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<document id="C5E70701A5AF256CE5A7E55201790FA6" ID="10.11646/zootaxa.4200.1.1" ID-DOI="10.11646/zootaxa.4200.1.1" ID-GBIF-Dataset="834295d1-63b4-4951-a5ea-f2d3ec0228f3" ID-ISSN="1175-5326" ID-Zenodo-Dep="177598" ID-ZooBank="911ECACD-8903-405E-AAA4-8300901D69C7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480318716440" checkinUser="plazi" docAuthor="Silva, Gabriel S. C., Melo, Bruno F., Oliveira, Claudio &amp; Benine, Ricardo C." docDate="2016" docId="03F47776FFEC5977FF5CBA8535FA4159" docLanguage="en" docName="zootaxa.4200.1.1.pdf" docOrigin="Zootaxa 4200 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetragonopterus ommatus Silva, Melo, Oliveira &amp; Benine, 2016, new species" docType="treatment" docVersion="12" lastPageNumber="40" masterDocId="FFCD0F0EFFC95950FFCBB83134144519" masterDocTitle="Revision of the South American genus Tetragonopterus Cuvier, 1816 (Teleostei: Characidae) with description of four new species" masterLastPageNumber="46" masterPageNumber="1" pageNumber="38" updateTime="1698394834132" updateUser="plazi">
<mods:mods id="299B37652744F72BF48D5257252EED22" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9869981CD1ED2B4CE0075D2B28F6648A">
<mods:title id="F92B5AD8CE1C752730095B5561BD60B1">Revision of the South American genus Tetragonopterus Cuvier, 1816 (Teleostei: Characidae) with description of four new species</mods:title>
</mods:titleInfo>
<mods:name id="FB0D79FE0269816D01A6C6BD2AEE57A1" type="personal">
<mods:role id="5FDFBA3E3B4523A34C2F1ACA4AF8A700">
<mods:roleTerm id="2A8D62A03BE13528C85A99A36F57ED84">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="15ACD8857DD13D7CE2F34C391467F36D">Silva, Gabriel S. C.</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="A74D8DE900FCD39BDD1419A1D7BA21D4">Melo, Bruno F.</mods:namePart>
</mods:name>
<mods:name id="7926CF7E6C7C13D45145641DEB3AE917" type="personal">
<mods:role id="BD59D3112F2FBA5E93B1DCD1B2110E14">
<mods:roleTerm id="81F31F8F9BFFD891B69962B26BA44ED0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="36E1AFA92D19BF37F2FAD5774ED79AA5">Oliveira, Claudio</mods:namePart>
</mods:name>
<mods:name id="72B373753DB76E93A9AEA340A700619C" type="personal">
<mods:role id="50029641A0C5CD06C4E1C06FC8FB7158">
<mods:roleTerm id="992784AA61BB08D01135570978434A38">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="355443859C24D7A4F72D1329A73EC98A">Benine, Ricardo C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="EF2A84389DAA1D25B6C05262F926FFFA">text</mods:typeOfResource>
<mods:relatedItem id="BA6A078575A2DF7EE824A4190F350D98" type="host">
<mods:titleInfo id="13594BE8AC75579E8E37F92C7602E716">
<mods:title id="EFE8390A1C5F58DB780E9978FB3CFA2E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="FFD175F1CC5741F208A176104F21C13D">
<mods:date id="A6255BE33D2827C0BC88B73A18667882">2016</mods:date>
<mods:detail id="2C02042ED004F014EF47757A0C7D51A7" type="volume">
<mods:number id="296FC343CD1DA17C72AD654F3152B469">4200</mods:number>
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<mods:classification id="90D660CD9BAEEA3D0330EB91AB3138AC">journal article</mods:classification>
<mods:identifier id="B16E9AF82E786D6710E4DEBF723B9F00" type="DOI">10.11646/zootaxa.4200.1.1</mods:identifier>
<mods:identifier id="8AC594A83A85C3C4D6F517EB824CC991" type="GBIF-Dataset">834295d1-63b4-4951-a5ea-f2d3ec0228f3</mods:identifier>
<mods:identifier id="9B3CE34AAA79ABC3FE6F811E607C9C88" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F3175CC4E94B749D2DEA039962B53ECF" type="Zenodo-Dep">177598</mods:identifier>
<mods:identifier id="BEDC2E2CCA2601CAE138B3DA385B2D19" type="ZooBank">911ECACD-8903-405E-AAA4-8300901D69C7</mods:identifier>
</mods:mods>
<treatment id="03F47776FFEC5977FF5CBA8535FA4159" ID-DOI="http://doi.org/10.5281/zenodo.4566273" ID-GBIF-Taxon="124864092" ID-Zenodo-Dep="4566273" LSID="urn:lsid:plazi:treatment:03F47776FFEC5977FF5CBA8535FA4159" httpUri="http://treatment.plazi.org/id/03F47776FFEC5977FF5CBA8535FA4159" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
<subSubSection id="C34795EBFFEC5975FF5CBA85355347F6" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BE2C660FFEC5975FF5CBA85366547D7" blockId="37.[151,625,692,751]" box="[151,625,692,718]" pageId="37" pageNumber="38">
<heading id="D0AA710CFFEC5975FF5CBA85366547D7" bold="true" box="[151,625,692,718]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
<emphasis id="B9291A72FFEC5975FF5CBA85366547D7" bold="true" box="[151,625,692,718]" pageId="37" pageNumber="38">
<taxonomicName id="4C5DBDE3FFEC5975FF5CBA8535C747D7" box="[151,467,692,718]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus" status="sp. nov.">
<emphasis id="B9291A72FFEC5975FF5CBA8535C747D7" bold="true" box="[151,467,692,718]" italics="true" pageId="37" pageNumber="38">Tetragonopterus ommatus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21AA709FFEC5975FE2ABA85366547D7" box="[481,625,692,718]" pageId="37" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE2C660FFEC5975FF5CBAE7355347F6" blockId="37.[151,625,692,751]" box="[151,327,726,751]" pageId="37" pageNumber="38">
<figureCitation id="1366DAE5FFEC5975FF5CBAE734FC47F6" box="[151,232,726,751]" captionStart="FIGURE 14" captionStartId="39.[151,250,791,813]" captionTargetBox="[151,1436,193,770]" captionTargetId="figure@39.[151,1436,193,770]" captionTargetPageId="39" captionText="FIGURE 14. Tetragonopterus ommatus. MZUSP 92667, holotype, 61.1 mm SL, Brazil, Pará, Itaituba, Rio Tapajós, Amazon basin." httpUri="https://zenodo.org/record/177611/files/figure.png" pageId="37" pageNumber="38">Fig. 14</figureCitation>
,
<tableCitation id="C6DFF3DBFFEC5975FF3FBAE7355347F6" box="[244,327,726,751]" captionStart="TABLE 8" captionStartId="38.[151,239,580,603]" captionText="TABLE 8. Morphometric data for Tetragonopterus ommatus (n = 5). Range includes holotype and paratypes." pageId="37" pageNumber="38">Table 8</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFEC5975FF5CBB1137DF462D" box="[151,971,798,821]" pageId="37" pageNumber="38" type="description">
<paragraph id="8BE2C660FFEC5975FF5CBB1137DF462D" blockId="37.[151,971,798,821]" box="[151,971,798,821]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="AB4DE14EFFEC5975FF5CBB1137DF462D" box="[151,971,798,821]" pageId="37" pageNumber="38">
<taxonomicName id="4C5DBDE3FFEC5975FF5CBB113529462C" box="[151,317,800,821]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="genus">
<emphasis id="B9291A72FFEC5975FF5CBB113529462C" box="[151,317,800,821]" italics="true" pageId="37" pageNumber="38">Tetragonopterus</emphasis>
</taxonomicName>
sp. Tapajós:
<treatmentCitation id="0AFCE071FFEC5975FE08BB2F36B7462D" author="Melo" box="[451,675,798,821]" page="709" pageId="37" pageNumber="38" year="2016">
<bibRefCitation id="EFCCBB91FFEC5975FE08BB2F3660462D" author="Melo" box="[451,628,798,821]" pageId="37" pageNumber="38" refString="Melo, B. F., Benine, R. C., Silva, G. S. C., Avelino, G. S. &amp; Oliveira, C. (2016) Molecular phylogeny of the Neotropical fish genus Tetragonopterus (Teleostei: Characiformes: Characidae). Molecular Phylogenetics and Evolution, 94, 709 - 717. http: // dx. doi. org / 10.1016 / j. ympev. 2015.10.022" type="journal article" year="2016">
Melo
<emphasis id="B9291A72FFEC5975FE35BB113625462D" box="[510,561,798,821]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
, 2016
</bibRefCitation>
: 709
</treatmentCitation>
–717 (molecular phylogeny).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFEC5975FF5CBB5036824113" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="8BE2C660FFEC5975FF5CBB5035BE4687" blockId="37.[151,1437,865,2006]" pageId="37" pageNumber="38">
<materialsCitation id="3B35CC3DFFEC5975FF5CBB5035BE4687" ID-GBIF-Occurrence="1412767517" collectingDate="2006-11-08" collectionCode="MZUSP" collectorName="J. Birindelli" country="Brazil" latitude="-4.359444" location="Rio Tapajos" longLatPrecision="21" longitude="-56.1675" municipality="Itaituba" pageId="37" pageNumber="38" specimenCode="MZUSP 92667, 64.1" specimenCount="1" stateProvince="Para" typeStatus="holotype">
<emphasis id="B9291A72FFEC5975FF5CBB50351F4663" bold="true" box="[151,267,865,890]" pageId="37" pageNumber="38">
<typeStatus id="54E678C2FFEC5975FF5CBB5035124663" box="[151,262,865,890]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBFB6E1BFFEC5975FED9BB5335EF4663" box="[274,507,865,890]" pageId="37" pageNumber="38">
<collectionCode id="ED4C5EA5FFEC5975FED9BB53357B4663" box="[274,367,866,890]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="37" pageNumber="38">MZUSP</collectionCode>
92667,
<quantity id="4CA56B85FFEC5975FE01BB5035EF4663" box="[458,507,865,890]" metricMagnitude="-2" metricUnit="m" metricValue="6.409999999999999" pageId="37" pageNumber="38" unit="mm" value="64.1">64.1</quantity>
</specimenCode>
mm SL,
<collectingCountry id="F34A86F0FFEC5975FDA9BB5036BD4663" box="[610,681,865,890]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
,
<collectingRegion id="49990882FFEC5975FD7EBB5336FE4663" box="[693,746,866,890]" country="Brazil" name="Para" pageId="37" pageNumber="38">Pará</collectingRegion>
,
<collectingMunicipality id="6B865C1AFFEC5975FD3FBB5037584663" box="[756,844,865,890]" pageId="37" pageNumber="38">Itaituba</collectingMunicipality>
,
<location id="8E8290BBFFEC5975FC9CBB5037F54663" LSID="urn:lsid:plazi:treatment:03F47776FFEC5977FF5CBA8535FA4159:8E8290BBFFEC5975FC9CBB5037F54663" box="[855,993,865,890]" country="Brazil" latitude="-4.359444" longLatPrecision="21" longitude="-56.1675" municipality="Itaituba" name="Rio Tapajos" pageId="37" pageNumber="38" stateProvince="Para">Rio Tapajós</location>
,
<geoCoordinate id="EE69A0A7FFEC5975FC20BB5330754663" box="[1003,1121,866,890]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="15" value="-4.359444">4°21’34”S</geoCoordinate>
<geoCoordinate id="EE69A0A7FFEC5975FBA2BB5030EC4663" box="[1129,1272,865,890]" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="15" value="-56.1675">56°10’03”W</geoCoordinate>
, L. Souza &amp;
<collectorName id="26A8A3B6FFEC5975FA41BB5335114687" pageId="37" pageNumber="38">J. Birindelli</collectorName>
, 0
<date id="FFE3E0A0FFEC5975FED4BBB735B24687" box="[287,422,901,926]" pageId="37" pageNumber="38" value="2006-11-08">
<collectingDate id="EFA71948FFEC5975FED4BBB735B24687" box="[287,422,901,926]" pageId="37" pageNumber="38" value="2006-11-08">8 Nov 2006</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE2C660FFEC5975FF0CBB9836824113" blockId="37.[151,1437,865,2006]" pageId="37" pageNumber="38">
<materialsCitation id="3B35CC3DFFEC5975FF0CBB98369046FF" ID-GBIF-Occurrence="1412767506" collectingDate="2011-09-25" country="Brazil" county="All" latitude="-4.4581943" location="Rio Tapajos" longLatPrecision="15" longitude="-56.267498" municipality="Itaituba" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<emphasis id="B9291A72FFEC5975FF0CBB98355246DB" bold="true" box="[199,326,937,962]" pageId="37" pageNumber="38">
<typeStatus id="54E678C2FFEC5975FF0CBB98355546DB" box="[199,321,937,962]" pageId="37" pageNumber="38" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectingCounty id="6283BEECFFEC5975FE99BB98356246DB" box="[338,374,937,962]" pageId="37" pageNumber="38">All</collectingCounty>
from
<collectingCountry id="F34A86F0FFEC5975FE0EBB98361A46DB" box="[453,526,937,962]" name="Brazil" pageId="37" pageNumber="38">Brazil</collectingCountry>
: LBP 13949, 3,
<quantity id="4CA56B85FFEC5975FD12BB98379646D8" box="[729,898,937,962]" metricMagnitude="-2" metricUnit="m" metricValue="6.435" metricValueMax="6.59" metricValueMin="6.279999999999999" pageId="37" pageNumber="38" unit="mm" value="64.35" valueMax="65.9" valueMin="62.8">62.8–65.9 mm</quantity>
SL,
<collectingRegion id="49990882FFEC5975FC09BB9B37E346DB" box="[962,1015,938,962]" country="Brazil" name="Para" pageId="37" pageNumber="38">Pará</collectingRegion>
,
<collectingMunicipality id="6B865C1AFFEC5975FBC3BB98307446DB" box="[1032,1120,937,962]" pageId="37" pageNumber="38">Itaituba</collectingMunicipality>
,
<location id="8E8290BBFFEC5975FBBABB98311446DB" LSID="urn:lsid:plazi:treatment:03F47776FFEC5977FF5CBA8535FA4159:8E8290BBFFEC5975FBBABB98311446DB" box="[1137,1280,937,962]" country="Brazil" county="All" latitude="-4.4581943" longLatPrecision="15" longitude="-56.267498" municipality="Itaituba" name="Rio Tapajos" pageId="37" pageNumber="38" stateProvince="Para">Rio Tapajós</location>
,
<geoCoordinate id="EE69A0A7FFEC5975FADBBB98318846DB" box="[1296,1436,937,962]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="1" value="-4.4581943">4°27’29.5”S</geoCoordinate>
<geoCoordinate id="EE69A0A7FFEC5975FF5CBBFC353246FF" box="[151,294,973,998]" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="15" value="-56.267498">56°16’03”W</geoCoordinate>
, R.
<location id="8E8290BBFFEC5975FE9FBBFC35B246FF" LSID="urn:lsid:plazi:treatment:03F47776FFEC5977FF5CBA8535FA4159:8E8290BBFFEC5975FE9FBBFC35B246FF" box="[340,422,973,998]" country="Brazil" county="All" latitude="-4.4581943" longLatPrecision="15" longitude="-56.267498" municipality="Itaituba" name="Britzke" pageId="37" pageNumber="38" stateProvince="Para">Britzke</location>
<emphasis id="B9291A72FFEC5975FE66BBFF35F246FF" box="[429,486,973,998]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
,
<collectingDate id="EFA71948FFEC5975FE3EBBFC369046FF" box="[501,644,973,998]" pageId="37" pageNumber="38" value="2011-09-25">
<date id="FFE3E0A0FFEC5975FE3EBBFC369046FF" box="[501,644,973,998]" pageId="37" pageNumber="38" value="2011-09-25">25 Sep 2011</date>
</collectingDate>
</materialsCitation>
.
<materialsCitation id="3B35CC3DFFEC5975FD44BBFF36864113" ID-GBIF-Occurrence="1412767489" collectingDate="2012-06-11" country="Brazil" county="Itaituba" latitude="-4.552695" location="Britzke" longLatPrecision="1" longitude="-56.29989" municipality="Rio Tapajos" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Para" typeStatus="paratype">
LBP 16126, 1,
<quantity id="4CA56B85FFEC5975FCF7BBFC37B446FC" box="[828,928,973,998]" metricMagnitude="-2" metricUnit="m" metricValue="4.76" pageId="37" pageNumber="38" unit="mm" value="47.6">47.6 mm</quantity>
SL,
<collectingRegion id="49990882FFEC5975FC1CBBFF301846FF" box="[983,1036,974,998]" country="Brazil" name="Para" pageId="37" pageNumber="38">Pará</collectingRegion>
,
<collectingCounty id="6283BEECFFEC5975FBDCBBFC307B46FF" box="[1047,1135,973,998]" pageId="37" pageNumber="38">Itaituba</collectingCounty>
,
<collectingMunicipality id="6B865C1AFFEC5975FBB0BBFC311046FF" box="[1147,1284,973,998]" pageId="37" pageNumber="38">Rio Tapajós</collectingMunicipality>
,
<geoCoordinate id="EE69A0A7FFEC5975FADABBFF318846FF" box="[1297,1436,974,998]" direction="south" orientation="latitude" pageId="37" pageNumber="38" precision="1" value="-4.552695">4°33’09.7”S</geoCoordinate>
<geoCoordinate id="EE69A0A7FFEC5975FF5CBBC035284113" box="[151,316,1009,1034]" direction="west" orientation="longitude" pageId="37" pageNumber="38" precision="1" value="-56.29989">56°17’59.6”W</geoCoordinate>
, R.
<location id="8E8290BBFFEC5975FEACBBC035AD4113" LSID="urn:lsid:plazi:treatment:03F47776FFEC5977FF5CBA8535FA4159:8E8290BBFFEC5975FEACBBC035AD4113" box="[359,441,1009,1034]" country="Brazil" county="Itaituba" latitude="-4.552695" longLatPrecision="1" longitude="-56.29989" municipality="Rio Tapajos" name="Britzke" pageId="37" pageNumber="38" stateProvince="Para">Britzke</location>
<emphasis id="B9291A72FFEC5975FE0ABBC335ED4113" box="[449,505,1009,1034]" italics="true" pageId="37" pageNumber="38">et al.</emphasis>
,
<collectingDate id="EFA71948FFEC5975FDCCBBC336864113" box="[519,658,1010,1034]" pageId="37" pageNumber="38" value="2012-06-11">
<date id="FFE3E0A0FFEC5975FDCCBBC336864113" box="[519,658,1010,1034]" pageId="37" pageNumber="38" value="2012-06-11">11 Jun 2012</date>
</collectingDate>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFEC5975FF0CBC24369140C7" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="8BE2C660FFEC5975FF0CBC24369140C7" blockId="37.[151,1437,865,2006]" pageId="37" pageNumber="38">
<emphasis id="B9291A72FFEC5975FF0CBC2435554134" bold="true" box="[199,321,1045,1070]" pageId="37" pageNumber="38">Diagnosis.</emphasis>
<taxonomicName id="4C5DBDE3FFEC5975FE80BC2736664134" box="[331,626,1046,1069]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEC5975FE80BC2736664134" box="[331,626,1046,1069]" italics="true" pageId="37" pageNumber="38">Tetragonopterus ommatus</emphasis>
</taxonomicName>
is distinguished from all congeners by the presence of 7–8 maxillary teeth (
<emphasis id="B9291A72FFEC5975FF15BC0B34E84148" box="[222,252,1082,1105]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
1–6) and by having a dark, vertically-oriented patch of dark pigmentation limited to the posterior portion of the caudal peduncle (
<emphasis id="B9291A72FFEC5975FE63BC6F35D3416C" box="[424,455,1118,1141]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
mark centered on the caudal peduncle).
<taxonomicName id="4C5DBDE3FFEC5975FC5EBC6F30AE416C" box="[917,1210,1118,1141]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEC5975FC5EBC6F30AE416C" box="[917,1210,1118,1141]" italics="true" pageId="37" pageNumber="38">Tetragonopterus ommatus</emphasis>
</taxonomicName>
also differs from
<taxonomicName id="4C5DBDE3FFEC5975FA4CBC6F351B4180" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="anostomus">
<emphasis id="B9291A72FFEC5975FA4CBC6F351B4180" italics="true" pageId="37" pageNumber="38">T. anostomus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5DBDE3FFEC5975FED4BCB335D24183" box="[287,454,1153,1178]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="denticulatus">
<emphasis id="B9291A72FFEC5975FED4BCB335D24183" box="[287,454,1153,1178]" italics="true" pageId="37" pageNumber="38">T. denticulatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5DBDE3FFEC5975FDC9BCB3366C4183" box="[514,632,1153,1178]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="kuluene">
<emphasis id="B9291A72FFEC5975FDC9BCB3366C4183" box="[514,632,1153,1178]" italics="true" pageId="37" pageNumber="38">T. kuluene</emphasis>
</taxonomicName>
by having 4 principal teeth on dentary (
<emphasis id="B9291A72FFEC5975FB84BCB3307A4180" box="[1103,1134,1154,1177]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
5–6) and by having more robust dentary teeth (
<emphasis id="B9291A72FFEC5975FE4DBC9735B041A4" box="[390,420,1190,1213]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
thinner and sharper teeth).
<taxonomicName id="4C5DBDE3FFEC5975FD12BC9737E941A4" box="[729,1021,1190,1213]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEC5975FD12BC9737E941A4" box="[729,1021,1190,1213]" italics="true" pageId="37" pageNumber="38">Tetragonopterus ommatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5DBDE3FFEC5975FB59BC97313141A4" box="[1170,1317,1190,1213]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="anostomus">
<emphasis id="B9291A72FFEC5975FB59BC97313141A4" box="[1170,1317,1190,1213]" italics="true" pageId="37" pageNumber="38">T. anostomus</emphasis>
</taxonomicName>
by having a terminal mouth (
<emphasis id="B9291A72FFEC5975FEA6BCFB359841F8" box="[365,396,1226,1249]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
subsuperior mouth) and by having two conspicuous humeral marks (
<emphasis id="B9291A72FFEC5975FB67BCFB30DE41F8" box="[1196,1226,1226,1249]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
only one humeral mark). In addition,
<taxonomicName id="4C5DBDE3FFEC5975FEB8BCDF35E1401C" box="[371,501,1262,1285]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEC5975FEB8BCDF35E1401C" box="[371,501,1262,1285]" italics="true" pageId="37" pageNumber="38">T. ommatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5DBDE3FFEC5975FD47BCDF3735401C" box="[652,801,1262,1285]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="anostomus">
<emphasis id="B9291A72FFEC5975FD47BCDF3735401C" box="[652,801,1262,1285]" italics="true" pageId="37" pageNumber="38">T. anostomus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5DBDE3FFEC5975FC91BCDF301C401C" box="[858,1032,1262,1285]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="araguaiensis">
<emphasis id="B9291A72FFEC5975FC91BCDF301C401C" box="[858,1032,1262,1285]" italics="true" pageId="37" pageNumber="38">T. araguaiensis</emphasis>
</taxonomicName>
by the number of gill rakers on the lower (13–15) and upper (8–9) limbs of the first gill arch (
<emphasis id="B9291A72FFEC5975FC81BD23377C4030" box="[842,872,1298,1321]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
<date id="FFE3E0A0FFEC5975FCB8BD2330554033" box="[883,1089,1297,1322]" pageId="37" pageNumber="38" value="1917-12-20" valueMax="1917-12-10" valueMin="1917-12-20">17–20 and 10–12</date>
).
<taxonomicName id="4C5DBDE3FFEC5975FB9CBD23316B4030" box="[1111,1407,1298,1321]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEC5975FB9CBD23316B4030" box="[1111,1407,1298,1321]" italics="true" pageId="37" pageNumber="38">Tetragonopterus ommatus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C5DBDE3FFEC5975FEB1BD07361C4054" box="[378,520,1334,1357]" class="Actinopterygii" family="Prochilodontidae" genus="Prochilodus" kingdom="Animalia" order="Characiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="argenteus">
<emphasis id="B9291A72FFEC5975FEB1BD07361C4054" box="[378,520,1334,1357]" italics="true" pageId="37" pageNumber="38">T. argenteus</emphasis>
</taxonomicName>
by having 8 predorsal scales (
<emphasis id="B9291A72FFEC5975FCB0BD07378D4054" box="[891,921,1334,1357]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
11–17). It differs from
<taxonomicName id="4C5DBDE3FFEC5975FB71BD0731524057" box="[1210,1350,1333,1358]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="carvalhoi">
<emphasis id="B9291A72FFEC5975FB71BD0731524057" box="[1210,1350,1333,1358]" italics="true" pageId="37" pageNumber="38">T. carvalhoi</emphasis>
</taxonomicName>
by the presence of a rounded dark mark on the caudal peduncle (
<emphasis id="B9291A72FFEC5975FCEABD6B37544068" box="[801,832,1370,1393]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
a lozenge-shaped dark mark on the caudal peduncle).
<taxonomicName id="4C5DBDE3FFEC5975FF5CBD4F35A8408C" box="[151,444,1406,1429]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEC5975FF5CBD4F35A8408C" box="[151,444,1406,1429]" italics="true" pageId="37" pageNumber="38">Tetragonopterus ommatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5DBDE3FFEC5975FD9FBD4F36BA408C" box="[596,686,1406,1429]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFEC5975FD9FBD4F36BA408C" box="[596,686,1406,1429]" italics="true" pageId="37" pageNumber="38">T. rarus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5DBDE3FFEC5975FD2DBD4F3772408C" box="[742,870,1406,1429]" class="Actinopterygii" family="Characidae" genus="Moenkhausia" kingdom="Animalia" order="Characiformes" pageId="37" pageNumber="38" phylum="Chordata" rank="species" species="georgiae">
<emphasis id="B9291A72FFEC5975FD2DBD4F3772408C" box="[742,870,1406,1429]" italics="true" pageId="37" pageNumber="38">T. georgiae</emphasis>
</taxonomicName>
by the presence of 3.5 scale rows between lateral line and pelvic-fin origin (
<emphasis id="B9291A72FFEC5975FE77BD9335CE40A0" box="[444,474,1442,1465]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
4.5–5.5). Finally,
<taxonomicName id="4C5DBDE3FFEC5975FD6EBD93373140A0" box="[677,805,1442,1465]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEC5975FD6EBD93373140A0" box="[677,805,1442,1465]" italics="true" pageId="37" pageNumber="38">T. ommatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C5DBDE3FFEC5975FC7CBD93301A40A0" box="[951,1038,1442,1465]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="rarus">
<emphasis id="B9291A72FFEC5975FC7CBD93301A40A0" box="[951,1038,1442,1465]" italics="true" pageId="37" pageNumber="38">T. rarus</emphasis>
</taxonomicName>
by the absence of dark longitudinal stripes on the trunk (
<emphasis id="B9291A72FFEC5975FEB5BDF7358940C4" box="[382,413,1478,1501]" italics="true" pageId="37" pageNumber="38">vs.</emphasis>
presence of stripes).
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFEC5977FF0CBDD8360146A9" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="description">
<paragraph id="8BE2C660FFEC5975FF0CBDD8317943AF" blockId="37.[151,1437,865,2006]" pageId="37" pageNumber="38">
<emphasis id="B9291A72FFEC5975FF0CBDD8354D431B" bold="true" box="[199,345,1513,1538]" pageId="37" pageNumber="38">Description.</emphasis>
Morphometric data presented in
<tableCitation id="C6DFF3DBFFEC5975FD20BDD83752431B" box="[747,838,1513,1538]" captionStart="TABLE 8" captionStartId="38.[151,239,580,603]" captionText="TABLE 8. Morphometric data for Tetragonopterus ommatus (n = 5). Range includes holotype and paratypes." pageId="37" pageNumber="38">Table 8</tableCitation>
. Body compressed and moderately elongate when compared with congeners. Greatest body depth at dorsal-fin origin. Dorsal profile slightly convex from snout tip to vertical through middle of orbit; straight from this point to end of occipital process; convex from that point to dorsal-fin origin; straight or slightly convex along dorsal-fin base to rear of adipose fin. Prepelvic region transversely flattened with distinct, longitudinally aligned lateral keels. Ventral profile slightly convex from lower lip to anal-fin origin; straight along base of anal fin. Caudal peduncle with concave dorsal and ventral profiles.
</paragraph>
<paragraph id="8BE2C660FFEC5975FF0CBEF335CD4270" blockId="37.[151,1437,865,2006]" pageId="37" pageNumber="38">Snout shorter than orbital diameter. Mouth terminal. Upper and lower jaw of similar size. Premaxillary teeth in two rows of relatively small teeth with sharp cusps. Outer row with 6 (1) or 7 (4)* teeth with three cusps, central cusp slightly longer. Inner row with 5 (4)* or 6 (1) teeth with four or five cusps, central cusp, when present, twice as long as the lateral cusps. Maxilla with 7 (1)* or 8 (4) teeth with three cusps. Dentary bearing 4 (5)* anteriormost teeth with four or five cusps.</paragraph>
<paragraph id="8BE2C660FFEC5976FF0CBF4435DA4404" blockId="37.[151,1437,865,2006]" lastBlockId="38.[151,1436,151,536]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">Dorsal-fin rays ii,9 (5)*. First unbranched ray shorter than second one. Dorsal-fin origin at middle of the body in SL. Distal margin of fin straight with anterior rays longer. Anal-fin rays v,25 (2) or v,26 (3)*; posterior unbranched rays and anterior branched rays slightly longer than following rays. Anal-fin origin at vertical through two or three scales posterior to last branched dorsal-fin ray. Pelvic-fin rays i,7 (5)*. Pelvic-fin origin located at vertical through dorsal-fin origin. Distal margin of pelvic-fin angled, anterior rays longest. Tip of adpressed pelvic fin not reaching the anal-fin origin. Pectoral-fin rays i,14 (2)*or i,15 (3). Caudal fin forked with i,16,i principal rays (three radiographed adults).</paragraph>
<paragraph id="8BE2C660FFEF5976FF0CB916306C44B4" blockId="38.[151,1436,151,536]" pageId="38" pageNumber="39">Scales large and cycloids. Lateral line complete and distinctly bent downward anteriorly. Longitudinal scales 30 (3) or 31 (2)*. Scale rows between lateral line and dorsal-fin origin 6 (5)*. Scale rows between lateral line and pelvic-fin origin 3.5 (5)*. Predorsal scales 8 (5)*. Scale rows around caudal peduncle 12 (1), 13 (3)* or 14 (1). Anal-fin base covered by a single row of small scales. Lobes of caudal fin poorly scaled.</paragraph>
<paragraph id="8BE2C660FFEF5976FF0CB98636804701" blockId="38.[151,1436,151,536]" pageId="38" pageNumber="39">First gill arch with 8 (3)* or 9 (2) rakers on upper limb and 13 (2)*, 14 (2) or 15 (1) rakers on lower limb. Total vertebrae 30 (3 radiographs), precaudal vertebrae 12, intermediate vertebrae 2, and caudal vertebrae 16 (3 radiographs). Supraneurals 3 (3 radiographs).</paragraph>
<caption id="DF2296E8FFEF5976FF5CBA75310C4742" box="[151,1304,580,603]" pageId="38" pageNumber="39">
<paragraph id="8BE2C660FFEF5976FF5CBA75310C4742" blockId="38.[151,1304,580,603]" box="[151,1304,580,603]" pageId="38" pageNumber="39">
<emphasis id="B9291A72FFEF5976FF5CBA75351E4742" bold="true" box="[151,266,580,603]" pageId="38" pageNumber="39">TABLE 8.</emphasis>
Morphometric data for
<taxonomicName id="4C5DBDE3FFEF5976FDC1BA7537344742" box="[522,800,580,603]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEF5976FDC1BA7537344742" box="[522,800,580,603]" italics="true" pageId="38" pageNumber="39">Tetragonopterus ommatus</emphasis>
</taxonomicName>
(n=5). Range includes holotype and paratypes.
</paragraph>
</caption>
<paragraph id="8BE2C660FFEF5976FD1CBA4735C047C6" blockId="38.[159,1274,630,1644]" pageId="38" pageNumber="39">
<typeStatus id="54E678C2FFEF5976FD1CBA4737214795" box="[727,821,630,652]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
Range Mean Standard length (mm) 64.1 47.6–65.9 61.2 Percentages of standard length
</paragraph>
<paragraph id="8BE2C660FFEF5976FF54BAC335BB40DE" blockId="38.[159,1274,630,1644]" pageId="38" pageNumber="39">Greatest depth 42.2 42.1–44.7 43.5 Predorsal length 50.2 50.2–52.7 50.9 Prepectoral length 31.0 30.8–33.8 31.7 Prepelvic length 51.8 51.8–55.5 53.2 Preanal length 69.9 69.9–73.9 71.4 Caudal peduncle depth 9.36 9.22–10.2 9.71 Caudal peduncle length 9.16 9.03–10.7 9.62 Pectoral-fin length 24.2 24.2–25.5 24.9 Pelvic-fin length 18.5 18.5–20.9 19.8 Dorsal-fin length 31.1 31.1–36.1 33.6 Length of dorsal-fin base 15.7 15.6–17.9 16.5 Anal-fin length 15.9 15.4–19.5 16.4 Length of anal-fin base 29.2 28.4–30.3 29.0 Distance from eye to dorsal-fin origin 46.3 45.9–48.5 46.7 Distance from dorsal-fin origin to caudal-fin base 53.2 51.6–57.4 54.8 Head length 32.6 32.0–35.3 33.1 Head depth 24.6 24.0–27.3 25.2 Percentages of head length</paragraph>
<paragraph id="8BE2C660FFEF5976FF54BDEB37EB42DF" blockId="38.[159,1274,630,1644]" lastBlockId="38.[151,1437,1713,1990]" pageId="38" pageNumber="39">
Snout length 18 18–22 20.2 Maxillary length 46 46–50 48.5 Horizontal orbital diameter 51 49–52 50.6 Least interorbital width 28 25–30 27.9
<emphasis id="B9291A72FFEF5976FF0CBE80358C43D3" bold="true" box="[199,408,1713,1738]" pageId="38" pageNumber="39">Color in alcohol.</emphasis>
General body color pale brown. Dorsal portions of head and body darkly pigmented. Opercular and infraorbital bones silvery. Dorsolateral portion of body with few chromatophores along distal margins of scales. Ventrolateral portion of body unpigmented. Two vertical dark humeral marks spaced by one scale row, with anterior more evident. Anterior humeral mark extending over three horizontal scale rows above lateral line. Posterior humeral mark extending over three horizontal scale rows above lateral line. Caudal peduncle with a ventrally-oriented patch of pigmentation limited to posterior portion of caudal peduncle. Midlateral silvery stripe broad, extending from supracleithrum to caudal peduncle. Anal, pelvic, and dorsal fin hyaline outlined by small chromatophores at distal portions of rays. Adipose fin hyaline (
<figureCitation id="1366DAE5FFEF5976FC54BF9C37E642DF" box="[927,1010,1965,1990]" captionStart="FIGURE 14" captionStartId="39.[151,250,791,813]" captionTargetBox="[151,1436,193,770]" captionTargetId="figure@39.[151,1436,193,770]" captionTargetPageId="39" captionText="FIGURE 14. Tetragonopterus ommatus. MZUSP 92667, holotype, 61.1 mm SL, Brazil, Pará, Itaituba, Rio Tapajós, Amazon basin." httpUri="https://zenodo.org/record/177611/files/figure.png" pageId="38" pageNumber="39">Fig. 14</figureCitation>
).
</paragraph>
<caption id="DF2296E8FFEE5977FF5CBB2634C74655" httpUri="https://zenodo.org/record/177611/files/figure.png" pageId="39" pageNumber="40" targetBox="[151,1436,193,770]" targetPageId="39">
<paragraph id="8BE2C660FFEE5977FF5CBB2634C74655" blockId="39.[151,1436,791,844]" pageId="39" pageNumber="40">
<emphasis id="B9291A72FFEE5977FF5CBB2635344635" bold="true" box="[151,288,791,813]" pageId="39" pageNumber="40">FIGURE 14.</emphasis>
<emphasis id="B9291A72FFEE5977FEE3BB29362D4634" box="[296,569,792,813]" italics="true" pageId="39" pageNumber="40">
<taxonomicName id="4C5DBDE3FFEE5977FEE3BB2936214634" box="[296,565,792,813]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="ommatus">Tetragonopterus ommatus</taxonomicName>
.
</emphasis>
MZUSP 92667, holotype, 61.1 mm SL, Brazil, Pará, Itaituba, Rio Tapajós, Amazon basin.
</paragraph>
</caption>
<paragraph id="8BE2C660FFEE5977FF0CBB42360146A9" blockId="39.[151,1436,883,1088]" pageId="39" pageNumber="40">
<emphasis id="B9291A72FFEE5977FF0CBB4235A34695" bold="true" box="[199,439,883,908]" pageId="39" pageNumber="40">Sexual dimorphism.</emphasis>
Adult males with bony hooks on distal segments of the first 12 branched rays of the anal fin. Two bony hooks per segment.
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFEE5977FF0CBB8A368F46E1" pageId="39" pageNumber="40" type="distribution">
<paragraph id="8BE2C660FFEE5977FF0CBB8A368F46E1" blockId="39.[151,1436,883,1088]" pageId="39" pageNumber="40">
<emphasis id="B9291A72FFEE5977FF0CBB8A357446CD" bold="true" box="[199,352,955,980]" pageId="39" pageNumber="40">Distribution.</emphasis>
<taxonomicName id="4C5DBDE3FFEE5977FEA2BB8D369A46CA" box="[361,654,956,979]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEE5977FEA2BB8D369A46CA" box="[361,654,956,979]" italics="true" pageId="39" pageNumber="40">Tetragonopterus ommatus</emphasis>
</taxonomicName>
is only known near the Cachoeira de São Luiz, middle Rio Tapajós, municipality of Itaituba,
<collectingRegion id="49990882FFEE5977FE66BBD135CB46E1" box="[429,479,992,1016]" country="Brazil" name="Para" pageId="39" pageNumber="40">Pará</collectingRegion>
<collectingCountry id="F34A86F0FFEE5977FE2DBBEE363E46E1" box="[486,554,991,1016]" name="Brazil" pageId="39" pageNumber="40">Brazil</collectingCountry>
(
<figureCitation id="1366DAE5FFEE5977FDF1BBEE366946E1" box="[570,637,991,1016]" captionStart="FIGURE 2" captionStartId="4.[151,250,2001,2023]" captionText="FIGURE 2. Distribution of Tetragonopterus species throughout South America." pageId="39" pageNumber="40">Fig. 2</figureCitation>
b).
</paragraph>
</subSubSection>
<subSubSection id="C34795EBFFEE5977FF0CBC3235FA4159" pageId="39" pageNumber="40" type="etymology">
<paragraph id="8BE2C660FFEE5977FF0CBC3235FA4159" blockId="39.[151,1436,883,1088]" pageId="39" pageNumber="40">
<emphasis id="B9291A72FFEE5977FF0CBC32355A4105" bold="true" box="[199,334,1027,1052]" pageId="39" pageNumber="40">Etymology.</emphasis>
The name
<taxonomicName id="4C5DBDE3FFEE5977FE1BBC3536214102" box="[464,565,1028,1051]" class="Insecta" family="Anthribidae" genus="Tetragonopterus" kingdom="Animalia" order="Coleoptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="ommatus">
<emphasis id="B9291A72FFEE5977FE1BBC3536214102" box="[464,565,1028,1051]" italics="true" pageId="39" pageNumber="40">ommatus</emphasis>
</taxonomicName>
is from Greek
<emphasis id="B9291A72FFEE5977FD2DBC35373E4102" box="[742,810,1028,1051]" italics="true" pageId="39" pageNumber="40">omma</emphasis>
meaning eye, in reference to the great diameter of the orbits observed in this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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