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<mods:title id="C9423FC84295E0A4690E937C40F7C4F1">Review of the genus Ceraturgus Wiedemann (Diptera: Asilidae) in North America north of Mexico</mods:title>
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<paragraph id="8B88BA5BFFE3FFF9FF51FF61FDDAA889" blockId="4.[151,530,151,177]" box="[151,530,151,177]" pageId="4" pageNumber="5">
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<taxonomicName id="4C37C1D8FFE3FFF9FF51FF61FDDAA889" ID-CoL="3KXZ" ID-ENA="468703" authority="Wiedemann (1824)" authorityName="Wiedemann" authorityYear="1824" box="[151,530,151,177]" class="Insecta" family="Asilidae" genus="Ceraturgus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE3FFF9FF51FF61FDDAA889" bold="true" box="[151,530,151,177]" pageId="4" pageNumber="5">
<emphasis id="B9436649FFE3FFF9FF51FF61FEE8A889" bold="true" box="[151,288,151,177]" italics="true" pageId="4" pageNumber="5">Ceraturgus</emphasis>
<bibRefCitation id="EFA6C7AAFFE3FFF9FEE0FF61FDDAA889" author="Wiedemann" box="[294,530,151,177]" pageId="4" pageNumber="45" refString="Wiedemann, C. R. W. (1824) Munus rectoris in Academia Christiana Albertina aditurus analecta entomologica ex Museo Regio Havniensi. Kiliae, 60 pp." type="book" year="1824">Wiedemann (1824)</bibRefCitation>
</emphasis>
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<taxonomicName id="4C37C1D8FFE3FFF9FF51FF14FDCCA8C1" ID-CoL="3KXZ" authority="Wiedemann, 1824: 12" authorityName="Wiedemann" authorityPageNumber="12" authorityYear="1824" box="[151,516,226,249]" class="Insecta" family="Asilidae" genus="Ceraturgus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE3FFF9FF51FF14FED8A8C1" box="[151,272,226,249]" italics="true" pageId="4" pageNumber="5">Ceraturgus</emphasis>
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<bibRefCitation id="EFA6C7AAFFE3FFF9FED3FF14FE14A8C1" author="Wiedemann" box="[277,476,226,249]" pageId="4" pageNumber="45" refString="Wiedemann, C. R. W. (1824) Munus rectoris in Academia Christiana Albertina aditurus analecta entomologica ex Museo Regio Havniensi. Kiliae, 60 pp." type="book" year="1824">Wiedemann, 1824</bibRefCitation>
: 12
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</taxonomicName>
.
<typeStatus id="548C04F9FFE3FFF9FDC8FF14FD8BA8C1" box="[526,579,226,249]" pageId="4" pageNumber="5">Type</typeStatus>
species:
<taxonomicName id="4C37C1D8FFE3FFF9FD63FF14FBCCA8C1" ID-CoL="884KV" authority="Fabricius" authorityName="Fabricius" box="[677,1028,226,249]" class="Insecta" family="Asilidae" genus="Dasypogon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aurulentus">
<emphasis id="B9436649FFE3FFF9FD63FF14FC5FA8C1" box="[677,919,226,249]" italics="true" pageId="4" pageNumber="5">Dasypogon aurulentus</emphasis>
Fabricius
</taxonomicName>
, by monotypy.
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</paragraph>
<paragraph id="8B88BA5BFFE3FFF9FF51FEDCFD3DAC85" blockId="4.[151,1437,298,2016]" pageId="4" pageNumber="5">
Body length, excluding antennae, less than
<quantity id="4CCF17BEFFE3FFF9FD5EFEDCFD38A97C" box="[664,752,298,324]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="4" pageNumber="5" unit="mm" value="28.0">28 mm</quantity>
. Head width 1.51.6 times eye height; face in profile produced beyond eye margin, lacking pronounced tentorial pits; mystax occupying lower half or more of face; sides of frons more or less parallel sided, not strongly converging or diverging dorsally; compound eye with anterior ommatidia greatly enlarged; ocellar tubercle low, bearing several stout setae and shorter hair-like setae; occipital pile dense. Proboscis laterally compressed, apically blunt; palpus 2-segmented. Antennae exceptionally long, much longer than head, with 5 antennomeres, including 3 elongate flagellomeres; scape and pedicel subequal; first flagellomere less than twice length of scape and pedicel combined; second flagellomere short, about half length of scape; antennal bases on raised area. Thorax with scutum typically vittate, bearing sparse or dense pile of short or long, erect or reclinate, hair-like setae shorter or longer than scape; anepisternum and katepisternum bearing many long or short setae; katatergite bearing fan of long setae; anatergite tomentose, lacking long setae; scutellem low convex, bearing band of long marginal bristles; postmetacoxal area membranous. Femora stout but not swollen, usually with apical circlet of stout setae; fore and mid femora with 05 stout anterior setae; hind femur usually with anterior row of 2-8 stout setae. Fore tibia with all apical bristles straight, usually with anterodorsal and posterodorsal rows of short, stout setae and posteroventral row of long, stout setae; mid tibia with apical ventral bristles directed distally, not toward body, usually with anterodorsal, anteroventral, posterodorsal, and posteroventral rows of stout setae; hind tibia with anterior, dorsal, and ventral rows of stout setae; pulvilli present. Wing hyaline to moderately infuscated; vein C continuing around posterior wing margin to wing base; R2+3 ending in C, its apex not directed sharply forward before ending in C; R4 not unusually arched and sinuate; apical portion of M3 at an angle to crossvein mm; CuA2 and A1 ending separately; cell m3 open at wing margin; cell cu
<emphasis id="B9436649FFE3FFF9FC3BFBDFFBC4AC7A" box="[1021,1036,1065,1090]" italics="true" pageId="4" pageNumber="5">p</emphasis>
closed or narrowly open at wing margin. Prosternum not fused to proepisternum. Abdomen usually black with conspicuous white to yellow pollinose posterior fascia; segment 2 not unusually elongate; gonocoxite often bilobed, bearing tuft of long, stout setae; tergite 10 with strong bristles in female.
</paragraph>
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<emphasis id="B9436649FFE3FFF9FF00FB3DFE82ACDD" bold="true" box="[198,330,1227,1253]" pageId="4" pageNumber="5">Discussion</emphasis>
.
<bibRefCitation id="EFA6C7AAFFE3FFF9FE9CFB3DFDF6ACDD" author="Wiedemann" box="[346,574,1227,1253]" pageId="4" pageNumber="45" refString="Wiedemann, C. R. W. (1824) Munus rectoris in Academia Christiana Albertina aditurus analecta entomologica ex Museo Regio Havniensi. Kiliae, 60 pp." type="book" year="1824">Wiedemann (1824)</bibRefCitation>
described the genus
<taxonomicName id="4C37C1D8FFE3FFF9FCFAFB3DFC09ACDC" box="[828,961,1227,1252]" class="Insecta" family="Asilidae" genus="Ceraturgus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE3FFF9FCFAFB3DFC09ACDC" box="[828,961,1227,1252]" italics="true" pageId="4" pageNumber="5">Ceraturgus</emphasis>
</taxonomicName>
to receive the single distinctive species
<taxonomicName id="4C37C1D8FFE3FFF9FF51FB05FDD0AD35" authority="Fabricius" authorityName="Fabricius" box="[151,536,1267,1293]" class="Insecta" family="Asilidae" genus="Dasypogon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="aurulentus">
<emphasis id="B9436649FFE3FFF9FF51FB05FE68AD34" box="[151,416,1267,1292]" italics="true" pageId="4" pageNumber="5">Dasypogon aurulentus</emphasis>
Fabricius
</taxonomicName>
, known at that time from a single specimen, “Habitat in New-York Americes Dom. de Sehestedt” (
<bibRefCitation id="EFA6C7AAFFE3FFF9FE5AFAEDFD9DAD0D" author="Fabricius" box="[412,597,1307,1333]" pageId="4" pageNumber="44" refString="Fabricius, J. C. (1805) Systema antiatorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Brunsvigae, 373 pp." type="book" year="1805">Fabricius 1805</bibRefCitation>
). His brief description distinguished the genus primarily on antennal characters: antennae porrect, long, 5-segmented: segments 13 subclavate, segment 4 cup-shaped, 5 with obtuse apex; body naked.
</paragraph>
</subSubSection>
<subSubSection id="C32DE9D0FFE3FFF8FF00FA65FB2AA912" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="reference_group">
<paragraph id="8B88BA5BFFE3FFF9FF00FA65FB54AE98" blockId="4.[151,1437,298,2016]" pageId="4" pageNumber="5">
The
<taxonomicName id="4C37C1D8FFE3FFF9FF3BFA65FE97AD95" ID-CoL="6SY" box="[253,351,1427,1453]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Asilidae</taxonomicName>
have not yet been subjected to thorough phylogenetic analysis, although work is presently underway. Higher classification within the family remains unstable.
<bibRefCitation id="EFA6C7AAFFE3FFF9FC0FFA4DFAE6ADED" author="Artigas" box="[969,1326,1467,1493]" pageId="4" pageNumber="44" refString="Artigas, J. N. &amp; Papavero, N. (1991 a) The American genera of Asilidae (Diptera): keys for identification with an atlas of female spermathecae and other morphological details. VII. 1. Subfamily Stenopogoninae Hull. A preliminary classification into tribes. Gayana Zoologia, 55 (2), 139 - 144." type="journal article" year="1991" yearSuffix="a">Artigas and Papavero (1991a)</bibRefCitation>
included the genera
<taxonomicName id="4C37C1D8FFE3FFF9FEDEFA15FE54ADC4" ID-CoL="3KXZ" box="[280,412,1507,1532]" class="Insecta" family="Asilidae" genus="Ceraturgus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE3FFF9FEDEFA15FE54ADC4" box="[280,412,1507,1532]" italics="true" pageId="4" pageNumber="5">Ceraturgus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C37C1D8FFE3FFF9FE10FA15FD9FADC4" ID-CoL="5W8C" box="[470,599,1507,1532]" class="Insecta" family="Asilidae" genus="Myelaphus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE3FFF9FE10FA15FD9FADC4" box="[470,599,1507,1532]" italics="true" pageId="4" pageNumber="5">Myelaphus</emphasis>
</taxonomicName>
in their new tribe Ceraturgini of the subfamily
<taxonomicName id="4C37C1D8FFE3FFF9FB43FA15FA8DADC5" ID-CoL="89P4B" box="[1157,1349,1507,1533]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Stenopogoninae">Stenopogoninae</taxonomicName>
.
<taxonomicName id="4C37C1D8FFE3FFF9FA96FA15FED9AE1D" ID-CoL="89P4B" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Stenopogoninae">Stenopogoninae</taxonomicName>
are recognized by the following combination of characters: prothoracic tibia lacking a differentiated apical spur (stouter than others, or twisted and sigmoid), vein R2+3 ending in vein C, prosternum not fused to proepisternum, ovipositor with acanthophorite spines (
<bibRefCitation id="EFA6C7AAFFE3FFF9FCF5F9A8FC77AE40" author="Wood" box="[819,959,1630,1656]" pageId="4" pageNumber="45" refString="Wood, G. C. (1981) Asilidae. In: McAlpine, J. F., Peterson, B. V., Shewell, G. E., Vockeroth, J. R. &amp; Wood, D. M. (Coords.), Manual of Nearctic Diptera, Volume 1, Research Branch, Agriculture Canada, Monographs, 27, 549 - 573." type="book chapter" year="1981">Wood 1981</bibRefCitation>
,
<bibRefCitation id="EFA6C7AAFFE3FFF9FC0DF9A8FAC5AE40" author="Artigas" box="[971,1293,1630,1656]" lastPageId="4" lastPageNumber="44" pageId="4" pageNumber="43" refString="Artigas, J. N. &amp; Papavero, N. (1988) The American genera of Asilidae (Diptera): keys for identification with an atlas of female spermathecae and other morphological details. I. Key to subfamilies and subfamily Leptogastrinae Schiner. Gayana Zoologia, 52 (1 - 2), 95 - 114." type="journal article" year="1988">Artigas and Papavero 1988</bibRefCitation>
). These features are presumed to be plesiomorphic, so the subfamily probably is not monophyletic.
</paragraph>
<paragraph id="8B88BA5BFFE3FFF9FF00F958FEABAFD8" blockId="4.[151,1437,298,2016]" pageId="4" pageNumber="5">
Female
<taxonomicName id="4C37C1D8FFE3FFF9FEE2F958FE60AEFF" ID-CoL="3KXZ" box="[292,424,1710,1735]" class="Insecta" family="Asilidae" genus="Ceraturgus" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE3FFF9FEE2F958FE60AEFF" box="[292,424,1710,1735]" italics="true" pageId="4" pageNumber="5">Ceraturgus</emphasis>
</taxonomicName>
, like many other orthorrhaphous Brachycera (e.g.
<taxonomicName id="4C37C1D8FFE3FFF9FC3AF958FBAFAEF0" ID-CoL="D2N" box="[1020,1127,1710,1736]" class="Insecta" family="Mydidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Mydidae</taxonomicName>
,
<taxonomicName id="4C37C1D8FFE3FFF9FBB5F958FACDAEF0" ID-CoL="6KF" box="[1139,1285,1710,1736]" class="Insecta" family="Apioceridae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Apioceridae</taxonomicName>
,
<taxonomicName id="4C37C1D8FFE3FFF9FAD7F958FA50AEF0" ID-CoL="H6L" box="[1297,1432,1710,1736]" class="Insecta" family="Therevidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Therevidae</taxonomicName>
,
<taxonomicName id="4C37C1D8FFE3FFF9FF51F920FEF0AEC8" ID-CoL="G26" box="[151,312,1750,1776]" class="Insecta" family="Scenopinidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Scenopinidae</taxonomicName>
, and
<taxonomicName id="4C37C1D8FFE3FFF9FEBDF920FDF1AEC8" ID-CoL="623XN" box="[379,569,1750,1776]" class="Insecta" family="Dolichopodidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Dolichopodidae</taxonomicName>
, and some other
<taxonomicName id="4C37C1D8FFE3FFF9FCCAF920FCBAAEC8" ID-CoL="6SY" box="[780,882,1750,1776]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="family">Asilidae</taxonomicName>
), have tergite 10 divided into a pair of spinebearing hemitergites called ancathophorites, used for digging during oviposition (
<bibRefCitation id="EFA6C7AAFFE3FFF9FB88F908FAC4AF20" author="McAlpine" box="[1102,1292,1790,1816]" pageId="4" pageNumber="45" refString="McAlpine, J. F. (1981) Morphology and Terminology - Adults. In: McAlpine, J. F., Peterson, B. V., Shewell, G. E., Vockeroth, J. R. &amp; Wood, D. M. (Coords.), Manual of Nearctic Diptera, Volume 1, Research Branch, Agriculture Canada, Monographs, 27, 2 - 63." type="book chapter" year="1981">McAlpine 1981</bibRefCitation>
). According to
<bibRefCitation id="EFA6C7AAFFE3FFF9FF70F8D0FEB5AF78" author="Papavero" box="[182,381,1830,1856]" pageId="4" pageNumber="45" refString="Papavero, N. (1973) Studies of Asilidae (Diptera) systematics and evolution. I. A preliminary classification in subfamilies. Arquivos de Zoologia, 23 (3), 217 - 274." type="journal article" year="1973">Papavero (1973)</bibRefCitation>
, primitive asilid female terminalia have spined acanthophorites. Reduction of spines and fusion and elongation of the two acanthophorites led to the formation of several
<typeStatus id="548C04F9FFE3FFF9FB8FF8B8FB4FAF50" box="[1097,1159,1870,1896]" pageId="4" pageNumber="5">types</typeStatus>
of ovipositors found in other groups of robber flies. He found spined acanthophorites in only some genera of
<taxonomicName id="4C37C1D8FFE3FFF9FB4CF880FA81AFA8" ID-CoL="89P4B" box="[1162,1353,1910,1936]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Stenopogoninae">Stenopogoninae</taxonomicName>
,
<taxonomicName id="4C37C1D8FFE3FFF9FA90F880FEDCAF80" ID-CoL="87CGK" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Dasypogoninae">Dasypogoninae</taxonomicName>
,
<taxonomicName id="4C37C1D8FFE3FFF9FEE7F868FE62AF80" ID-CoL="89NVV" box="[289,426,1950,1976]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Apocleinae">Apocleinae</taxonomicName>
, and Asilinae. He found them lacking in Laphystiinae,
<taxonomicName id="4C37C1D8FFE3FFF9FB86F868FB0AAF80" ID-CoL="87CJ8" box="[1088,1218,1950,1976]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Laphriinae">Laphriinae</taxonomicName>
,
<taxonomicName id="4C37C1D8FFE3FFF9FB16F868FAACAF80" ID-CoL="89P28" box="[1232,1380,1950,1976]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Ommatiinae">Ommatiinae</taxonomicName>
, and
<taxonomicName id="4C37C1D8FFE3FFF9FF51F830FE97AFD8" ID-CoL="89P58" box="[151,351,1990,2016]" class="Insecta" family="Asilidae" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="subFamily" subFamily="Trigonomiminae">Trigonomiminae</taxonomicName>
.
</paragraph>
<paragraph id="8B88BA5BFFE2FFF8FF00FF6EFB2AA912" blockId="5.[151,1436,152,298]" pageId="5" pageNumber="6">
Ceraturgini have the head wider than high, antennae with 3 flagellomeres (total of 5 antennomeres), third flagellomere pubescent and as wide or wider than first flagellomere (
<bibRefCitation id="EFA6C7AAFFE2FFF8FC0DFF36FAE8A8E2" author="Artigas" box="[971,1312,192,218]" pageId="5" pageNumber="44" refString="Artigas, J. N. &amp; Papavero, N. (1991 a) The American genera of Asilidae (Diptera): keys for identification with an atlas of female spermathecae and other morphological details. VII. 1. Subfamily Stenopogoninae Hull. A preliminary classification into tribes. Gayana Zoologia, 55 (2), 139 - 144." type="journal article" year="1991" yearSuffix="a">Artigas and Papavero 1991a</bibRefCitation>
).
<taxonomicName id="4C37C1D8FFE2FFF8FAF3FF36FF08A939" ID-CoL="3KXZ" class="Insecta" family="Asilidae" genus="Ceraturgus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE2FFF8FAF3FF36FF08A939" italics="true" pageId="5" pageNumber="6">Ceraturgus</emphasis>
</taxonomicName>
lacks the deep clefts at the apices of the first and second flagellomeres that are found in
<taxonomicName id="4C37C1D8FFE2FFF8FB08FF1EFA86A939" ID-CoL="5W8C" box="[1230,1358,232,257]" class="Insecta" family="Asilidae" genus="Myelaphus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE2FFF8FB08FF1EFA86A939" box="[1230,1358,232,257]" italics="true" pageId="5" pageNumber="6">Myelaphus</emphasis>
</taxonomicName>
. The name
<taxonomicName id="4C37C1D8FFE2FFF8FE84FEE6FE0EA911" ID-CoL="3KXZ" box="[322,454,272,297]" class="Insecta" family="Asilidae" genus="Ceraturgus" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B9436649FFE2FFF8FE84FEE6FE0EA911" box="[322,454,272,297]" italics="true" pageId="5" pageNumber="6">Ceraturgus</emphasis>
</taxonomicName>
is derived from the Greek word
<emphasis id="B9436649FFE2FFF8FC83FEE6FC10A911" box="[837,984,272,297]" italics="true" pageId="5" pageNumber="6">Keratourgos</emphasis>
, meaning horn-maker.
</paragraph>
</subSubSection>
</treatment>
</document>