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<document id="53AC01770F5BBCF17BF7BFD5151ED28B" ID-CLB-Dataset="8324" ID-DOI="10.11646/zootaxa.4920.4.3" ID-GBIF-Dataset="3f6c02fc-66e9-4ad6-a759-ef23da595c48" ID-ISSN="1175-5326" ID-Zenodo-Dep="4491181" ID-ZooBank="22E8AB54-654D-4882-9084-41F0D229D010" IM.bibliography_approvedBy="tatiana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="tatiana" checkinTime="1612249726991" checkinUser="plazi" docAuthor="Ruschel, Tatiana P. &amp; Sanborn, Allen F." docDate="2021" docId="03D787C8EC48FFCDFFE05979FDDE36F6" docLanguage="en" docName="zootaxa.4920.4.3.pdf" docOrigin="Zootaxa 4920 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Nosola Stal 1866" docType="treatment" docVersion="9" lastPageNumber="511" masterDocId="FFEEFFB0EC4AFFCEFF775816FFDA377F" masterDocTitle="Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae)" masterLastPageNumber="527" masterPageNumber="509" pageNumber="511" updateTime="1698926442746" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="1AC2CC0E3736F77B6018322B2AF2346A">Two new genera and species of Fidicinini Distant, 1905 with a re-description of Nosola Stål, 1866 (Hemiptera, Auchenorrhyncha, Cicadidae)</mods:title>
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<mods:namePart id="232F9D42F021652171B5021128016C4C">Ruschel, Tatiana P.</mods:namePart>
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<mods:affiliation id="60AF4187CC5C08BE413B1411E294F9E7">Plazi, Rio Grande do Sul, Porto Alegre, RS, Brazil. urn: lsid: zoobank. org: author: 9 D 376223 - 5637 - 4376 - 9 CBC- 9 F 98441455 A 6 tatiana. ruschel @ gmail. com; https: // orcid. org / 0000 - 0002 - 9052 - 1760</mods:affiliation>
<mods:nameIdentifier id="3F3BE3557753D1302595E85126D4F3D7" type="email">tatiana.ruschel@gmail.com</mods:nameIdentifier>
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<mods:namePart id="027F3C9B0DC076083B35C400E4E74D5F">Sanborn, Allen F.</mods:namePart>
<mods:affiliation id="EDDF11C27A7DA51B0F655E16EA55A96A">Department of Biology, Barry University, 11300 NE Second Avenue, Miami Shores, FL 33161 - 6695, USA. urn: lsid: zoobank. org: author: 9544 FF 3 F- 9 B 1 F- 457 A- 90 A 2 - 8 A 1 C 0 C 1 F 22 A 0</mods:affiliation>
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<mods:title id="0D3D8D0EFE0DBCA3962562B6A5202865">Zootaxa</mods:title>
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<mods:date id="3BE51A1D69F43DE11E43BA54D21F918F">2021</mods:date>
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<treatment id="03D787C8EC48FFCDFFE05979FDDE36F6" ID-DOI="http://doi.org/10.5281/zenodo.4531850" ID-GBIF-Taxon="177317070" ID-Zenodo-Dep="4531850" LSID="urn:lsid:plazi:treatment:03D787C8EC48FFCDFFE05979FDDE36F6" httpUri="http://treatment.plazi.org/id/03D787C8EC48FFCDFFE05979FDDE36F6" lastPageId="3" lastPageNumber="511" pageId="2" pageNumber="511">
<subSubSection id="C3646555EC48FFCCFFE05979FEA636F6" box="[151,380,366,394]" pageId="2" pageNumber="511" type="nomenclature">
<paragraph id="8BC136DEEC48FFCCFFE05979FEA636F6" blockId="2.[151,380,366,429]" box="[151,380,366,394]" pageId="2" pageNumber="511">
<heading id="D08981B2EC48FFCCFFE05979FEA636F6" bold="true" box="[151,380,366,394]" fontSize="11" level="1" pageId="2" pageNumber="511" reason="1">
<taxonomicName id="4C7E4D5DEC48FFCCFFE05979FEA636F6" ID-CoL="8HLGX" ID-ENA="2218838" authority="Stal, 1866 a" authorityName="Stal" authorityYear="1866" box="[151,380,366,394]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACCEC48FFCCFFE05979FEA636F6" bold="true" box="[151,380,366,394]" pageId="2" pageNumber="511">
<emphasis id="B90AEACCEC48FFCCFFE05979FF3636F5" bold="true" box="[151,236,367,394]" italics="true" pageId="2" pageNumber="511">Nosola</emphasis>
<bibRefCitation id="EFEF4B2FEC48FFCCFF845978FEA636F6" author="Stal" box="[243,380,366,393]" firstAuthor="Stal" pageId="2" pageNumber="511" pagination="1 - 276" refId="ref10871" refString="Stal, C. (1866 a) Hemiptera Homoptera Latr. Hemiptera Africana, 4, 1 - 276, pl. 1. https: // doi. org / 10.5962 / bhl. title. 8566" type="journal article" year="1866" yearSuffix="a">Stål, 1866a</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="C3646555EC48FFCCFFE05985FEC836D2" box="[151,274,403,429]" pageId="2" pageNumber="511" type="description">
<paragraph id="8BC136DEEC48FFCCFFE05985FEC836D2" blockId="2.[151,380,366,429]" box="[151,274,403,429]" pageId="2" pageNumber="511">
(
<figureCitation id="13452A5BEC48FFCCFFE85985FED036D2" box="[159,266,403,429]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1031,1055]" captionStartId-1="3.[151,250,1778,1802]" captionStartId-2="5.[151,250,1689,1713]" captionStartId-3="6.[151,250,1898,1922]" captionTargetBox-0="[151,1435,345,1007]" captionTargetBox-1="[151,1435,417,1755]" captionTargetBox-2="[151,1436,186,1665]" captionTargetBox-3="[161,1427,309,1875]" captionTargetId-0="figure-108@1.[151,1435,345,1007]" captionTargetId-1="figure-129@3.[151,1435,417,1755]" captionTargetId-2="figure-19@5.[151,1436,181,1665]" captionTargetId-3="figure-97@6.[161,1427,309,1875]" captionTargetPageId-0="1" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Nosola paradoxa Stål, 1866, holotype male in dorsal view and labels. Scale bar = 50 mm. [Photograph courtesy of Gunvi Lindberg, Swedish Museum of Natural History]." captionText-1="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." captionText-2="FIGURE 3. Nosola paradoxa Stål, 1866, male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in lateroventral view; D, phallus in left view; E, phallus in right view. Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), basal plate (bp), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). The arrow indicates the bud-like lobe developed between the dorsal crest and the lateral branches. Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." captionText-3="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4491183" figureDoi-1="http://doi.org/10.5281/zenodo.4491187" figureDoi-2="http://doi.org/10.5281/zenodo.4491189" figureDoi-3="http://doi.org/10.5281/zenodo.4491192" httpUri-0="https://zenodo.org/record/4491183/files/figure.png" httpUri-1="https://zenodo.org/record/4491187/files/figure.png" httpUri-2="https://zenodo.org/record/4491189/files/figure.png" httpUri-3="https://zenodo.org/record/4491192/files/figure.png" pageId="2" pageNumber="511">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC48FFCCFFE059CEFDCD368F" box="[151,535,471,496]" pageId="2" pageNumber="511" type="reference_group">
<paragraph id="8BC136DEEC48FFCCFFE059CEFDCD368F" blockId="2.[151,535,471,496]" box="[151,535,471,496]" pageId="2" pageNumber="511">
<treatmentCitationGroup id="AB6E11F0EC48FFCCFFE059CEFDCD368F" box="[151,535,471,496]" pageId="2" pageNumber="511">
<treatmentCitation id="0ADF10CFEC48FFCCFFE059CEFEA83690" author="Stal, C." box="[151,370,471,496]" page="7" pageId="2" pageNumber="511" year="1866">
<taxonomicName id="4C7E4D5DEC48FFCCFFE059CEFEA83690" ID-CoL="8HLGX" authority="Stal, 1866 a: 7" authorityName="Stal" authorityPageNumber="7" authorityYear="1866" box="[151,370,471,496]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="511" phylum="Arthropoda" rank="genus">
<emphasis id="B90AEACCEC48FFCCFFE059CEFF05368F" box="[151,223,472,496]" italics="true" pageId="2" pageNumber="511">Nosola</emphasis>
<bibRefCitation id="EFEF4B2FEC48FFCCFF9259C1FEA83690" author="Stal" box="[229,370,471,495]" firstAuthor="Stal" page="7" pageId="2" pageNumber="511" pagination="1 - 276" refId="ref10871" refString="Stal, C. (1866 a) Hemiptera Homoptera Latr. Hemiptera Africana, 4, 1 - 276, pl. 1. https: // doi. org / 10.5962 / bhl. title. 8566" type="journal article" year="1866" yearSuffix="a">Stål, 1866a: 7</bibRefCitation>
</taxonomicName>
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(gen. nov.; key)
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</paragraph>
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<subSubSection id="C3646555EC48FFCCFFE05A03FC563552" box="[151,908,533,557]" pageId="2" pageNumber="511" type="type_taxon">
<paragraph id="8BC136DEEC48FFCCFFE05A03FC563552" blockId="2.[151,908,533,557]" box="[151,908,533,557]" pageId="2" pageNumber="511">
<emphasis id="B90AEACCEC48FFCCFFE05A03FEF93552" bold="true" box="[151,291,533,557]" pageId="2" pageNumber="511">
<typeStatus id="54C5887CEC48FFCCFFE05A03FF163552" box="[151,204,533,557]" pageId="2" pageNumber="511">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C7E4D5DEC48FFCCFE5D5A03FD8F3552" ID-CoL="6SB43" authority="Stal, 1866 b" authorityName="Stal" authorityYear="1866" box="[298,597,533,557]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="511" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B90AEACCEC48FFCCFE5D5A03FE033552" box="[298,473,533,557]" italics="true" pageId="2" pageNumber="511">Nosola paradoxa</emphasis>
<bibRefCitation id="EFEF4B2FEC48FFCCFEA85A03FD8F3552" author="Stal" box="[479,597,533,557]" firstAuthor="Stal" pageId="2" pageNumber="511" pagination="151 - 172" refId="ref10910" refString="Stal, C. (1866 b) Analecta Hemipterologica. Berliner Entomologischer Zeitschrift, 10, 151 - 172. https: // doi. org / 10.1002 / mmnd. 18660100112" type="journal article" year="1866" yearSuffix="b">Stål, 1866b</bibRefCitation>
</taxonomicName>
, designated by
<bibRefCitation id="EFEF4B2FEC48FFCCFD855A03FC523552" author="Distant" box="[754,904,533,557]" firstAuthor="Distant" pageId="2" pageNumber="511" refId="ref9367" refString="Distant, W. L. (1906) A Synonymic Catalogue of Homoptera. Part 1. Cicadidae. Trustees of the British Museum, London, 207 pp. https: // doi. org / 10.5962 / bhl. title. 8554" type="book" year="1906">Distant (1906)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC48FFCCFFE05A4DFD60350A" box="[151,698,603,630]" pageId="2" pageNumber="511" type="taxon_list">
<paragraph id="8BC136DEEC48FFCCFFE05A4DFD60350A" blockId="2.[151,1437,603,2034]" box="[151,698,603,630]" pageId="2" pageNumber="511">
<emphasis id="B90AEACCEC48FFCCFFE05A4DFEB9350A" bold="true" box="[151,355,603,629]" pageId="2" pageNumber="511">Species included.</emphasis>
<taxonomicName id="4C7E4D5DEC48FFCCFE1D5A4DFD6C350A" authority="Stal, 1866 b" authorityName="Stal" authorityYear="1866" box="[362,694,603,630]" class="Insecta" family="Cicadidae" genus="Nosola" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="511" phylum="Arthropoda" rank="species" species="paradoxa">
<emphasis id="B90AEACCEC48FFCCFE1D5A4DFDF1350A" box="[362,555,603,629]" italics="true" pageId="2" pageNumber="511">Nosola paradoxa</emphasis>
<bibRefCitation id="EFEF4B2FEC48FFCCFD445A4DFD6C350A" author="Stal" box="[563,694,603,630]" firstAuthor="Stal" pageId="2" pageNumber="511" pagination="151 - 172" refId="ref10910" refString="Stal, C. (1866 b) Analecta Hemipterologica. Berliner Entomologischer Zeitschrift, 10, 151 - 172. https: // doi. org / 10.1002 / mmnd. 18660100112" type="journal article" year="1866" yearSuffix="b">Stål, 1866b</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC48FFCCFFB05A69FDE9359E" pageId="2" pageNumber="511" type="discussion">
<paragraph id="8BC136DEEC48FFCCFFB05A69FDE9359E" blockId="2.[151,1437,603,2034]" pageId="2" pageNumber="511">
<emphasis id="B90AEACCEC48FFCCFFB05A69FEE135E6" bold="true" box="[199,315,639,665]" pageId="2" pageNumber="511">Remarks.</emphasis>
The genus was designated in a key based on the obtusely conical and anteriorly produced postclypeus and the strongly curved base of the costal margin of the fore wing (
<bibRefCitation id="EFEF4B2FEC48FFCCFCDB5AB5FBEB35C1" author="Stal" box="[940,1073,675,702]" firstAuthor="Stal" pageId="2" pageNumber="511" pagination="151 - 172" refId="ref10910" refString="Stal, C. (1866 b) Analecta Hemipterologica. Berliner Entomologischer Zeitschrift, 10, 151 - 172. https: // doi. org / 10.1002 / mmnd. 18660100112" type="journal article" year="1866" yearSuffix="b">Stål, 1866b</bibRefCitation>
). No complete description of the genus has been produced previously.
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC48FFCCFFB05AFDFCA93066" pageId="2" pageNumber="511" type="description">
<paragraph id="8BC136DEEC48FFCCFFB05AFDFC623479" blockId="2.[151,1437,603,2034]" box="[199,952,747,774]" pageId="2" pageNumber="511">
<emphasis id="B90AEACCEC48FFCCFFB05AFDFE83347A" bold="true" box="[199,345,747,773]" pageId="2" pageNumber="511">Description.</emphasis>
Medium sized cicada (body length about
<quantity id="4C869B3BEC48FFCCFC585AFDFC763479" box="[815,940,747,774]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" metricValueMax="2.9" metricValueMin="2.5" pageId="2" pageNumber="511" unit="mm" value="27.0" valueMax="29.0" valueMin="25.0">25–29 mm</quantity>
).
</paragraph>
<paragraph id="8BC136DEEC48FFCCFFB05B19FB8931D1" blockId="2.[151,1437,603,2034]" pageId="2" pageNumber="511">
Head broader than anterior pronotum but not as wide as pronotal collar, about as wide as mesonotum; eyes larger than space of vertex between internal margin of eyes and frontoclypeal suture; lateral ocelli widely separated, separated by a greater distance from each other than from median ocellus (
<figureCitation id="13452A5BEC48FFCCFCAB5B41FBEF340E" box="[988,1077,855,881]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2B</figureCitation>
). Supra–antennal plate reaching half distance to eye (
<figureCitation id="13452A5BEC48FFCCFEF75B6DFE0234EA" box="[384,472,891,917]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2B</figureCitation>
). Postclypeus very prominent extending well beyond anterior margin of head forming acute angle in dorsal view, longer than vertex (
<figureCitation id="13452A5BEC48FFCCFDD45B89FD2134C6" box="[675,763,927,953]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2B</figureCitation>
), rectangular rectangular in ventral view (
<figureCitation id="13452A5BEC48FFCCFBA65B89FAF334C6" box="[1233,1321,927,953]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2C</figureCitation>
), obtusely angled in lateral view. Anteclypeus with ventralsurface tumid, lateroposterior margins inclined forming acute angle, posterolateral flaps absent (
<figureCitation id="13452A5BEC48FFCCFEA75BF1FDF1337E" box="[464,555,999,1025]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2C</figureCitation>
). Pronotum shorter than mesonotum, lateral part of pronotal collar broad and convex (
<figureCitation id="13452A5BEC48FFCCFF8E5C1DFE88335A" box="[249,338,1035,1061]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2B</figureCitation>
). Mesonotum covering dorsal metanotum; cruciform elevation short, reaching anterior margin of abdominal tergite 1, central area swollen, lateral areas flat, angle of posterolateral projections obtuse (
<figureCitation id="13452A5BEC48FFCCFA725C39FA853336" box="[1285,1375,1071,1097]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2A</figureCitation>
). Anterior basisternum 3 obtusely angled, not prominent relative to mesocoxae (
<figureCitation id="13452A5BEC48FFCCFC9C5C45FB9C3312" box="[1003,1094,1107,1133]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2D</figureCitation>
); three segmented tarsi; wings hyaline, forewing costal margin strongly curved and widened at base, radial cell expanded, eight apical cells long, extending about one third of forewing length, apical cell 2 about half apical cell 1 length, infuscation present, marginal area present; hindwing with six apical cells, cubital cell 1 expanding to junction with apical cell 6, anal lobe broader than cubital cell 1, anal vein 3 curved at distal terminus, marginal area present (
<figureCitation id="13452A5BEC48FFCCFB025CF5FB153382" box="[1141,1231,1251,1277]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2A</figureCitation>
). Male operculum semilunar, not meeting medially; meracanthus short not reaching middle operculum; lateral metascutellar plate not covering posterolateral timbal chamber (
<figureCitation id="13452A5BEC48FFCCFD2D5D3DFD6B323A" box="[602,689,1323,1349]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2E</figureCitation>
). Female operculum triangular, short, apex not surpassing posterior sternite II (
<figureCitation id="13452A5BEC48FFCCFE615D59FEAB3216" box="[278,369,1359,1385]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="2" pageNumber="511">Fig. 4C</figureCitation>
). Male abdomen longer than distance between apex of posclypeus and cruciform elevation in dorsal view, tergite 1 visible, about as long as median portion length of tergite 2 but longer than cruciform elevation (
<figureCitation id="13452A5BEC48FFCCFFA55D81FEF732CE" box="[210,301,1431,1457]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2A</figureCitation>
). Timbal cover flat, incomplete dorsally exposing dorsal timbal (
<figureCitation id="13452A5BEC48FFCCFB725D81FB8132CE" box="[1029,1115,1431,1457]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2F</figureCitation>
). Male sternite I about twice metacoxae length, sternite I swollen medially (
<figureCitation id="13452A5BEC48FFCCFDDE5DADFCD932AA" box="[681,771,1467,1493]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2D</figureCitation>
), sternite VII sub–rectangular, medial epipleurites reflexed dorsally (
<figureCitation id="13452A5BEC48FFCCFE755DC9FE873286" box="[258,349,1503,1529]" captionStart="FIGURE 2" captionStartId="3.[151,250,1778,1802]" captionTargetBox="[151,1435,417,1755]" captionTargetId="figure-129@3.[151,1435,417,1755]" captionTargetPageId="3" captionText="FIGURE 2. Nosola paradoxa Stål, 1866. A, male specimen in dorsal habitus; B, head, pronotum and part of mesonotum in dorsal view; C, head in ventral view; D, thorax in ventral view; E, right operculum in latero-ventral view; F, timbal cover in latero-ventral view; G, sternites VII and VIII in ventral view. Abbreviations: basisternum 3 (bs3), operculum (op), tergite (t), timbal cover (tc), timbal (tim), sternite (st). Scale bar = A, 1 cm; B–D, G, 2 mm; E, F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491187" httpUri="https://zenodo.org/record/4491187/files/figure.png" pageId="2" pageNumber="511">Fig. 2G</figureCitation>
). Female sternite VII rectangular, wider than long, notch in posterior margin (
<figureCitation id="13452A5BEC48FFCCFBA55DC9FAF13286" box="[1234,1323,1503,1529]" captionStart="FIGURE 4" captionStartId="6.[151,250,1898,1922]" captionTargetBox="[161,1427,309,1875]" captionTargetId="figure-97@6.[161,1427,309,1875]" captionTargetPageId="6" captionText="FIGURE 4. Nosola paradoxa Stål, 1866. A, female specimen in dorsal habitus; B, thorax in ventral view; C, left operculum in latero-ventral view; D, terminalia in lateral view; E, terminalia in ventral view. Abbreviations: basisternum 3 (bs3), dorsal beak (db), gonocoxites (gx), gonapophyses (gy), meracanthus (mc), ovipositor sheath (os), operculum (op), tergite (t). Scale bar = A, 1 cm; B, D, E, 2 mm; C, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4491192" httpUri="https://zenodo.org/record/4491192/files/figure.png" pageId="2" pageNumber="511">Fig. 4E</figureCitation>
). Anterior margin of pygofer distal shoulder terminating in an acute projection, dorsal beak absent, pygofer upper lobe absent, pygofer basal lobe well-developed (
<figureCitation id="13452A5BEC48FFCCFD595E31FD5D313E" box="[558,647,1575,1601]" captionStart="FIGURE 3" captionStartId="5.[151,250,1689,1713]" captionTargetBox="[151,1436,186,1665]" captionTargetId="figure-19@5.[151,1436,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Nosola paradoxa Stål, 1866, male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in lateroventral view; D, phallus in left view; E, phallus in right view. Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), basal plate (bp), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). The arrow indicates the bud-like lobe developed between the dorsal crest and the lateral branches. Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4491189" httpUri="https://zenodo.org/record/4491189/files/figure.png" pageId="2" pageNumber="511">Fig. 3C</figureCitation>
). Uncal dorsal crest developed, projected posteriorly, lateral branches of uncus present, projected anteriorly, ventral apophyses fused with lateral branches of uncus (
<figureCitation id="13452A5BEC48FFCCFBB15E5DFA983119" box="[1222,1346,1611,1638]" captionStart="FIGURE 3" captionStartId="5.[151,250,1689,1713]" captionTargetBox="[151,1436,186,1665]" captionTargetId="figure-19@5.[151,1436,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Nosola paradoxa Stål, 1866, male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in lateroventral view; D, phallus in left view; E, phallus in right view. Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), basal plate (bp), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). The arrow indicates the bud-like lobe developed between the dorsal crest and the lateral branches. Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4491189" httpUri="https://zenodo.org/record/4491189/files/figure.png" pageId="2" pageNumber="511">Fig. 3A, B</figureCitation>
). Theca a simple bent tube with fissure in distal third where vesica originates; thecal and vesical processes absent; vesica extruded, long and slender, ornamented with cornuti and spine of vesica (
<figureCitation id="13452A5BEC48FFCCFCBA5E85FB9C31D1" box="[973,1094,1683,1710]" captionStart="FIGURE 3" captionStartId="5.[151,250,1689,1713]" captionTargetBox="[151,1436,186,1665]" captionTargetId="figure-19@5.[151,1436,181,1665]" captionTargetPageId="5" captionText="FIGURE 3. Nosola paradoxa Stål, 1866, male genitalia. A, uncus in ventral view; B, uncus in lateral view; C, pygofer in lateroventral view; D, phallus in left view; E, phallus in right view. Abbreviations: aedeagus (aed), anal styles (as), basal lobe (bl), basal plate (bp), cornuti (cor), distal shoulder (ds), lateral branches of uncus (lbu), spine of vesica (sv); theca (th), uncal dorsal crest (udc), uncus (un), ventral apophyses (va), vesica (ve). The arrow indicates the bud-like lobe developed between the dorsal crest and the lateral branches. Scale bar = A, B, 0.5 mm; C, 1 mm; D, E, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4491189" httpUri="https://zenodo.org/record/4491189/files/figure.png" pageId="2" pageNumber="511">Fig. 3D, E</figureCitation>
).
</paragraph>
<paragraph id="8BC136DEEC48FFCCFFB05EA1FCA93066" blockId="2.[151,1437,603,2034]" pageId="2" pageNumber="511">
<emphasis id="B90AEACCEC48FFCCFFB05EA1FE1231AE" bold="true" box="[199,456,1719,1745]" pageId="2" pageNumber="511">Measurements (mm).</emphasis>
Length of body: 25.67–29.72; length of fore wing: 32.33–34.84; width of fore wing: 10.27–12.04; length of head: 4.05–4.41; width of head including eyes: 8.64–9.81; width of pronotum including suprahumeral plates: 9.27–11.25; width of mesonotum: 8.1–9.09.
</paragraph>
</subSubSection>
<subSubSection id="C3646555EC48FFCDFFB05F35FDDE36F6" lastPageId="3" lastPageNumber="512" pageId="2" pageNumber="511" type="diagnosis">
<paragraph id="8BC136DEEC48FFCDFFB05F35FDDE36F6" blockId="2.[151,1437,603,2034]" lastBlockId="3.[151,1437,151,393]" lastPageId="3" lastPageNumber="512" pageId="2" pageNumber="511">
<emphasis id="B90AEACCEC48FFCCFFB05F35FD633042" bold="true" box="[199,697,1827,1853]" pageId="2" pageNumber="511">
Diagnosis (extended from
<bibRefCitation id="EFEF4B2FEC48FFCCFE8A5F35FD763042" author="Sanborn" box="[509,684,1827,1853]" firstAuthor="Sanborn" pageId="2" pageNumber="511" pagination="1 - 104" refId="ref10381" refString="Sanborn, A. F. (2019) The cicadas (Hemiptera: Cicadidae) of Bolivia including the descriptions of fifteen new species, the resurrection of one genus and two species, seven new combinations, six new synonymies, and twenty-eight new records. Zootaxa, 4655 (1), 1 - 104. https: // doi. org / 10.11646 / zootaxa. 4655.1.1" type="journal article" year="2019">Sanborn, 2019</bibRefCitation>
).
</emphasis>
The genus can be distinguished from all other genera in
<taxonomicName id="4C7E4D5DEC48FFCCFA585F35FA463042" authorityName="Distant" authorityYear="1905" box="[1327,1436,1827,1853]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="511" phylum="Arthropoda" rank="tribe" tribe="Fidicinini">Fidicinini</taxonomicName>
by the following combination of characters: body with small spines on surface; head including eyes broader than anterior pronotum; supra–antennal plates not meeting eyes; prominent postclypeus extending well beyond anterior margin of head, postclypeus twice the length of the supra–antennal plate in dorsal view, rectangular in ventral view, obtusely angled in lateral view; anteclypeus tumid, posterolateral margins inclined forming an acute angle, posterolateral flaps on anteclypeus absent; anterior basisternum 3 obtusely angled, not prominent relative to mesocoxae; three segmented tarsi; male operculum semilunar; forewings with costal vein strongly arched and expanded at base, expanded proximal costal margin forming an arc bringing the leading edge of the fore wing almost to the apex of the postclypeus; apical cells about one third forewing length; distal shoulder of pygofer hook-like; uncal dorsal crest fused and projected posteriorly; lateral branches of uncus slender and anteriorly projected; ventral apophyses fused with lateral branches of uncus; theca a simple bent tube with a vertical row of cornuti toward the fissure in distal third where vesica originates; thecal and vesical processes absent; vesica extruded, long and slender, ornamented with cornuti and spine of vesica.
</paragraph>
</subSubSection>
</treatment>
</document>
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