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<document id="932455DFBA45672FFB46E885DF9FC464" ID-DOI="10.5281/zenodo.275511" ID-GBIF-Dataset="ab41269b-1b8d-4772-9b13-df9cbec5ecdf" ID-ISSN="1175-5326" ID-Zenodo-Dep="275511" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460128395670" checkinUser="plazi" docAuthor="Riedel, Alexander" docDate="2010" docId="03EFCD6EFFBB8669FF14FB72FD28F849" docLanguage="en" docName="zt02338p034.pdf" docOrigin="Zootaxa 2338" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Eupholus euphrosyne Porion" docType="treatment" docVersion="7" lastPageNumber="24" masterDocId="FFD6B516FFBA8668FF83FFF7FFE4FFE1" masterDocTitle="Two new species of Eupholus Boisduval (Coleoptera, Curculionidae, Entiminae), with observations on coloured cuticular exudates in weevils" masterLastPageNumber="34" masterPageNumber="23" pageNumber="24" updateTime="1698235076378" updateUser="plazi">
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<mods:title id="7AD9127C9CE604E15DE252B5CC0B2DB0">Two new species of Eupholus Boisduval (Coleoptera, Curculionidae, Entiminae), with observations on coloured cuticular exudates in weevils</mods:title>
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<mods:namePart id="64C21F78FD154CBC0870BD618F6AEA0C">Riedel, Alexander</mods:namePart>
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<mods:date id="78E789A8AC8609DD0F6D5E9151F06B7D">2010</mods:date>
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<treatment id="03EFCD6EFFBB8669FF14FB72FD28F849" ID-DOI="http://doi.org/10.5281/zenodo.6198413" ID-GBIF-Taxon="127654681" ID-Zenodo-Dep="6198413" LSID="urn:lsid:plazi:treatment:03EFCD6EFFBB8669FF14FB72FD28F849" httpUri="http://treatment.plazi.org/id/03EFCD6EFFBB8669FF14FB72FD28F849" lastPageNumber="24" pageId="1" pageNumber="24">
<subSubSection id="C35C2FF3FFBB8669FF14FB72FD24FB23" pageId="1" pageNumber="24" type="nomenclature">
<paragraph id="8BF97C78FFBB8669FF14FB72FDEBFB7E" blockId="1.[151,704,1157,1218]" box="[151,527,1157,1183]" pageId="1" pageNumber="24">
<heading id="D0B1CB14FFBB8669FF14FB72FDEBFB7E" bold="true" box="[151,527,1157,1183]" fontSize="11" level="1" pageId="1" pageNumber="24" reason="1">
<taxonomicName id="4C4607FBFFBB8669FF14FB72FDEBFB7E" authority="Porion" authorityName="Porion" box="[151,527,1157,1183]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="B932A06AFFBB8669FF14FB72FDEBFB7E" bold="true" box="[151,527,1157,1183]" pageId="1" pageNumber="24">
<emphasis id="B932A06AFFBB8669FF14FB72FE44FB7E" bold="true" box="[151,416,1157,1183]" italics="true" pageId="1" pageNumber="24">Eupholus euphrosyne</emphasis>
(Porion)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF97C78FFBB8669FF14FB5FFD24FB23" blockId="1.[151,704,1157,1218]" box="[151,704,1192,1218]" pageId="1" pageNumber="24">
(
<figureCitation id="137D60FDFFBB8669FF1CFB5FFF15FB23" box="[159,241,1192,1218]" captionStart="FIGURES 1 4" captionStartId="2.[151,269,1957,1981]" captionTargetBox="[266,1292,190,1903]" captionTargetId="figure@2.[266,1307,178,1911]" captionTargetPageId="2" captionText="FIGURES 1 4. Habitus of Eupholus spp.: (1) E. mimicus, holotype (2) E. sedlaceki, holotype; (3) E. euphrosyne, female (Aseki); (4) E. sedlaceki, female (Wau)." httpUri="https://zenodo.org/record/275512/files/figure.png" pageId="1" pageNumber="24">Figs. 3</figureCitation>
,
<figureCitation id="137D60FDFFBB8669FF7CFB5FFEEBFB23" box="[255,271,1192,1218]" captionStart="FIGURES 5 7" captionStartId="3.[151,269,850,874]" captionTargetBox="[266,1320,220,828]" captionTargetId="figure@3.[266,1321,194,828]" captionTargetPageId="3" captionText="FIGURES 5 7. Head and rostrum of Eupholus spp.: (5) E. euphrosyne; (6) E. sedlaceki; (7) E. mimicus." httpUri="https://zenodo.org/record/275513/files/figure.png" pageId="1" pageNumber="24">5</figureCitation>
,
<figureCitation id="137D60FDFFBB8669FE9FFB5FFEBDFB23" box="[284,345,1192,1218]" captionStart="FIGURES 8 10" captionStartId="4.[151,270,1683,1707]" captionTargetBox="[316,1272,554,1662]" captionTargetId="figure@4.[304,1282,546,1662]" captionTargetPageId="4" captionText="FIGURES 8 10. Male genitalia of Eupholus euphrosyne: (8) aedeagus in lateral aspect; (9) aedeagus in dorsal aspect; (10) detail of transfer apparatus." httpUri="https://zenodo.org/record/275514/files/figure.png" pageId="1" pageNumber="24">810</figureCitation>
,
<figureCitation id="137D60FDFFBB8669FEE6FB5FFE39FB23" box="[357,477,1192,1218]" captionStart="FIGURES 14 22" captionStartId="5.[151,269,1944,1968]" captionTargetBox="[311,1267,402,1924]" captionTargetId="figure@5.[308,1278,391,1929]" captionTargetPageId="5" captionText="FIGURES 14 22. Female genitalia of Eupholus spp., upper row: overview in ventral aspect; middle row: vagina, bursa copulatrix and base of hemisternites in ventral aspect; lower row: left hemisternite. (14, 17, 20) E. euphrosyne; (15, 18, 21) E. sedlaceki; (16, 19, 22) E. mimicus." httpUri="https://zenodo.org/record/275515/files/figure.png" pageId="1" pageNumber="24">14, 17, 20</figureCitation>
,
<figureCitation id="137D60FDFFBB8669FE6AFB5FFD85FB23" box="[489,609,1192,1218]" captionStart="FIGURES 23 31" captionStartId="6.[151,270,1269,1293]" captionTargetBox="[276,1319,200,1230]" captionTargetId="figure@6.[264,1323,179,1244]" captionTargetPageId="6" captionText="FIGURES 23 31. Female genitalia of Eupholus spp.: (23 25) sternite VIII; (26 28) tergite VIII; (29 31) spermatheca. (23, 26, 30) E. euphrosyne; (24, 27, 31) E. sedlaceki; (25, 28, 29) E. mimicus." httpUri="https://zenodo.org/record/275516/files/figure.png" pageId="1" pageNumber="24">23, 26, 30</figureCitation>
,
<figureCitation id="137D60FDFFBB8669FDEEFB5FFD5CFB23" box="[621,696,1192,1218]" captionStart="FIGURES 32 37" captionStartId="8.[151,273,1538,1562]" captionTargetBox="[264,1322,298,1516]" captionTargetId="figure@8.[264,1323,298,1517]" captionTargetPageId="8" captionText="FIGURES 32 37. Base of left elytron of Eupholini with yellow coloration; overview (left); detail (right): (32 33) Eupholus mimicus; note yellow scales; (34 35) Eupholus euphrosyne; note cream-coloured setae holding yellow particles; (36 37) Rhinoscapha cf. thomsoni; note incrustation of yellow substance." httpUri="https://zenodo.org/record/275517/files/figure.png" pageId="1" pageNumber="24">3435</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35C2FF3FFBB8669FF14FB05FBC4FAE8" box="[151,1056,1266,1289]" pageId="1" pageNumber="24" type="reference_group">
<paragraph id="8BF97C78FFBB8669FF14FB05FBC4FAE8" blockId="1.[151,1056,1266,1289]" box="[151,1056,1266,1289]" pageId="1" pageNumber="24">
<treatmentCitationGroup id="AB565B56FFBB8669FF14FB05FBC4FAE8" box="[151,1056,1266,1289]" pageId="1" pageNumber="24">
<taxonomicName id="4C4607FBFFBB8669FF14FB05FDD5FAE8" authority="Porion 1993: 12" authorityName="Porion" authorityPageNumber="12" authorityYear="1993" box="[151,561,1266,1289]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="euphrosyne">
<emphasis id="B932A06AFFBB8669FF14FB05FE98FAE8" box="[151,380,1266,1289]" italics="true" pageId="1" pageNumber="24">Eupholus euphrosyne</emphasis>
<treatmentCitation id="0AE75A69FFBB8669FE01FB05FDD5FAE8" author="Porion" box="[386,561,1266,1289]" page="12" pageId="1" pageNumber="24" year="1993">
<bibRefCitation id="EFD70189FFBB8669FE01FB05FDEDFAE8" author="Porion" box="[386,521,1266,1289]" pageId="1" pageNumber="34" refString="Porion, T. (1993) Eupholus. Les Coleopteres du Monde - The Beetles of the World, Vol. 19. 112 pp. Sciences Nat, Venette." type="book" year="1993">Porion 1993</bibRefCitation>
: 12
</treatmentCitation>
</taxonomicName>
. (
<taxonomicName id="4C4607FBFFBB8669FDC7FB05FC96FAE8" ID-CoL="8HJKK" authority="Guerin-Meneville" authorityName="Guerin-Meneville" box="[580,882,1266,1289]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B932A06AFFBB8669FDC7FB05FD4DFAE8" box="[580,681,1266,1289]" italics="true" pageId="1" pageNumber="24">Eupholus</emphasis>
Guérin-Méneville
</taxonomicName>
, non Boisduval)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35C2FF3FFBB8669FF14FAC0FC22F9B9" pageId="1" pageNumber="24" type="diagnosis">
<paragraph id="8BF97C78FFBB8669FF14FAC0FC22F9B9" blockId="1.[151,1437,1335,1960]" pageId="1" pageNumber="24">
<emphasis id="B932A06AFFBB8669FF14FAC0FEF4FAB0" bold="true" box="[151,272,1335,1361]" pageId="1" pageNumber="24">Diagnosis</emphasis>
. A
<taxonomicName id="4C4607FBFFBB8669FEBAFACFFE4CFAB0" box="[313,424,1336,1361]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="genus">
<emphasis id="B932A06AFFBB8669FEBAFACFFE4CFAB0" box="[313,424,1336,1361]" italics="true" pageId="1" pageNumber="24">Eupholus</emphasis>
</taxonomicName>
species with two transverse yellow bands on black elytra. Easily distinguished from
<taxonomicName id="4C4607FBFFBB8669FA00FACFFF1EFA97" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="mimicus" status="sp. nov.">
<emphasis id="B932A06AFFBB8669FA00FACFFF1EFA97" italics="true" pageId="1" pageNumber="24">E. mimicus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2011D11FFBB8669FE81FAAAFEA6FA96" box="[258,322,1373,1399]" pageId="1" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
by the yellow bands being composed of yellow extracuticular particles (
<figureCitation id="137D60FDFFBB8669FB2FFAAAFADBFA96" box="[1196,1343,1373,1399]" captionStart="FIGURES 32 37" captionStartId="8.[151,273,1538,1562]" captionTargetBox="[264,1322,298,1516]" captionTargetId="figure@8.[264,1323,298,1517]" captionTargetPageId="8" captionText="FIGURES 32 37. Base of left elytron of Eupholini with yellow coloration; overview (left); detail (right): (32 33) Eupholus mimicus; note yellow scales; (34 35) Eupholus euphrosyne; note cream-coloured setae holding yellow particles; (36 37) Rhinoscapha cf. thomsoni; note incrustation of yellow substance." httpUri="https://zenodo.org/record/275517/files/figure.png" pageId="1" pageNumber="24">Figs. 3435</figureCitation>
), not of scales. Both dorsal and ventral surface subglabrous, without recumbent cream-coloured amygdaloid scales except on apical margin of abdominal ventrites, but often body with scattered, relatively long, suberect setae. Larger than
<taxonomicName id="4C4607FBFFBB8669FEA4FA39FE51FA06" box="[295,437,1486,1511]" class="Insecta" family="Curculionidae" genus="Eupholus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="24" phylum="Arthropoda" rank="species" species="sedlaceki">
<emphasis id="B932A06AFFBB8669FEA4FA39FE51FA06" box="[295,437,1486,1511]" italics="true" pageId="1" pageNumber="24">E. sedlaceki</emphasis>
</taxonomicName>
: total length of males 21.022.0 mm (n=4, =21.4 ± 0.5), of females 22.524.0 mm (n=2, =23.3); pronotum + elytron of males 16.017.0 mm (n=5, =16.4 ± 0.5), pronotum + elytron of females 18.019.0 mm (n=2, =18.5). Yellow elytral bands narrow, area between them wider than a band; anterior margin of posterior band evenly convex across elytral suture.
</paragraph>
</subSubSection>
<subSubSection id="C35C2FF3FFBB8669FF45F995FBEBF8F3" pageId="1" pageNumber="24" type="materials_examined">
<paragraph id="8BF97C78FFBB8669FF45F995FBEBF8F3" blockId="1.[151,1437,1335,1960]" pageId="1" pageNumber="24">
<emphasis id="B932A06AFFBB8669FF45F995FE56F99D" bold="true" box="[198,434,1634,1660]" pageId="1" pageNumber="24">Material examined</emphasis>
.
<collectingCountry id="F3513CE8FFBB8669FE42F994FD07F99C" box="[449,739,1635,1661]" name="Papua New Guinea" pageId="1" pageNumber="24">
<collectionCode id="ED57E4BDFFBB8669FE42F994FDFBF99C" box="[449,543,1635,1661]" pageId="1" pageNumber="24">PAPUA</collectionCode>
<collectionCode id="ED57E4BDFFBB8669FDA5F994FD8DF99C" box="[550,617,1635,1661]" httpUri="http://grbio.org/cool/93c9-w2q7" name="University of Newcastle" pageId="1" pageNumber="24">NEW</collectionCode>
<collectionCode id="ED57E4BDFFBB8669FDECF994FD07F99C" box="[623,739,1635,1661]" pageId="1" pageNumber="24">GUINEA</collectionCode>
</collectingCountry>
, Morobe Prov.:
<specimenCount id="9D40B7F1FFBB8669FC1DF994FC1FF99C" box="[926,1019,1635,1661]" pageId="1" pageNumber="24" type="male">2 males</specimenCount>
, Watut to Aseki, div nr Aseki, 1250 [m], leg. J.L.Gressitt,
<date id="FFF85AB8FFBB8669FE22F97FFDC8F943" box="[417,556,1672,1698]" pageId="1" pageNumber="24" value="1974-11-30">30.XI.1974</date>
, “K-956” (
<collectionCode id="ED57E4BDFFBB8669FD32F97FFCEFF943" LSID="urn:lsid:biocol.org:col:34697" box="[689,779,1672,1698]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="24">BPBM</collectionCode>
);
<specimenCount id="9D40B7F1FFBB8669FCA3F97FFC90F943" box="[800,884,1672,1698]" pageId="1" pageNumber="24" type="male">1 male</specimenCount>
, Aseki,
<quantity id="4CBED19DFFBB8669FC54F97FFBD0F943" box="[983,1076,1672,1698]" metricMagnitude="3" metricUnit="m" metricValue="1.1199999999999999" pageId="1" pageNumber="24" unit="m" value="1120.0">1120 m</quantity>
, leg. J.L.Gressitt,
<date id="FFF85AB8FFBB8669FA93F97FFA73F943" box="[1296,1431,1672,1698]" pageId="1" pageNumber="24" value="1974-04-13">13.IV.1974</date>
, “?
<collectionCode id="ED57E4BDFFBB8669FF2DF959FF12F929" box="[174,246,1710,1736]" pageId="1" pageNumber="24">PITT</collectionCode>
06P”;
<specimenCount id="9D40B7F1FFBB8669FEC3F959FE4AF929" box="[320,430,1710,1736]" pageId="1" pageNumber="24" type="female">1 female</specimenCount>
, Upper Watut,
<quantity id="4CBED19DFFBB8669FDEAF959FCE1F929" box="[617,773,1710,1736]" metricMagnitude="3" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="1" pageNumber="24" unit="m" value="1050.0" valueMax="1200.0" valueMin="900.0">9001200 m</quantity>
,
<date id="FFF85AB8FFBB8669FC90F959FC2AF929" box="[787,974,1710,1736]" pageId="1" pageNumber="24" value="1962-03-20" valueMax="1962-03-28" valueMin="1962-03-20">2028.III.1962</date>
, Native Collector (
<collectionCode id="ED57E4BDFFBB8669FB3AF959FAF0F929" LSID="urn:lsid:biocol.org:col:34697" box="[1209,1300,1710,1736]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="1" pageNumber="24">BPBM</collectionCode>
);
<specimenCount id="9D40B7F1FFBB8669FAA9F959FA7CF929" box="[1322,1432,1710,1736]" pageId="1" pageNumber="24" type="female">1 female</specimenCount>
, Aseki, Oiwa,
<quantity id="4CBED19DFFBB8669FEBDF924FE03F90C" box="[318,487,1747,1773]" metricMagnitude="3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="1" pageNumber="24" unit="m" value="1650.0" valueMax="1700.0" valueMin="1600.0">16001700 m</quantity>
,
<date id="FFF85AB8FFBB8669FE76F924FD4FF90C" box="[501,683,1747,1773]" pageId="1" pageNumber="24" value="1998-03-11" valueMax="1998-03-13" valueMin="1998-03-11">1113.III.1998</date>
, leg. A.Riedel (
<collectionCode id="ED57E4BDFFBB8669FCE9F924FC4AF90C" box="[874,942,1747,1773]" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" pageId="1" pageNumber="24">ARC</collectionCode>
);
<specimenCount id="9D40B7F1FFBB8669FC47F924FBFDF90C" box="[964,1049,1747,1773]" pageId="1" pageNumber="24" type="male">1 male</specimenCount>
, Aseki,
<date id="FFF85AB8FFBB8669FBFFF924FB07F90C" box="[1148,1251,1747,1773]" pageId="1" pageNumber="24" value="1992-06">VI.1992</date>
, local collector (
<collectionCode id="ED57E4BDFFBB8669FF1CF90FFF06F8F3" box="[159,226,1784,1810]" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" pageId="1" pageNumber="24">ARC</collectionCode>
);
<specimenCount id="9D40B7F1FFBB8669FF75F90FFEACF8F3" box="[246,328,1784,1810]" pageId="1" pageNumber="24" type="male">1 male</specimenCount>
, Aseki, Hiyewini village,
<date id="FFF85AB8FFBB8669FDFAF90FFD1AF8F3" box="[633,766,1784,1810]" pageId="1" pageNumber="24" value="1999-04-30">30.IV.1999</date>
, local collector (
<collectionCode id="ED57E4BDFFBB8669FC43F90FFBE7F8F3" box="[960,1027,1784,1810]" httpUri="http://grbio.org/cool/r82x-h8re" name="Atlantic Reference Centre" pageId="1" pageNumber="24">ARC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C35C2FF3FFBB8669FF45F8EAFD28F849" pageId="1" pageNumber="24" type="distribution">
<paragraph id="8BF97C78FFBB8669FF45F8EAFB0AF8D9" blockId="1.[151,1437,1335,1960]" box="[198,1262,1821,1848]" pageId="1" pageNumber="24">
<emphasis id="B932A06AFFBB8669FF45F8EAFEBBF8D6" bold="true" box="[198,351,1821,1847]" pageId="1" pageNumber="24">Distribution</emphasis>
. Only known from the area of Aseki (Morobe Province,
<collectingCountry id="F3513CE8FFBB8669FC79F8E9FB05F8D9" box="[1018,1249,1822,1848]" name="Papua New Guinea" pageId="1" pageNumber="24">Papua New Guinea</collectingCountry>
).
</paragraph>
<paragraph id="8BF97C78FFBB8669FF45F8B5FD28F849" blockId="1.[151,1437,1335,1960]" pageId="1" pageNumber="24">
<emphasis id="B932A06AFFBB8669FF45F8B5FEE8F8BD" bold="true" box="[198,268,1858,1884]" pageId="1" pageNumber="24">Notes</emphasis>
. The
<typeStatus id="54FDC2DAFFBB8669FED2F8B4FE53F8BC" box="[337,439,1859,1885]" pageId="1" pageNumber="24" type="holotype">holotype</typeStatus>
is still housed in TPCJ, not at the Natural History Museum Paris as stated by
<bibRefCitation id="EFD70189FFBB8669FACDF8B4FF0DF863" author="Porion" pageId="1" pageNumber="34" refString="Porion, T. (1993) Eupholus. Les Coleopteres du Monde - The Beetles of the World, Vol. 19. 112 pp. Sciences Nat, Venette." type="book" year="1993">Porion (1993)</bibRefCitation>
. It was not possible to examine it, but T. Porion provided a colour photograph of the
<typeStatus id="54FDC2DAFFBB8669FB5EF89FFAA3F863" box="[1245,1351,1896,1922]" pageId="1" pageNumber="24" type="holotype">holotype</typeStatus>
, which was sufficient to determine the species´identity.
</paragraph>
</subSubSection>
</treatment>
</document>