baseAuthorityName="Schmidt" baseAuthorityYear="1922" box="[1096,1264,1361,1387]" class="Insecta" family="Scarabaeidae" genus="Scybalocanthon" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="326" phylum="Arthropoda" rank="species" species="uniplagiatus">
<emphasis id="27CF88533A40FF87912EFD11FB01C07D" box="[1096,1264,1361,1387]" italics="true" pageId="25" pageNumber="326">S. uniplagiatus</emphasis>
</taxonomicName>
by the bilobate excavation of ventral margin of left paramere wide and deep, extending at least one-half of the paramere margin in the lateral view (
<figureCitation id="8D8048C43A40FF879434FDD9FE5BC0A5" box="[338,426,1433,1459]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="25" pageNumber="326">Fig. 4O</figureCitation>
) (other species with a short bilobate excavation, not reaching one-half of paramere margin in the lateral view (
<figureCitation id="8D8048C43A40FF879416FDFDFDFEC0C1" box="[368,527,1469,1495]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="25" pageNumber="326">Figs. 4I, W, R</figureCitation>
)).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A40FF8795A1FDA0FAC7C2BD" pageId="25" pageNumber="326" type="description">
<paragraph id="150454413A40FF8795A1FDA0FAC7C2BD" blockId="25.[151,1437,1001,2035]" pageId="25" pageNumber="326">
<emphasis id="27CF88533A40FF8795A1FDA0FE59C0ED" bold="true" box="[199,424,1504,1531]" pageId="25" pageNumber="326">Description. Body.</emphasis>
Oval-elongate. Surface opaque (except pronotum), completely microgranulate.
<emphasis id="27CF88533A40FF879037FDA0FA64C0ED" bold="true" box="[1361,1429,1504,1531]" pageId="25" pageNumber="326">Color</emphasis>
. Most of pronotum, hypomera (except internal margin), metaventrite, abdominal ventrites, pygidium, middle of femora, and tibiae yellow or light brown. Head, internal margin of hypomera, prosternum, mesoventrite, mesoepisternae, metaepisternae, elytra, trochanters, and proximal and distal portions of femora dark brown.
<emphasis id="27CF88533A40FF879075FE0DFA81C371" bold="true" box="[1299,1392,1613,1639]" pageId="25" pageNumber="326">Length.</emphasis>
<quantity id="D243F9A43A40FF87901FFE0CFF37C39D" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="25" pageNumber="326" unit="mm" value="8.5">8.5 mm</quantity>
.
<emphasis id="27CF88533A40FF8795B7FE31FEDBC39D" bold="true" box="[209,298,1649,1675]" pageId="25" pageNumber="326">Thorax</emphasis>
. Pronotum with one brown rounded spot at the anterocentral portion (
<figureCitation id="8D8048C43A40FF879125FE31FB69C39D" box="[1091,1176,1649,1675]" captionStart="FIGURE 1" captionStartId="8.[151,250,1804,1829]" captionTargetBox="[165,1428,599,1773]" captionTargetId="figure@8.[151,1436,594,1776]" captionTargetPageId="8" captionText="FIGURE 1. Species of Scybalocanthon, dorsal habitus. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. batesi; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N–O, S. maculatus; P, S. martinezi new species; Q–R; S. moniliatus; S. S. nigriceps; T. S. papaxibe new species. Scale bar: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3995510" httpUri="https://zenodo.org/record/3995510/files/figure.png" pageId="25" pageNumber="326">Fig. 1P</figureCitation>
). In some, spot absent. Anterior angles of pronotum approximately 80°. Lateral margin regularly curved outward, not forming an angle at the middle portion.
<emphasis id="27CF88533A40FF8794F1FEF9FE12C3C5" bold="true" box="[407,483,1721,1747]" pageId="25" pageNumber="326">Elytra</emphasis>
. Striae thin and shiny, punctures conspicuous. Eighth stria with a thin carina at the anterior portion.
<emphasis id="27CF88533A40FF87943EFE9CFE23C3E1" bold="true" box="[344,466,1756,1783]" pageId="25" pageNumber="326">Aedeagus.</emphasis>
Parameres strongly asymmetrical (
<figureCitation id="8D8048C43A40FF879603FE9DFC30C3E1" box="[869,961,1757,1783]" captionStart="FIGURE 4" captionStartId="14.[151,250,1663,1688]" captionTargetBox="[164,1424,190,1621]" captionTargetId="figure@14.[151,1436,181,1637]" captionTargetPageId="14" captionText="FIGURE 4. Aedeagus, detail of parameres in left and right view. A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. aereus (lectotype); E, S. arnaudi new species; F, S. chamorroi new species; G, S. cyanocephalus; H, S. darlingtoni; I, S. federicoescobari new species; J, S. haroldi new species; K, S. kaestneri; L, S. kelleri; M, S. korasakiae; N, S. maculatus; O, S. martinezi new species; P, S. moniliatus; Q, S. nigriceps; R, S. papaxibe new species; S, S. pinopterus; T, S. pygidialis; U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995518" httpUri="https://zenodo.org/record/3995518/files/figure.png" pageId="25" pageNumber="326">Fig. 4O</figureCitation>
). Dorsal margin of right paramere curved inward, apex obliquely truncate. Ventral margin of right paramere substraight, with a rounded excavation at the basal portion. Dorsal margin of left paramere curved inward from the basal to medial portions, apical portion with a short and pointed projection. Ventral margin of left paramere substraight, with a bilobate excavation from the basal to medial portions. SRP circular, with rounded handle-shaped extension (
<figureCitation id="8D8048C43A40FF8796ADFF2DFBD4C291" box="[971,1061,1900,1927]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="25" pageNumber="326">Fig. 5N</figureCitation>
). FLP short, comma shaped, with three sets of bristles right beside it (
<figureCitation id="8D8048C43A40FF879741FFD1FD70C2BD" box="[551,641,1936,1963]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="25" pageNumber="326">Fig. 5N</figureCitation>
). A+SA with two superposed and elongate sclerites (
<figureCitation id="8D8048C43A40FF8791A9FFD1FADBC2BD" box="[1231,1322,1936,1963]" captionStart="FIGURE 5" captionStartId="16.[151,250,1584,1609]" captionTargetBox="[170,1421,191,1543]" captionTargetId="figure@16.[151,1436,181,1558]" captionTargetPageId="16" captionText="FIGURE 5. Sclerites of endophallus. Superior right peripheral (SRP) sclerite. Fronto-lateral peripheral (FLP) sclerite. Complex of axial and subaxial (A+SA) sclerites.Additional sclerite (AS). Set of microbristles (Smb). Set of bristles (Sb). A, Scybalocanthon acrianus new species; B, S. adisi new species; C, S. aereus; D, S. arnaudi new species; E, S. chamorroi new species; F, S. cyanocephalus; G, S. darlingtoni; H, S. federicoescobari new species; I, S. haroldi new species; J. S. kaestneri; K, S. kelleri; L, S. korasakiae; M, S. maculatus; N, S. martinezi new species; O, S. moniliatus; P, S. nigriceps; Q, S. papaxibe new species; R, S. pinopterus; S, S. pygidialis (specimen from Amazonas, Brazil); T, S. pygidialis (specimen from Cayenne, French Guiana); U, S. sexspilotus; V, S. trimaculatus; W, S. uniplagiatus. Scale bar: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3995520" httpUri="https://zenodo.org/record/3995520/files/figure.png" pageId="25" pageNumber="326">Fig. 5N</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A40FF8495A1FFF5FAC5C455" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326" type="materials_examined">
<paragraph id="150454413A40FF8495A1FFF5FAC5C455" blockId="25.[151,1437,1001,2035]" lastBlockId="26.[151,1437,152,395]" lastPageId="26" lastPageNumber="327" pageId="25" pageNumber="326">
<emphasis id="27CF88533A40FF8795A1FFF5FE84C2D9" bold="true" box="[199,373,1972,1999]" pageId="25" pageNumber="326">Type material.</emphasis>
<materialsCitation id="A5D35E1C3A40FF87941AFFF5FF0AC2E5" ID-GBIF-Occurrence="2847261398" collectingDate="1989-09-20" collectionCode="CEMT" country="Ecuador" location="Bosque Sendero" municipality="Daimi" pageId="25" pageNumber="326" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="holotype">
<emphasis id="27CF88533A40FF87941AFFF5FD7FC2D9" bold="true" box="[380,654,1972,1999]" pageId="25" pageNumber="326">
<typeStatus id="CA00EAE33A40FF87941AFFF5FE1AC2D9" box="[380,491,1973,1999]" pageId="25" pageNumber="326" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="6DAC14D13A40FF879491FFF4FD78C2D9" box="[503,649,1972,1999]" name="Ecuador" pageId="25" pageNumber="326">ECUADOR</collectingCountry>
:
</emphasis>
<collectingRegion id="D77F9AA33A40FF8797F3FFF5FD17C2D9" box="[661,742,1973,1999]" country="Ecuador" name="Napo" pageId="25" pageNumber="326">NAPO</collectingRegion>
:
<collectingMunicipality id="F560CE3B3A40FF879796FFF5FCC7C2D9" box="[752,822,1973,1999]" pageId="25" pageNumber="326">Daimi</collectingMunicipality>
1,
<location id="1064029A3A40FF87963FFFF5FBE2C2D9" LSID="urn:lsid:plazi:treatment:9D12E5573A40FF8495F1FBC2FC16C49D:1064029A3A40FF87963FFFF5FBE2C2D9" box="[857,1043,1973,1999]" country="Ecuador" municipality="Daimi" name="Bosque Sendero" pageId="25" pageNumber="326" stateProvince="Napo">Bosque Sendero</location>
,
<date id="610572813A40FF879178FFF5FB69C2D9" box="[1054,1176,1973,1999]" pageId="25" pageNumber="326" value="1989-09-20">
<collectingDate id="71418B693A40FF879178FFF5FB69C2D9" box="[1054,1176,1973,1999]" pageId="25" pageNumber="326" value="1989-09-20">20.ix.1989</collectingDate>
</date>
,
<collectingMethod id="CCFA2C563A40FF8791C2FFF5FB1BC2D9" box="[1188,1258,1973,1999]" pageId="25" pageNumber="326">pitfall</collectingMethod>
, Sandoval—
<specimenCount id="03BD9FC83A40FF8790E7FFF5FA6DC2D9" box="[1409,1436,1973,1999]" pageId="25" pageNumber="326" type="male">1♂</specimenCount>
(
<collectionCode id="73AACC843A40FF8795F9FF99FF02C2E5" box="[159,243,2009,2035]" pageId="25" pageNumber="326">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A5D35E1C3A40FF849461FF99FE69C5A5" ID-GBIF-Occurrence="2847261568" collectingDate="1997-10-20" collectingDateMax="1997-10-21" collectingDateMin="1997-10-20" collectionCode="CEMT" collectorName="D. Padilla" country="Ecuador" lastPageId="26" lastPageNumber="327" location="Scyasuni" pageId="25" pageNumber="326" specimenCount="38" specimenCount-female="19" specimenCount-male="19" stateProvince="Napo" typeStatus="paratype">
<emphasis id="27CF88533A40FF879461FF99FD7EC2E5" bold="true" box="[263,655,2008,2035]" pageId="25" pageNumber="326">
<typeStatus id="CA00EAE33A40FF879461FF99FE8EC2E5" box="[263,383,2009,2035]" pageId="25" pageNumber="326" type="paratype">Paratypes</typeStatus>
[
<specimenCount id="03BD9FC83A40FF8794E8FF99FE0BC2E5" box="[398,506,2009,2035]" pageId="25" pageNumber="326" type="male">19 males</specimenCount>
,
<specimenCount id="03BD9FC83A40FF879761FF99FD79C2E5" box="[519,648,2008,2035]" pageId="25" pageNumber="326" type="female">18 females</specimenCount>
]
</emphasis>
.
<emphasis id="27CF88533A40FF8797F8FF98FCC4C2E5" bold="true" box="[670,821,2008,2035]" pageId="25" pageNumber="326">
<collectingCountry id="6DAC14D13A40FF8797F8FF98FCC1C2E5" box="[670,816,2008,2035]" name="Ecuador" pageId="25" pageNumber="326">ECUADOR</collectingCountry>
:
</emphasis>
<collectingRegion id="D77F9AA33A40FF87965BFF99FC7FC2E5" box="[829,910,2009,2035]" country="Ecuador" name="Napo" pageId="25" pageNumber="326">NAPO</collectingRegion>
:
<location id="1064029A3A40FF8796FFFF99FBF0C2E5" LSID="urn:lsid:plazi:treatment:9D12E5573A40FF8495F1FBC2FC16C49D:1064029A3A40FF8796FFFF99FBF0C2E5" box="[921,1025,2009,2035]" country="Ecuador" name="Scyasuni" pageId="25" pageNumber="326" stateProvince="Napo">Scyasuni</location>
,
<date id="610572813A40FF87916BFF99FB5BC2E5" box="[1037,1194,2009,2035]" pageId="25" pageNumber="326" value="1997-10-20" valueMax="1997-10-21" valueMin="1997-10-20">
<collectingDate id="71418B693A40FF87916BFF99FB5BC2E5" box="[1037,1194,2009,2035]" pageId="25" pageNumber="326" value="1997-10-20" valueMax="1997-10-21" valueMin="1997-10-20">
20–21.x.
<quantity id="D243F9A43A40FF879108FF99FB5BC2E5" box="[1134,1194,2009,2035]" metricMagnitude="6" metricUnit="m" metricValue="1.9972" pageId="25" pageNumber="326" unit="m" value="1997200.0">1997</quantity>
</collectingDate>
</date>
, 200 m,
<collectorName id="B84E31973A40FF87906BFF99FA8EC2E5" box="[1293,1407,2009,2035]" pageId="25" pageNumber="326">D. Padilla</collectorName>
&amp; I. Tapin—
<specimenCount id="03BD9FC83A43FF849470F8D9FEDCC5A5" box="[278,301,153,179]" pageId="26" pageNumber="327" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A43FF84945AF8D9FE61C5A5" box="[316,400,153,179]" pageId="26" pageNumber="327">CEMT</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="A5D35E1C3A43FF8494C5F8D8FE0CC5C1" ID-GBIF-Occurrence="2847261312" collectingDate="1970-07-10" collectingDateMax="1974-03-02" collectingDateMin="1970-07-10" collectionCode="CMNC, MZUFPA" collectorName="Peck &amp; Howden &amp; S. Peck" country="Colombia" elevation="215" latitude="-4.0691667" location="Leticia" longLatPrecision="21" longitude="-69.998886" pageId="26" pageNumber="327" specimenCount="36" specimenCount-female="17" specimenCount-male="19" typeStatus="paratype">
<collectingCountry id="6DAC14D13A43FF8494C5F8D8FDB3C5A5" box="[419,578,152,179]" name="Colombia" pageId="26" pageNumber="327">
<emphasis id="27CF88533A43FF8494C5F8D8FDB3C5A5" bold="true" box="[419,578,152,179]" pageId="26" pageNumber="327">COLOMBIA</emphasis>
</collectingCountry>
: AMAZONAS,
<location id="1064029A3A43FF84979AF8D9FCBCC5A5" LSID="urn:lsid:plazi:treatment:9D12E5573A40FF8495F1FBC2FC16C49D:1064029A3A43FF84979AF8D9FCBCC5A5" box="[764,845,153,179]" country="Colombia" latitude="-4.0691667" longLatPrecision="21" longitude="-69.998886" name="Leticia" pageId="26" pageNumber="327">Leticia</location>
,
<quantity id="D243F9A43A43FF84963FF8D8FC51C5A5" box="[857,928,152,179]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="327" unit="m" value="215.0">
<elevation id="9E96B3723A43FF84963FF8D8FC51C5A5" box="[857,928,152,179]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="327" unit="m" value="215.0">215 m</elevation>
</quantity>
(
<geoCoordinate id="708F32863A43FF8496D7F8D9FBDDC5A5" box="[945,1068,153,179]" degrees="4" direction="south" minutes="11" orientation="latitude" pageId="26" pageNumber="327" precision="15" seconds="20" value="-4.188889">4°11’20’’S</geoCoordinate>
,
<geoCoordinate id="708F32863A43FF84915FF8D8FB30C5A5" box="[1081,1217,152,179]" degrees="69" direction="west" minutes="56" orientation="longitude" pageId="26" pageNumber="327" precision="15" seconds="9" value="-69.93584">69°56’9’’W</geoCoordinate>
),
<date id="610572813A43FF8491B0F8D9FAA6C5A5" box="[1238,1367,153,179]" pageId="26" pageNumber="327" value="1970-07-10">
<collectingDate id="71418B693A43FF8491B0F8D9FAA6C5A5" box="[1238,1367,153,179]" pageId="26" pageNumber="327" value="1970-07-10">10.vii.1970</collectingDate>
</date>
, H. &amp; A. Howden—
<specimenCount id="03BD9FC83A43FF84945AF8FDFEA8C5C1" box="[316,345,189,215]" pageId="26" pageNumber="327" type="male">3♂</specimenCount>
<specimenCount id="03BD9FC83A43FF849405F8FDFE77C5C1" box="[355,390,189,215]" pageId="26" pageNumber="327" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A43FF8494FEF8FDFE00C5C1" box="[408,497,189,215]" pageId="26" pageNumber="327">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A43FF849761F8FDFEF5C5ED" ID-GBIF-Occurrence="2847261504" collectingDate="1970-07-10" collectingDateMax="1974-03-02" collectingDateMin="1970-07-10" collectionCode="CMNC, MZUFPA" collectorName="Peck &amp; Howden &amp; S. Peck" country="Colombia" elevation="215" latitude="-4.0691667" location="Leticia" longLatPrecision="21" longitude="-69.998886" pageId="26" pageNumber="327" specimenCount="36" specimenCount-female="17" specimenCount-male="19" typeStatus="paratype">
Leticia,
<quantity id="D243F9A43A43FF849703F8FCFD5FC5C1" box="[613,686,188,215]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="327" unit="m" value="215.0">
<elevation id="9E96B3723A43FF849703F8FCFD5FC5C1" box="[613,686,188,215]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="327" unit="m" value="215.0">215 m</elevation>
</quantity>
(
<geoCoordinate id="708F32863A43FF8497A7F8FDFCCDC5C1" box="[705,828,189,215]" degrees="4" direction="south" minutes="11" orientation="latitude" pageId="26" pageNumber="327" precision="15" seconds="20" value="-4.188889">4°11’20’’S</geoCoordinate>
,
<geoCoordinate id="708F32863A43FF84962DF8FCFC25C5C1" box="[843,980,188,215]" degrees="69" direction="west" minutes="56" orientation="longitude" pageId="26" pageNumber="327" precision="15" seconds="9" value="-69.93584">69°56’9’’W</geoCoordinate>
),
<date id="610572813A43FF84968DF8FCFB78C5C1" box="[1003,1161,188,215]" pageId="26" pageNumber="327" value="1972-02-19" valueMax="1972-02-25" valueMin="1972-02-19">
<collectingDate id="71418B693A43FF84968DF8FCFB78C5C1" box="[1003,1161,188,215]" pageId="26" pageNumber="327" value="1972-02-19" valueMax="1972-02-25" valueMin="1972-02-19">19–25.ii.1972</collectingDate>
</date>
, H. &amp; A. Howden—
<specimenCount id="03BD9FC83A43FF8490E0F8FDFA6CC5C1" box="[1414,1437,189,215]" pageId="26" pageNumber="327" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A43FF8495F9F8A1FF09C5ED" box="[159,248,225,251]" pageId="26" pageNumber="327">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A43FF84946CF8A1FEE6C409" ID-GBIF-Occurrence="2847261346" collectingDate="1970-07-10" collectingDateMax="1974-03-02" collectingDateMin="1970-07-10" collectionCode="CMNC, MZUFPA" collectorName="Peck &amp; Howden &amp; S. Peck" country="Colombia" elevation="215" latitude="-4.0691667" location="Leticia" longLatPrecision="21" longitude="-69.998886" pageId="26" pageNumber="327" specimenCount="36" specimenCount-female="17" specimenCount-male="19" typeStatus="paratype">
Leticia,
<quantity id="D243F9A43A43FF849403F8A0FE5AC5ED" box="[357,427,224,251]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="327" unit="m" value="215.0">
<elevation id="9E96B3723A43FF849403F8A0FE5AC5ED" box="[357,427,224,251]" metricMagnitude="2" metricUnit="m" metricValue="2.15" pageId="26" pageNumber="327" unit="m" value="215.0">215 m</elevation>
</quantity>
(
<geoCoordinate id="708F32863A43FF8494DCF8A1FDC4C5ED" box="[442,565,225,251]" degrees="4" direction="south" minutes="11" orientation="latitude" pageId="26" pageNumber="327" precision="15" seconds="20" value="-4.188889">4°11’20’’S</geoCoordinate>
,
<geoCoordinate id="708F32863A43FF849727F8A0FD38C5ED" box="[577,713,224,251]" degrees="69" direction="west" minutes="56" orientation="longitude" pageId="26" pageNumber="327" precision="15" seconds="9" value="-69.93584">69°56’9’’W</geoCoordinate>
),
<date id="610572813A43FF8497BAF8A0FC8BC5ED" box="[732,890,224,251]" pageId="26" pageNumber="327" value="1972-02-20" valueMax="1972-02-25" valueMin="1972-02-20">
<collectingDate id="71418B693A43FF8497BAF8A0FC8BC5ED" box="[732,890,224,251]" pageId="26" pageNumber="327" value="1972-02-20" valueMax="1972-02-25" valueMin="1972-02-20">20–25.ii.1972</collectingDate>
</date>
,
<collectorName id="B84E31973A43FF8496E3F8A1FC4AC5ED" box="[901,955,225,251]" pageId="26" pageNumber="327">Peck</collectorName>
&amp;
<collectorName id="B84E31973A43FF8496BBF8A1FBCEC5ED" box="[989,1087,225,251]" pageId="26" pageNumber="327">Howden</collectorName>
, dung traps—
<specimenCount id="03BD9FC83A43FF8491B8F8A1FAFAC5ED" box="[1246,1291,225,251]" pageId="26" pageNumber="327" type="male">10♂</specimenCount>
<specimenCount id="03BD9FC83A43FF849076F8A1FAC2C5ED" box="[1296,1331,225,251]" pageId="26" pageNumber="327" type="female">4♀</specimenCount>
(
<collectionCode id="73AACC843A43FF849027F8A1FA69C5ED" box="[1345,1432,225,251]" pageId="26" pageNumber="327">CMNC</collectionCode>
,
<collectionCode id="73AACC843A43FF8495F1F945FEFDC409" box="[151,268,261,287]" pageId="26" pageNumber="327">MZUFPA</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A43FF84947AF945FA6DC409" ID-GBIF-Occurrence="2847261410" box="[284,1436,260,287]" collectingDate="1970-07-10" collectingDateMax="1974-03-02" collectingDateMin="1970-07-10" collectionCode="CMNC, MZUFPA" collectorName="Peck &amp; Howden &amp; S. Peck" country="Colombia" elevation="215" latitude="-4.0691667" location="Leticia" longLatPrecision="21" longitude="-69.998886" pageId="26" pageNumber="327" specimenCount="36" specimenCount-female="17" specimenCount-male="19" typeStatus="paratype">
Los Alpes, Leticia,
<quantity id="D243F9A43A43FF849494F945FDC6C409" box="[498,567,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="26" pageNumber="327" unit="m" value="213.0">
<elevation id="9E96B3723A43FF849494F945FDC6C409" box="[498,567,261,287]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="26" pageNumber="327" unit="m" value="213.0">213 m</elevation>
</quantity>
(
<geoCoordinate id="708F32863A43FF849723F944FD42C409" box="[581,691,260,287]" degrees="4" direction="south" minutes="4" orientation="latitude" pageId="26" pageNumber="327" precision="15" seconds="25" value="-4.0736113">4°4’25’’S</geoCoordinate>
,
<geoCoordinate id="708F32863A43FF8497D8F944FCC9C409" box="[702,824,260,287]" degrees="70" direction="west" minutes="0" orientation="longitude" pageId="26" pageNumber="327" precision="15" seconds="5" value="-70.00139">70°0’5’’W</geoCoordinate>
), 23.ii–iii.1974,
<collectorName id="B84E31973A43FF849688F945FBB2C409" box="[1006,1091,261,287]" pageId="26" pageNumber="327">S. Peck</collectorName>
, dung trap—
<specimenCount id="03BD9FC83A43FF8491B3F945FB05C409" box="[1237,1268,261,287]" pageId="26" pageNumber="327" type="male">2♂</specimenCount>
<specimenCount id="03BD9FC83A43FF84919FF945FADBC409" box="[1273,1322,261,287]" pageId="26" pageNumber="327" type="female">10♀</specimenCount>
(
<collectionCode id="73AACC843A43FF849051F945FA61C409" box="[1335,1424,261,287]" pageId="26" pageNumber="327">CMNC</collectionCode>
);
</materialsCitation>
<materialsCitation id="A5D35E1C3A43FF8495F1F969FAC1C455" ID-GBIF-Occurrence="2847261488" box="[151,1328,296,323]" collectingDate="1970-07-10" collectingDateMax="1974-03-02" collectingDateMin="1970-07-10" collectionCode="CMNC, MZUFPA" collectorName="Peck &amp; Howden &amp; S. Peck" country="Colombia" elevation="215" latitude="-4.0691667" location="Leticia" longLatPrecision="21" longitude="-69.998886" pageId="26" pageNumber="327" specimenCount="36" specimenCount-female="17" specimenCount-male="19" typeStatus="paratype">
road end, Leticia,
<quantity id="D243F9A43A43FF849404F969FE58C455" box="[354,425,297,323]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="26" pageNumber="327" unit="m" value="213.0">
<elevation id="9E96B3723A43FF849404F969FE58C455" box="[354,425,297,323]" metricMagnitude="2" metricUnit="m" metricValue="2.13" pageId="26" pageNumber="327" unit="m" value="213.0">213 m</elevation>
</quantity>
(
<geoCoordinate id="708F32863A43FF8494DCF969FDEBC455" box="[442,538,297,323]" degrees="4" direction="south" minutes="4" orientation="latitude" pageId="26" pageNumber="327" precision="15" seconds="9" value="-4.0691667">4°4’9’’S</geoCoordinate>
,
<geoCoordinate id="708F32863A43FF849740F968FD4DC455" box="[550,700,296,323]" degrees="69" direction="west" minutes="59" orientation="longitude" pageId="26" pageNumber="327" precision="15" seconds="56" value="-69.998886">69°59’56’’W</geoCoordinate>
),
<date id="610572813A43FF8497B7F969FC8EC455" box="[721,895,297,323]" pageId="26" pageNumber="327" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">
<collectingDate id="71418B693A43FF8497B7F969FC8EC455" box="[721,895,297,323]" pageId="26" pageNumber="327" value="1974-02-23" valueMax="1974-03-02" valueMin="1974-02-23">23.ii–2.iii.1974</collectingDate>
</date>
,
<collectorName id="B84E31973A43FF8496EDF969FC10C455" box="[907,993,297,323]" pageId="26" pageNumber="327">S. Peck</collectorName>
, dung trap—
<specimenCount id="03BD9FC83A43FF849110F969FB67C455" box="[1142,1174,297,323]" pageId="26" pageNumber="327" type="male">4♂</specimenCount>
<specimenCount id="03BD9FC83A43FF8491FBF969FB31C455" box="[1181,1216,297,323]" pageId="26" pageNumber="327" type="female">1♀</specimenCount>
(
<collectionCode id="73AACC843A43FF8491A9F969FAD9C455" box="[1231,1320,297,323]" pageId="26" pageNumber="327">CMNC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="5DA107CA3A43FF8495A1F90CFC16C49D" pageId="26" pageNumber="327" type="distribution">
<paragraph id="150454413A43FF8495A1F90CFC16C49D" blockId="26.[151,1437,152,395]" pageId="26" pageNumber="327">
<emphasis id="27CF88533A43FF8495A1F90CFE90C471" bold="true" box="[199,353,332,359]" pageId="26" pageNumber="327">Distribution.</emphasis>
Known from
<collectingCountry id="6DAC14D13A43FF84949BF90DFDABC471" box="[509,602,333,359]" name="Ecuador" pageId="26" pageNumber="327">Ecuador</collectingCountry>
and
<collectingCountry id="6DAC14D13A43FF8497F6F90DFD0FC471" box="[656,766,333,359]" name="Colombia" pageId="26" pageNumber="327">Colombia</collectingCountry>
(
<figureCitation id="8D8048C43A43FF849668F90DFC99C47E" box="[782,872,333,360]" captionStart="FIGURE 6" captionStartId="17.[151,250,1784,1809]" captionTargetBox="[179,1376,186,1734]" captionTargetId="figure@17.[174,1414,181,1759]" captionTargetPageId="17" captionText="FIGURE 6. Known distribution of species of Scybalocanthon." figureDoi="http://doi.org/10.5281/zenodo.3995522" httpUri="https://zenodo.org/record/3995522/files/figure.png" pageId="26" pageNumber="327">Fig. 6D</figureCitation>
).
<emphasis id="27CF88533A43FF84961DF90CFBB1C471" bold="true" box="[891,1088,332,359]" pageId="26" pageNumber="327">Endemism areas</emphasis>
:
<emphasis id="27CF88533A43FF849129F90CFAB2C471" bold="true" box="[1103,1347,332,359]" pageId="26" pageNumber="327">Brazilian sub-region</emphasis>
: Boreal Brazilian dominion: Napo and Imerí provinces (see
<bibRefCitation id="712A29B03A43FF8497B8F931FC72C49D" author="Morrone, J. J." box="[734,899,369,395]" pageId="26" pageNumber="327" pagination="1 - 110" refId="ref47051" refString="Morrone, J. J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1 - 110. https: // doi. org / 10.11646 / zootaxa. 3782.1.1" type="journal article" year="2014">Morrone 2014</bibRefCitation>
; fig. 12).
</paragraph>
</subSubSection>
</treatment>
</document>
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