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<document id="11EC8212EE2B5F9F1E79C1A3011A425A" ID-CLB-Dataset="24881" ID-DOI="10.11646/zootaxa.4699.1.1" ID-GBIF-Dataset="c374bffb-02ad-4cfd-ab69-fc74b15c27a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="3543038" ID-ZooBank="469CF6FE-D2A3-499F-A9AF-E46B68FBAFD8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573813341551" checkinUser="plazi" docAuthor="Ruan, Yongying, Yang, Xingke, Konstantinov, Alexander S., Prathapan, Kaniyarikkal D. &amp; Zhang, Mengna" docDate="2019" docId="03CA4D00FF22BDDDFF733F27FB1454CE" docLanguage="en" docName="zootaxa.4699.1.1.pdf.imf" docOrigin="Zootaxa 4699 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Chaetocnema (Udorpes) paraumesaoi Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang 2019, New Species" docType="treatment" docVersion="9" lastPageNumber="182" masterDocId="FFF33578FF90BD68FFE43813FF805008" masterDocTitle="Revision of the Oriental Chaetocnema species (Coleoptera, Chrysomelidae, Galerucinae, Alticini)" masterLastPageNumber="206" masterPageNumber="1" pageNumber="179" updateTime="1698767055221" updateUser="ExternalLinkService">
<mods:mods id="C4689573D926C07CBE07C1C48674CE3E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8DAC8E001358D7B4DFB702D6CDE273D9">
<mods:title id="2CCC55DE6DA7AAE60155BFF30A69F736">Revision of the Oriental Chaetocnema species (Coleoptera, Chrysomelidae, Galerucinae, Alticini)</mods:title>
</mods:titleInfo>
<mods:name id="772F8856B30EB88BC8F4B7AB7011A219" type="personal">
<mods:role id="4CEDAB7534775664C5D76908C3BAE6FC">
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<mods:namePart id="D3E719716FF643DACEC1294FA2ABD3D9">Ruan, Yongying</mods:namePart>
</mods:name>
<mods:name id="135DB696FF86204B3E64840E21E2242C" type="personal">
<mods:role id="E5A9562CF1893AE105544723AE58A068">
<mods:roleTerm id="BEF0600BCD17D812E88BB2A01A9796E0">Author</mods:roleTerm>
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<mods:namePart id="C2EB4E14A1F92287F77354BDB573A556">Yang, Xingke</mods:namePart>
</mods:name>
<mods:name id="E944A72CE8C594167700C8BF7198A3D9" type="personal">
<mods:role id="9C771168173FCC7162C9F50DEFC1AFC6">
<mods:roleTerm id="3B612A799FA257FE13BD16ABA2F25D78">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0EC7E5B47CC9136A1985BC8B2537A560">Konstantinov, Alexander S.</mods:namePart>
</mods:name>
<mods:name id="557F073A00520AEC38C5F1695AAB3793" type="personal">
<mods:role id="02FEF4195332EE48CDAD37182FE07EFD">
<mods:roleTerm id="AD847781FC5D74FF87A3A28697E0C3D4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2254D4BCC6D58488FFC5BD9ED678AF6">Prathapan, Kaniyarikkal D.</mods:namePart>
</mods:name>
<mods:name id="46553A77C073DAB6B2F0025702098640" type="personal">
<mods:role id="C7D03C66C9612FB70F5EF81EF55F2820">
<mods:roleTerm id="1CF9DFE7E8A6773326FA65BD657D32DC">Author</mods:roleTerm>
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<mods:namePart id="E1062172BE016C13E5FCDA48A93A6A5F">Zhang, Mengna</mods:namePart>
</mods:name>
<mods:typeOfResource id="6893500B7597566C5E77C8708C20E57A">text</mods:typeOfResource>
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<mods:title id="715893F14AE15FCE6F0A78B04778BE2E">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C963C85724BE2532A384EA21BFE6C0CC">
<mods:date id="E0CA99E86D3F866B25794D63C1ABA54A">2019</mods:date>
<mods:detail id="3AC3213B0EF2FEB8960CA28EE1B9E04D" type="pubDate">
<mods:number id="9897025380D5506784423ECF6B3F1946">2019-11-15</mods:number>
</mods:detail>
<mods:detail id="6D28ABCBEEE7D27EC3F05954F6E9C9FB" type="volume">
<mods:number id="AA8B0A1915C7574CAAA75ACA0828B3D7">4699</mods:number>
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<mods:number id="BE623D7A273875A1E68FE9FCF9D201D9">1</mods:number>
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<mods:classification id="2414022C412149F5C5EA1CD85563756D">journal article</mods:classification>
<mods:identifier id="9F533E2AA39F5BE14083B3F86CF12CA1" type="CLB-Dataset">24881</mods:identifier>
<mods:identifier id="80D674E2CB6A5C4EA0284823C1F1E25D" type="DOI">10.11646/zootaxa.4699.1.1</mods:identifier>
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<mods:identifier id="216222A8EC944B6EED905001C4D995DB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CA4D00FF22BDDDFF733F27FB1454CE" ID-DOI="http://doi.org/10.5281/zenodo.5696751" ID-GBIF-Taxon="160406948" ID-Zenodo-Dep="5696751" LSID="urn:lsid:plazi:treatment:03CA4D00FF22BDDDFF733F27FB1454CE" httpUri="http://treatment.plazi.org/id/03CA4D00FF22BDDDFF733F27FB1454CE" lastPageId="181" lastPageNumber="182" pageId="178" pageNumber="179">
<subSubSection id="C379AF9DFF22BDDAFF733F27FB1F5746" box="[151,1183,1844,1871]" pageId="178" pageNumber="179" type="nomenclature">
<paragraph id="8BDCFC16FF22BDDAFF733F27FB1F5746" blockId="178.[151,1183,1844,1906]" box="[151,1183,1844,1871]" pageId="178" pageNumber="179">
<heading id="D0944B7AFF22BDDAFF733F27FB1F5746" bold="true" box="[151,1183,1844,1871]" fontSize="11" level="1" pageId="178" pageNumber="179" reason="1">
<emphasis id="B9172004FF22BDDAFF733F27FB1F5746" bold="true" box="[151,1183,1844,1871]" pageId="178" pageNumber="179">
75.
<taxonomicName id="4C638795FF22BDDAFF273F27FC785746" authority="Ruan, Konstantinov &amp; Yang" authorityName="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang" authorityYear="2019" box="[195,1016,1844,1871]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="paraumesaoi" status="sp. nov." subGenus="Udorpes">
<emphasis id="B9172004FF22BDDAFF273F27FEE95747" bold="true" box="[195,361,1844,1871]" italics="true" pageId="178" pageNumber="179">Chaetocnema</emphasis>
(
<emphasis id="B9172004FF22BDDAFE9F3F27FE605747" bold="true" box="[379,480,1844,1871]" italics="true" pageId="178" pageNumber="179">Udorpes</emphasis>
)
<emphasis id="B9172004FF22BDDAFE153F27FD0D5746" bold="true" box="[497,653,1844,1870]" italics="true" pageId="178" pageNumber="179">paraumesaoi</emphasis>
Ruan, Konstantinov &amp; Yang
</taxonomicName>
,
<taxonomicNameLabel id="A2249D7FFF22BDDAFBE13F27FB1F5746" box="[1029,1183,1844,1870]" pageId="178" pageNumber="179" rank="species">New Species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C379AF9DFF22BDDAFF733F4BFECA577A" box="[151,330,1879,1906]" pageId="178" pageNumber="179" type="description">
<paragraph id="8BDCFC16FF22BDDAFF733F4BFECA577A" blockId="178.[151,1183,1844,1906]" box="[151,330,1879,1906]" pageId="178" pageNumber="179">
(
<figureCitation id="1358E093FF22BDDAFF7B3F4BFE8B577A" box="[159,267,1880,1906]" captionStart="FIGURE 104" captionStartId="179.[151,250,1836,1861]" captionTargetBox="[180,1435,200,1785]" captionTargetId="figure@179.[151,1436,171,1813]" captionTargetPageId="179" captionText="FIGURE 104. Chaetocnema paraumesaoi sp. nov. Habitus, hand drawing." figureDoi="http://doi.org/10.5281/zenodo.3543268" httpUri="https://zenodo.org/record/3543268/files/figure.png" pageId="178" pageNumber="179">Figs. 104</figureCitation>
,
<figureCitation id="1358E093FF22BDDAFEF33F44FEC2577A" box="[279,322,1879,1906]" captionStart="FIGURE 105" captionStartId="180.[151,250,1852,1877]" captionTargetBox="[166,1436,314,1795]" captionTargetId="figure@180.[151,1436,314,1806]" captionTargetPageId="180" captionText="FIGURE 105. Chaetocnema paraumesaoi sp. nov. A Holotype, habitus. B Pronotum and peri-scutellar area (Holotype). C Head, frontal view (Holotype). D Aedeagus, ventral view (Paratype). E Aedeagus, lateral view (Paratype). F Apex of aedeagus, ventral view (Paratype)." figureDoi="http://doi.org/10.5281/zenodo.3543270" httpUri="https://zenodo.org/record/3543270/files/figure.png" pageId="178" pageNumber="179">105</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C379AF9DFF22BDDAFF733FB3FF4A57D6" pageId="178" pageNumber="179" type="etymology">
<paragraph id="8BDCFC16FF22BDDAFF733FB3FF4A57D6" blockId="178.[151,1436,1952,2014]" pageId="178" pageNumber="179">
<emphasis id="B9172004FF22BDDAFF733FB3FEA257B2" bold="true" box="[151,290,1952,1978]" pageId="178" pageNumber="179">Etymology:</emphasis>
This new species is named after its resemblance to
<emphasis id="B9172004FF22BDDAFC9D3FB2FC7757B3" box="[889,1015,1953,1979]" italics="true" pageId="178" pageNumber="179">
<taxonomicName id="4C638795FF22BDDAFC9D3FB2FC7357B3" box="[889,1011,1953,1979]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="178" pageNumber="179" phylum="Arthropoda" rank="species" species="umesaoi">C.umesaoi</taxonomicName>
.
</emphasis>
Specific epithet is a noun in apposition.
</paragraph>
</subSubSection>
<caption id="DF1CAC9EFF23BDDBFF733F3FFC1B574D" ID-DOI="http://doi.org/10.5281/zenodo.3543268" ID-Zenodo-Dep="3543268" box="[151,923,1836,1861]" httpUri="https://zenodo.org/record/3543268/files/figure.png" pageId="179" pageNumber="180" startId="179.[151,250,1836,1861]" targetBox="[180,1435,200,1785]" targetPageId="179">
<paragraph id="8BDCFC16FF23BDDBFF733F3FFC1B574D" blockId="179.[151,923,1836,1861]" box="[151,923,1836,1861]" pageId="179" pageNumber="180">
<emphasis id="B9172004FF23BDDBFF733F3FFD27574C" bold="true" box="[151,679,1836,1861]" pageId="179" pageNumber="180">
FIGURE 104.
<taxonomicName id="4C638795FF23BDDBFED73F3FFDD0574C" authority="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang, 2019" authorityName="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang" authorityYear="2019" box="[307,592,1836,1861]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="paraumesaoi" status="sp. nov.">
<emphasis id="B9172004FF23BDDBFED73F3FFDD0574C" bold="true" box="[307,592,1836,1861]" italics="true" pageId="179" pageNumber="180">Chaetocnema paraumesaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2249D7FFF23BDDBFDB23F3FFD27574C" box="[598,679,1836,1860]" pageId="179" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Habitus, hand drawing.
</paragraph>
</caption>
<subSubSection id="C379AF9DFF23BDDCFF233F85FB0C50DF" lastPageId="180" lastPageNumber="181" pageId="179" pageNumber="180" type="diagnosis">
<paragraph id="8BDCFC16FF23BDDCFF233F85FB0C50DF" blockId="179.[151,1436,1942,2041]" lastBlockId="180.[151,1436,153,287]" lastPageId="180" lastPageNumber="181" pageId="179" pageNumber="180">
<emphasis id="B9172004FF23BDDBFF233F85FEC457B9" bold="true" box="[199,324,1942,1969]" pageId="179" pageNumber="180">Diagnosis:</emphasis>
The new species strongly resembles
<taxonomicName id="4C638795FF23BDDBFD0C3F84FCE757B9" authority="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang, 2019" authorityName="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang" authorityYear="2019" box="[744,871,1943,1969]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="179" pageNumber="180" phylum="Arthropoda" rank="species" species="umesaoi" status="sp. nov.">
<emphasis id="B9172004FF23BDDBFD0C3F84FCE757B9" box="[744,871,1943,1969]" italics="true" pageId="179" pageNumber="180">C. umesaoi</emphasis>
</taxonomicName>
<emphasis id="B9172004FF23BDDBFC8B3F84FC4957B9" bold="true" box="[879,969,1943,1969]" pageId="179" pageNumber="180">
<taxonomicNameLabel id="A2249D7FFF23BDDBFC8B3F84FC4957B9" box="[879,969,1943,1969]" pageId="179" pageNumber="180" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
However, it differs from the latter by the following characters: smaller body size; peri-scutellar area of elytron is wider with more punctures; pronotal base obviously expanded and convex at middle; body strongly convex at sides, posterior end of body strongly narrowed and pointed; vertex with denser punctures, frontal ridge slightly wider. The new species is also close to
<taxonomicName id="4C638795FF24BDDCFAF4388AFF3550DF" authorityName="Jacoby" authorityYear="1896" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="modiglianii">
<emphasis id="B9172004FF24BDDCFAF4388AFF3550DF" italics="true" pageId="180" pageNumber="181">C. modiglianii</emphasis>
</taxonomicName>
on external characters, however, they can be easily separated by the shape of aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C379AF9DFF24BDDCFF2338F3FDA050F3" box="[199,544,224,251]" pageId="180" pageNumber="181" type="distribution">
<paragraph id="8BDCFC16FF24BDDCFF2338F3FDA050F3" blockId="180.[151,1436,153,287]" box="[199,544,224,251]" pageId="180" pageNumber="181">
<emphasis id="B9172004FF24BDDCFF2338F3FEE350F3" bold="true" box="[199,355,224,251]" pageId="180" pageNumber="181">Distribution:</emphasis>
<collectingCountry id="F374BC86FF24BDDCFE8E38F2FE5350F3" box="[362,467,225,251]" name="Malaysia" pageId="180" pageNumber="181">Malaysia</collectingCountry>
,
<collectingCountry id="F374BC86FF24BDDCFE3A38F2FD9B50F3" box="[478,539,225,251]" name="India" pageId="180" pageNumber="181">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C379AF9DFF24BDDCFF233916FE565117" box="[199,470,261,287]" pageId="180" pageNumber="181" type="biology_ecology">
<paragraph id="8BDCFC16FF24BDDCFF233916FE565117" blockId="180.[151,1436,153,287]" box="[199,470,261,287]" pageId="180" pageNumber="181">
<emphasis id="B9172004FF24BDDCFF233916FED95117" bold="true" box="[199,345,261,287]" pageId="180" pageNumber="181">
<collectingRegion id="49A732F4FF24BDDCFF233916FF7F5117" box="[199,255,261,287]" country="Afghanistan" name="Khost" pageId="180" pageNumber="181">Host</collectingRegion>
plants:
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<caption id="DF1CAC9EFF24BDDCFF733F2FFE0C5795" ID-DOI="http://doi.org/10.5281/zenodo.3543270" ID-Zenodo-Dep="3543270" httpUri="https://zenodo.org/record/3543270/files/figure.png" pageId="180" pageNumber="181" startId="180.[151,250,1852,1877]" targetBox="[166,1436,314,1795]" targetPageId="180">
<paragraph id="8BDCFC16FF24BDDCFF733F2FFE0C5795" blockId="180.[151,1436,1852,1949]" pageId="180" pageNumber="181">
<emphasis id="B9172004FF24BDDCFF733F2FFD4C575D" bold="true" box="[151,716,1852,1877]" pageId="180" pageNumber="181">
FIGURE 105.
<taxonomicName id="4C638795FF24BDDCFED33F2FFDD6575D" authority="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang, 2019" authorityName="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang" authorityYear="2019" box="[311,598,1852,1877]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="180" pageNumber="181" phylum="Arthropoda" rank="species" species="paraumesaoi" status="sp. nov.">
<emphasis id="B9172004FF24BDDCFED33F2FFDD6575D" bold="true" box="[311,598,1852,1877]" italics="true" pageId="180" pageNumber="181">Chaetocnema paraumesaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2249D7FFF24BDDCFDBA3F2EFD31575D" box="[606,689,1853,1877]" pageId="180" pageNumber="181" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
Holotype, habitus.
<emphasis id="B9172004FF24BDDCFC7D3F2EFC2A575D" bold="true" box="[921,938,1853,1877]" pageId="180" pageNumber="181">B</emphasis>
Pronotum and peri-scutellar area (Holotype).
<emphasis id="B9172004FF24BDDCFA6E3F2FFA1C575D" bold="true" box="[1418,1436,1852,1877]" pageId="180" pageNumber="181">C</emphasis>
Head, frontal view (Holotype).
<emphasis id="B9172004FF24BDDCFE313F72FE675771" bold="true" box="[469,487,1889,1913]" pageId="180" pageNumber="181">D</emphasis>
Aedeagus, ventral view (Paratype).
<emphasis id="B9172004FF24BDDCFCB13F72FCE65771" bold="true" box="[853,870,1889,1913]" pageId="180" pageNumber="181">E</emphasis>
Aedeagus, lateral view (Paratype).
<emphasis id="B9172004FF24BDDCFB293F72FB5C5771" bold="true" box="[1229,1244,1889,1913]" pageId="180" pageNumber="181">F</emphasis>
Apex of aedeagus, ventral view (Paratype).
</paragraph>
</caption>
<subSubSection id="C379AF9DFF25BDDDFF23388BFB1454CE" pageId="181" pageNumber="182" type="description">
<paragraph id="8BDCFC16FF25BDDDFF23388BFAFC5117" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">
<emphasis id="B9172004FF25BDDDFF23388BFEDB50BB" bold="true" box="[199,347,152,179]" pageId="181" pageNumber="182">Description:</emphasis>
Body length
<quantity id="4C9B51F3FF25BDDDFE17388BFD1950BB" box="[499,665,152,179]" metricMagnitude="-3" metricUnit="m" metricValue="1.575" metricValueMax="1.65" metricValueMin="1.5" pageId="181" pageNumber="182" unit="mm" value="1.575" valueMax="1.65" valueMin="1.5">1.50–1.65 mm</quantity>
, body width
<quantity id="4C9B51F3FF25BDDDFCCF388BFC5150BB" box="[811,977,152,179]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.0" metricValueMin="8.5" pageId="181" pageNumber="182" unit="mm" value="0.875" valueMax="0.9" valueMin="0.85">0.85–0.90 mm</quantity>
. Ratio of elytron length (along suture) to width (maximum): 2.20–2.25. Ratio of pronotum width (at base) to length: 1.50–1.55. Ratio of length of elytron to length of pronotum (along middle): 2.15–2.25. Ratio of width of elytra at base to width of pronotum at base (in middle of humeral calli): 1.05–1.10. Ratio of maximum width of elytra to maximum width of pronotum: 1.30–1.40.
</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF23393AFDF75182" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">Dorsum coppery, rarely bronzy. Antennae slender, slightly reach beyond humeral callus. Antennomeres 1–5 more or less yellow brown, 6–11 brown, distal ones gradually turning darker. Tibiae and tarsi brown. Pro- and mesofemora brown. Metafemora deep brown.</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF233987FF655237" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">Head hypognathous. Frontal ridge wide and flat, with numerous closely placed punctures. Frontolateral sulcus absent. Suprafrontal sulcus present, moderately developed. Ratio of width of frontal ridge to width of antennal socket (counting surrounding ridges): 1.95–2.10. Frons evenly covered with punctures. Vertex flat, situated on same level as orbit. Vertex evenly and closely covered with punctures, punctures more or less sparser in middle of vertex.</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF233A5AFDD452FB" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">Base of pronotum without short longitudinal impressions. Deep row of large punctures at base of pronotum absent. Pronotal base strongly convex at middle.Area adjacent to mid basal margin of pronotum covered with punctures. Pronotum without shallow antebasal transverse impression. Lateral margins of pronotum strongly convex. Anterolateral and posterolateral prothoracic callosities poorly developed. Diameter of pronotal punctures subequal to distance between adjacent punctures.</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF233AEFFB8F5332" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">Elytra strongly converging towards apex, with strongly convex lateral sides. Scutellar row of punctures tripled. Remaining rows of elytral punctures regular. Humeral callus poorly developed.</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF233B57FCDF538A" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">First male protarsomeres and mesotarsomeres strongly enlarged, obviously larger than those in females. Large lateral denticle on metatibia obtuse. Metatibial serration absent.</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF233B9EFEEC53E6" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">Apex of aedeagus narrowing abruptly in ventral view. Ventral longitudinal groove present. In ventral view, apical denticle not well developed. Minute transverse wrinkles on ventral side absent.Aedeagus in lateral view strongly and evenly curved.</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF233BEAFAD9543E" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">
<emphasis id="B9172004FF25BDDDFF233BEAFE96541B" bold="true" box="[199,278,1017,1043]" pageId="181" pageNumber="182">Types:</emphasis>
<materialsCitation id="3B0BF64BFF25BDDDFEC43BEBFDD0543F" ID-GBIF-Occurrence="2451575832" collectingDate="1983-09-07" collectionCode="USNM" collectorName="G. F. Hevel &amp; W. E. Steiner" country="Malaysia" elevation="1560" location="Headquarters area" municipality="Kinabalu National Park" pageId="181" pageNumber="182" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54D842B4FF25BDDDFEC43BEBFE0D541A" box="[288,397,1016,1042]" pageId="181" pageNumber="182" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D65379FFF25BDDDFE7E3BEAFE2F541B" box="[410,431,1017,1043]" pageId="181" pageNumber="182" type="male">♂</specimenCount>
(
<collectionCode id="ED7264D3FF25BDDDFE253BEAFD99541B" box="[449,537,1017,1043]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
): 1)
<collectingCountry id="F374BC86FF25BDDDFDAB3BEBFD38541A" box="[591,696,1016,1042]" name="Malaysia" pageId="181" pageNumber="182">Malaysia</collectingCountry>
,
<collectingRegion id="49A732F4FF25BDDDFD233BEBFC8E541A" box="[711,782,1016,1042]" country="Malaysia" name="Sabah" pageId="181" pageNumber="182">Sabah</collectingRegion>
,
<collectingMunicipality id="6BB8666CFF25BDDDFCF83BEBFBAD541A" box="[796,1069,1016,1042]" pageId="181" pageNumber="182">Kinabalu National Park</collectingMunicipality>
,
<location id="8EBCAACDFF25BDDDFBD83BEBFA8C541B" LSID="urn:lsid:plazi:treatment:03CA4D00FF22BDDDFF733F27FB1454CE:8EBCAACDFF25BDDDFBD83BEBFA8C541B" box="[1084,1292,1016,1043]" country="Malaysia" municipality="Kinabalu National Park" name="Headquarters area" pageId="181" pageNumber="182" stateProvince="Sabah">Headquarters area</location>
, alt.
<quantity id="4C9B51F3FF25BDDDFAA33BEBFA18541B" box="[1351,1432,1016,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="181" pageNumber="182" unit="m" value="1560.0">
<elevation id="004E1B25FF25BDDDFAA33BEBFA18541B" box="[1351,1432,1016,1043]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="181" pageNumber="182" unit="m" value="1560.0">1560m</elevation>
</quantity>
,
<date id="FFDDDAD6FF25BDDDFF733C0EFE8C543F" box="[151,268,1053,1079]" pageId="181" pageNumber="182" value="1983-09-07">
<collectingDate id="EF99233EFF25BDDDFF733C0EFE8C543F" box="[151,268,1053,1079]" pageId="181" pageNumber="182" value="1983-09-07">7.IX.1983</collectingDate>
</date>
,
<collectorName id="269699C0FF25BDDDFEF33C0FFE0A543E" box="[279,394,1052,1078]" pageId="181" pageNumber="182">G.F.Hevel</collectorName>
&amp;
<collectorName id="269699C0FF25BDDDFE483C0FFDB6543E" box="[428,566,1052,1078]" pageId="181" pageNumber="182">W.E.Steiner</collectorName>
; 2
</materialsCitation>
)
<typeStatus id="54D842B4FF25BDDDFD843C0FFD4C543E" box="[608,716,1052,1078]" pageId="181" pageNumber="182" type="holotype">Holotype</typeStatus>
; 3)
<taxonomicName id="4C638795FF25BDDDFD113C0FFBA8543E" authority="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang, 2019" authorityName="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang" authorityYear="2019" box="[757,1064,1052,1078]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="paraumesaoi" status="sp. nov.">
<emphasis id="B9172004FF25BDDDFD113C0FFBA8543E" box="[757,1064,1052,1078]" italics="true" pageId="181" pageNumber="182">Chaetocnema paraumesaoi</emphasis>
</taxonomicName>
<emphasis id="B9172004FF25BDDDFBCB3C0EFB09543F" bold="true" box="[1071,1161,1053,1079]" pageId="181" pageNumber="182">
<taxonomicNameLabel id="A2249D7FFF25BDDDFBCB3C0EFB09543F" box="[1071,1161,1053,1079]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Des. Ruan, 2016.
</paragraph>
<paragraph id="8BDCFC16FF25BDDDFF233C52FB1454CE" blockId="181.[151,1437,152,1223]" pageId="181" pageNumber="182">
<materialsCitation id="3B0BF64BFF25BDDDFF233C52FC0A5476" ID-GBIF-Occurrence="2451575822" collectingDate="2003-11-14" collectionCode="USNM" collectorName="Konstantinov &amp; Prathapan &amp; Saluk" country="India" county="Mudigere" latitude="13.1325" location="South" longLatPrecision="21" longitude="75.54472" municipality="Western Ghats" pageId="181" pageNumber="182" specimenCount="14" specimenCount-female="10" specimenCount-male="4" typeStatus="paratype">
<typeStatus id="54D842B4FF25BDDDFF233C52FEB85453" box="[199,312,1089,1115]" pageId="181" pageNumber="182" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D65379FFF25BDDDFEA13C52FEEB5453" box="[325,363,1089,1115]" pageId="181" pageNumber="182" type="male">4♂</specimenCount>
<specimenCount id="9D65379FFF25BDDDFE8F3C52FE185453" box="[363,408,1089,1115]" pageId="181" pageNumber="182" type="female">10♀</specimenCount>
(
<collectionCode id="ED7264D3FF25BDDDFE4D3C52FD805453" box="[425,512,1089,1115]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
),
<location id="8EBCAACDFF25BDDDFDF03C53FDD55452" LSID="urn:lsid:plazi:treatment:03CA4D00FF22BDDDFF733F27FB1454CE:8EBCAACDFF25BDDDFDF03C53FDD55452" box="[532,597,1088,1114]" country="India" county="Mudigere" latitude="13.1325" longLatPrecision="21" longitude="75.54472" municipality="Western Ghats" name="South" pageId="181" pageNumber="182">South</location>
<collectingCountry id="F374BC86FF25BDDDFDBA3C53FD1A5452" box="[606,666,1088,1114]" name="India" pageId="181" pageNumber="182">India</collectingCountry>
,
<collectingMunicipality id="6BB8666CFF25BDDDFD433C52FCCC5452" box="[679,844,1088,1115]" pageId="181" pageNumber="182">Western Ghats</collectingMunicipality>
<taxonomicName id="4C638795FF25BDDDFCB13C53FB875453" authority="Prov" authorityName="Prov" box="[853,1031,1088,1115]" class="Magnoliopsida" family="Apiaceae" genus="Karnataka" kingdom="Plantae" order="Apiales" pageId="181" pageNumber="182" phylum="Tracheophyta" rank="genus">Karnataka Prov</taxonomicName>
, env.
<collectingCounty id="62BD849AFF25BDDDFBAE3C53FB3A5452" box="[1098,1210,1088,1114]" pageId="181" pageNumber="182">Mudigere</collectingCounty>
,
<date id="FFDDDAD6FF25BDDDFB223C52FAC85453" box="[1222,1352,1089,1115]" pageId="181" pageNumber="182" value="2003-11-14">
<collectingDate id="EF99233EFF25BDDDFB223C52FAC85453" box="[1222,1352,1089,1115]" pageId="181" pageNumber="182" value="2003-11-14">
14.XI.
<quantity id="4C9B51F3FF25BDDDFAEE3C52FAC85453" box="[1290,1352,1089,1115]" metricMagnitude="6" metricUnit="m" metricValue="2.003904" pageId="181" pageNumber="182" unit="m" value="2003904.0">2003</quantity>
</collectingDate>
</date>
, 904m, river,
<geoCoordinate id="EE579AD1FF25BDDDFF3C3C77FEDF5477" box="[216,351,1124,1151]" degrees="13" direction="north" minutes="07" orientation="latitude" pageId="181" pageNumber="182" precision="15" seconds="57" value="13.1325">N13°07'57&quot;</geoCoordinate>
,
<geoCoordinate id="EE579AD1FF25BDDDFE8E3C77FE6D5477" box="[362,493,1124,1151]" degrees="75" direction="east" minutes="32" orientation="longitude" pageId="181" pageNumber="182" precision="15" seconds="41" value="75.54472">E75°32'41&quot;</geoCoordinate>
, leg.
<collectorName id="269699C0FF25BDDDFDC33C77FD3F5476" box="[551,703,1124,1150]" pageId="181" pageNumber="182">Konstantinov</collectorName>
,
<collectorName id="269699C0FF25BDDDFD2E3C77FCBD5476" box="[714,829,1124,1150]" pageId="181" pageNumber="182">Prathapan</collectorName>
,
<collectorName id="269699C0FF25BDDDFCA33C77FC0A5476" box="[839,906,1124,1150]" pageId="181" pageNumber="182">Saluk</collectorName>
</materialsCitation>
,
<materialsCitation id="3B0BF64BFF25BDDDFC703C76FE3554CE" ID-GBIF-Occurrence="2451575877" collectingDate="1983-09-07" collectionCode="USNM" collectorName="G. F. Hevel &amp; W. E. Steiner" country="Malaysia" elevation="1560" location="Headquarters area" municipality="Kinabalu National Park" pageId="181" pageNumber="182" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="54D842B4FF25BDDDFC703C76FC795477" box="[916,1017,1125,1151]" pageId="181" pageNumber="182" type="paratype">Paratype</typeStatus>
,
<taxonomicName id="4C638795FF25BDDDFBE73C77FAB65476" authority="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang, 2019" authorityName="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang" authorityYear="2019" box="[1027,1334,1124,1150]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="paraumesaoi" status="sp. nov.">
<emphasis id="B9172004FF25BDDDFBE73C77FAB65476" box="[1027,1334,1124,1150]" italics="true" pageId="181" pageNumber="182">Chaetocnema paraumesaoi</emphasis>
</taxonomicName>
<emphasis id="B9172004FF25BDDDFAD83C76FA155477" bold="true" box="[1340,1429,1125,1151]" pageId="181" pageNumber="182">
<taxonomicNameLabel id="A2249D7FFF25BDDDFAD83C76FA155477" box="[1340,1429,1125,1151]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Des. Ruan, 2016.
<specimenCount id="9D65379FFF25BDDDFEB83C9AFEFF54AB" box="[348,383,1161,1187]" pageId="181" pageNumber="182" type="male">1♂</specimenCount>
(
<collectionCode id="ED7264D3FF25BDDDFE683C9AFE6354AB" box="[396,483,1161,1187]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="181" pageNumber="182" type="Museum">USNM</collectionCode>
),
<collectingCountry id="F374BC86FF25BDDDFE103C9BFDDD54AA" box="[500,605,1160,1186]" name="Malaysia" pageId="181" pageNumber="182">Malaysia</collectingCountry>
,
<collectingRegion id="49A732F4FF25BDDDFD8C3C9BFD2F54AA" box="[616,687,1160,1186]" country="Malaysia" name="Sabah" pageId="181" pageNumber="182">Sabah</collectingRegion>
,
<collectingMunicipality id="6BB8666CFF25BDDDFD5D3C9BFC4154AA" box="[697,961,1160,1186]" pageId="181" pageNumber="182">Kinabalu National Park</collectingMunicipality>
,
<location id="8EBCAACDFF25BDDDFC2F3C9BFB1654AB" LSID="urn:lsid:plazi:treatment:03CA4D00FF22BDDDFF733F27FB1454CE:8EBCAACDFF25BDDDFC2F3C9BFB1654AB" box="[971,1174,1160,1187]" country="Malaysia" municipality="Kinabalu National Park" name="Headquarters area" pageId="181" pageNumber="182" stateProvince="Sabah">Headquarters area</location>
, alt.
<quantity id="4C9B51F3FF25BDDDFB2D3C9BFA9954AB" box="[1225,1305,1160,1187]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="181" pageNumber="182" unit="m" value="1560.0">
<elevation id="004E1B25FF25BDDDFB2D3C9BFA9954AB" box="[1225,1305,1160,1187]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="181" pageNumber="182" unit="m" value="1560.0">1560m</elevation>
</quantity>
,
<date id="FFDDDAD6FF25BDDDFAC03C9AFA1854AB" box="[1316,1432,1161,1187]" pageId="181" pageNumber="182" value="1983-09-07">
<collectingDate id="EF99233EFF25BDDDFAC03C9AFA1854AB" box="[1316,1432,1161,1187]" pageId="181" pageNumber="182" value="1983-09-07">7.IX.1983</collectingDate>
</date>
,
<collectorName id="269699C0FF25BDDDFF733CBFFE8A54CE" box="[151,266,1196,1222]" pageId="181" pageNumber="182">G.F.Hevel</collectorName>
&amp;
<collectorName id="269699C0FF25BDDDFEC93CBFFE3554CE" box="[301,437,1196,1222]" pageId="181" pageNumber="182">W.E.Steiner</collectorName>
</materialsCitation>
,
<typeStatus id="54D842B4FF25BDDDFE243CBEFDA554CF" box="[448,549,1197,1223]" pageId="181" pageNumber="182" type="paratype">Paratype</typeStatus>
,
<taxonomicName id="4C638795FF25BDDDFDD43CBFFCE354CE" authority="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang, 2019" authorityName="Ruan &amp; Yang &amp; Konstantinov &amp; Prathapan &amp; Zhang" authorityYear="2019" box="[560,867,1196,1222]" class="Insecta" family="Chrysomelidae" genus="Chaetocnema" kingdom="Animalia" order="Coleoptera" pageId="181" pageNumber="182" phylum="Arthropoda" rank="species" species="paraumesaoi" status="sp. nov.">
<emphasis id="B9172004FF25BDDDFDD43CBFFCE354CE" box="[560,867,1196,1222]" italics="true" pageId="181" pageNumber="182">Chaetocnema paraumesaoi</emphasis>
</taxonomicName>
<emphasis id="B9172004FF25BDDDFC8E3CBEFC4454CF" bold="true" box="[874,964,1197,1223]" pageId="181" pageNumber="182">
<taxonomicNameLabel id="A2249D7FFF25BDDDFC8E3CBEFC4454CF" box="[874,964,1197,1223]" pageId="181" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Des. Ruan, 2016.
</paragraph>
</subSubSection>
</treatment>
</document>
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