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<mods:title id="B268E3812CB07819DAB7311F26E816B5">To the knowledge of the Near East species of the genus Odocnemis Allard, 1876 (Coleoptera: Tenebrionidae: Helopini)</mods:title>
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<mods:namePart id="DD460BC8D336CEA061799041E5A42AF3">Nabozhenko, Maxim V.</mods:namePart>
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<paragraph id="8B9E9D27FFDDFFDFFF6E46F9577BF84B" blockId="5.[151,625,1937,1999]" box="[151,625,1937,1964]" pageId="5" pageNumber="324">
<heading id="D0D62A4BFFDDFFDFFF6E46F9577BF84B" bold="true" box="[151,625,1937,1964]" fontSize="10" level="2" pageId="5" pageNumber="324" reason="2">
<taxonomicName id="4C21E6A4FFDDFFDFFF6E46F9577BF84B" authority="(Sahlberg, 1908)" baseAuthorityName="Sahlberg" baseAuthorityYear="1908" box="[151,625,1937,1964]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="324" phylum="Arthropoda" rank="species" species="moabitica">
<emphasis id="B9554135FFDDFFDFFF6E46F9577BF84B" bold="true" box="[151,625,1937,1964]" pageId="5" pageNumber="324">
<emphasis id="B9554135FFDDFFDFFF6E46F95497F84C" bold="true" box="[151,413,1937,1964]" italics="true" pageId="5" pageNumber="324">Odocnemis moabitica</emphasis>
(
<bibRefCitation id="EFB0E0D6FFDDFFDFFE5546F95763F84B" author="Sahlberg" box="[428,617,1937,1963]" firstAuthor="Sahlberg" pageId="5" pageNumber="324" pagination="1 - 94" refId="ref9358" refString="Sahlberg, J. (1908) Coleoptera mediterranea et rosso-asiatica nova et minus cognita, maxima ex parte itineribus annis 1895 - 1896, 1898 - 1899 et 1903 - 1904 collecta. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar, 50, 1 - 94. [1907 - 1908]" type="journal article" year="1908">Sahlberg, 1908</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C33BCEACFFDDFFDFFF6E46DD5416F82F" box="[151,284,1973,1999]" pageId="5" pageNumber="324" type="description">
<paragraph id="8B9E9D27FFDDFFDFFF6E46DD5416F82F" blockId="5.[151,625,1937,1999]" box="[151,284,1973,1999]" pageId="5" pageNumber="324">
(
<figureCitation id="131A81A2FFDDFFDFFF6646DD55EEF82F" box="[159,228,1973,1999]" captionStart="FIGURE 3" captionStartId="4.[151,255,1912,1936]" captionTargetBox="[188,1400,184,1883]" captionTargetId="figure@4.[185,1402,180,1885]" captionTargetPageId="4" captionText="FIGURE 3. Odocnemis spp. from the Near East, details of the structure, males: AG—O. praelonga (AC—holotype of O. caudata); H, I—O. clara (holotype); JN—O. moabitica (holotype); OR—O. punctata; SV—O. valga. A, H, Jpronotum; B, I, K—protibia; C, L—mesotibia; D, O, S—aedeagus ventrally; E, P, T—aedeagus laterally; F, Q, U—male inner sternite VIII; J, R, V—spicula gastrale; M—metatibia; N—apical piece of aedeagus, ventrally. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3771525" httpUri="https://zenodo.org/record/3771525/files/figure.png" pageId="5" pageNumber="324">Figs 3</figureCitation>
JN)
</paragraph>
</subSubSection>
<subSubSection id="C33BCEACFFDEFFDCFF6E41F05639FF0E" pageId="6" pageNumber="325" type="discussion">
<paragraph id="8B9E9D27FFDEFFDCFF6E41F05639FF0E" blockId="6.[151,1436,152,238]" pageId="6" pageNumber="325">
<bibRefCitation id="EFB0E0D6FFDEFFDCFF6E41F0545EFF51" author="Sahlberg" box="[151,340,152,177]" firstAuthor="Sahlberg" pageId="6" pageNumber="325" pagination="1 - 94" refId="ref9358" refString="Sahlberg, J. (1908) Coleoptera mediterranea et rosso-asiatica nova et minus cognita, maxima ex parte itineribus annis 1895 - 1896, 1898 - 1899 et 1903 - 1904 collecta. Ofversigt af Finska Vetenskaps-Societetens Forhandlingar, 50, 1 - 94. [1907 - 1908]" type="journal article" year="1908">Sahlberg, 1908: 88</bibRefCitation>
(
<taxonomicName id="4C21E6A4FFDEFFDCFE9B41F154A0FF51" box="[354,426,153,177]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B9554135FFDEFFDCFE9B41F154A0FF51" box="[354,426,153,177]" italics="true" pageId="6" pageNumber="325">Helops</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFB0E0D6FFDEFFDCFE4641F05779FF51" author="Reitter" box="[447,627,152,177]" firstAuthor="Reitter" pageId="6" pageNumber="325" pagination="113 - 171" refId="ref9313" refString="Reitter, E. (1922) Bestimmungstabelle der palaearktischen Helopinae (Col. Tenebrionidae). Wiener Entomologische Zeitung, 39, 1 - 44, 113 - 171. https: // doi. org / 10.5962 / bhl. part. 2572" type="journal article" year="1922">Reitter, 1922: 129</bibRefCitation>
[
<taxonomicName id="4C21E6A4FFDEFFDCFD7841F15690FF51" authority="" box="[641,922,153,177]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="subGenus" subGenus="Odocnemis">
<emphasis id="B9554135FFDEFFDCFD7841F1561BFF51" box="[641,785,153,177]" italics="true" pageId="6" pageNumber="325">Cylindronotus</emphasis>
(
<emphasis id="B9554135FFDEFFDCFCE741F1569AFF51" box="[798,912,153,177]" italics="true" pageId="6" pageNumber="325">Odocnemis</emphasis>
)
</taxonomicName>
];
<bibRefCitation id="EFB0E0D6FFDEFFDCFC5441F0516DFF51" author="Gebien" box="[941,1127,152,177]" firstAuthor="Gebien" pageId="6" pageNumber="325" pagination="399 - 430" refId="ref8399" refString="Gebien, H. (1943) Katalog der Tenebrioniden. Teil. 3. Mitteilungen der Munchner Entomologischen Gesellschaft, 33, 399 - 430." type="journal article" year="1943">Gebien, 1943: 426</bibRefCitation>
[
<taxonomicName id="4C21E6A4FFDEFFDCFB8C41F15084FF51" authority="" box="[1141,1422,153,177]" class="Insecta" family="Tenebrionidae" genus="Cylindronotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="subGenus" subGenus="Odocnemis">
<emphasis id="B9554135FFDEFFDCFB8C41F1500FFF51" box="[1141,1285,153,177]" italics="true" pageId="6" pageNumber="325">Cylindronotus</emphasis>
(
<emphasis id="B9554135FFDEFFDCFAEB41F1508EFF51" box="[1298,1412,153,177]" italics="true" pageId="6" pageNumber="325">Odocnemis</emphasis>
)
</taxonomicName>
];
<bibRefCitation id="EFB0E0D6FFDEFFDCFF3E41DF54BEFF2F" author="Nabozhenko, M. V." box="[199,436,183,208]" pageId="6" pageNumber="325" pagination="36 - 38" refId="ref8634" refString="Nabozhenko, M. V. (2008) New nomenclatural and taxonomic acts, and comments. Tenebrionidae: Helopini. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera. Vol. 5. Tenebrionoidea. Stenstrup, Apollo books, pp. 36 - 38." type="book chapter" year="2008">Nabozhenko, 2008: 37</bibRefCitation>
(
<taxonomicName id="4C21E6A4FFDEFFDCFE3F41DF5732FF2F" authorityName="Allard" authorityYear="1876" box="[454,568,183,207]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B9554135FFDEFFDCFE3F41DF5732FF2F" box="[454,568,183,207]" italics="true" pageId="6" pageNumber="325">Odocnemis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFB0E0D6FFDEFFDCFDAB41DF56A9FF2F" author="Nabozhenko &amp; Lobl" box="[594,931,183,208]" firstAuthor="Nabozhenko" pageId="6" pageNumber="325" pagination="241 - 257" refId="ref8940" refString="Nabozhenko, M. V. &amp; Lobl, I. (2008) Tribe Helopini Latreille, 1802. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palearctic Coleoptera. Vol. 5. Tenebrionoidea. Stenstrup, Apollo books, pp. 241 - 257." type="book chapter" year="2008">Nabozhenko &amp; Löbl, 2008: 244</bibRefCitation>
(
<taxonomicName id="4C21E6A4FFDEFFDCFC4C41DF512DFF2F" authorityName="Allard" authorityYear="1876" box="[949,1063,183,207]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B9554135FFDEFFDCFC4C41DF512DFF2F" box="[949,1063,183,207]" italics="true" pageId="6" pageNumber="325">Odocnemis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFB0E0D6FFDEFFDCFBB941DF5096FF2F" author="Nabozhenko &amp; Keskin" box="[1088,1436,183,208]" firstAuthor="Nabozhenko" pageId="6" pageNumber="325" pagination="255 - 268" refId="ref9237" refString="Nabozhenko, M. V., Keskin, B. &amp; Alpagut Keskin, N. (2016) Taxonomic review of the genus Armenohelops Nabozhenko, 2002 (Coleoptera: Tenebrionidae) with additional support of the mitochondrial COI gene sequences. Caucasian Entomological Bulletin, 12, 255 - 268. https: // doi. org / 10.23885 / 1814 - 3326 - 2016 - 12 - 2 - 255 - 268" type="journal article" year="2016">Nabozhenko &amp; Keskin, 2016: 22</bibRefCitation>
(
<taxonomicName id="4C21E6A4FFDEFFDCFF2941BE5448FF0E" authorityName="Allard" authorityYear="1876" box="[208,322,214,238]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B9554135FFDEFFDCFF2941BE5448FF0E" box="[208,322,214,238]" italics="true" pageId="6" pageNumber="325">Odocnemis</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFB0E0D6FFDEFFDCFEAE41BD5786FF0E" author="Nabozhenko &amp; Tichy" box="[343,652,213,238]" firstAuthor="Nabozhenko" pageId="6" pageNumber="325" pagination="1 - 9" refId="ref9126" refString="Nabozhenko, M. V. &amp; Tichy, V. (2019) Darkling beetles of the tribe Helopini (Coleoptera: Tenebrionidae) of Jordan. Far Eastern Entomologist, 384, 1 - 9. https: // doi. org / 10.25221 / fee. 384.1" type="journal article" year="2019">Nabozhenko &amp; Tichý, 2019: 6</bibRefCitation>
, 7 (
<taxonomicName id="4C21E6A4FFDEFFDCFD4A41BE562FFF0E" authorityName="Allard" authorityYear="1876" box="[691,805,214,238]" class="Insecta" family="Tenebrionidae" genus="Odocnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="325" phylum="Arthropoda" rank="genus">
<emphasis id="B9554135FFDEFFDCFD4A41BE562FFF0E" box="[691,805,214,238]" italics="true" pageId="6" pageNumber="325">Odocnemis</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33BCEACFFDEFFDCFF6E407554AFFE9F" pageId="6" pageNumber="325" type="materials_examined">
<paragraph id="8B9E9D27FFDEFFDCFF6E407554ADFED6" blockId="6.[151,1437,284,1355]" box="[151,423,284,311]" pageId="6" pageNumber="325">
<emphasis id="B9554135FFDEFFDCFF6E4075543CFED7" bold="true" box="[151,310,284,311]" pageId="6" pageNumber="325">
<typeStatus id="549A2385FFDEFFDCFF6E407555D8FED7" box="[151,210,285,311]" pageId="6" pageNumber="325">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F336DDB7FFDEFFDCFEBF40745493FED6" box="[326,409,284,310]" name="Jordan" pageId="6" pageNumber="325">Jordan</collectingCountry>
”.
</paragraph>
<paragraph id="8B9E9D27FFDEFFDCFF3E402954AFFE9F" blockId="6.[151,1437,284,1355]" pageId="6" pageNumber="325">
<emphasis id="B9554135FFDEFFDCFF3E4029547BFEBB" bold="true" box="[199,369,320,347]" pageId="6" pageNumber="325">Type material</emphasis>
was studied, deposited in NHMH. Information about the
<typeStatus id="549A2385FFDEFFDCFBEF40285172FEBA" box="[1046,1144,320,346]" pageId="6" pageNumber="325" type="holotype">holotype</typeStatus>
was published by
<bibRefCitation id="EFB0E0D6FFDEFFDCFAAF402854ABFE9F" author="Nabozhenko &amp; Tichy" firstAuthor="Nabozhenko" pageId="6" pageNumber="325" pagination="1 - 9" refId="ref9126" refString="Nabozhenko, M. V. &amp; Tichy, V. (2019) Darkling beetles of the tribe Helopini (Coleoptera: Tenebrionidae) of Jordan. Far Eastern Entomologist, 384, 1 - 9. https: // doi. org / 10.25221 / fee. 384.1" type="journal article" year="2019">Nabozhenko &amp; Tichý (2019)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33BCEACFFDEFFDCFF3E40E05752FAC7" pageId="6" pageNumber="325" type="description">
<paragraph id="8B9E9D27FFDEFFDCFF3E40E0571DFDB7" blockId="6.[151,1437,284,1355]" pageId="6" pageNumber="325">
<emphasis id="B9554135FFDEFFDCFF3E40E0547FFE43" bold="true" box="[199,373,392,419]" pageId="6" pageNumber="325">Redescription.</emphasis>
Male,
<typeStatus id="549A2385FFDEFFDCFE3C40E05721FE42" box="[453,555,392,418]" pageId="6" pageNumber="325" type="holotype">holotype</typeStatus>
. Body strongly elongate, slender, light-brown, with dull shiny. Head widest at eye level. Eyes large, convex. Ratio of head width at eyes to distance between eyes 1.7. Genae strongly rounded. Lateral margin of head between genae and epistoma weakly emarginated. Anterior margin of epistoma weakly rounded. Fronto-epistomal depression narrow and deep. Punctuation of head moderately coarse, dense (puncture diameter 23 times as long as interpuncture distance). Antennae thin and long, with four apical antennomeres extending beyond base of pronotum.
</paragraph>
<paragraph id="8B9E9D27FFDEFFDCFF3E4309546DFCEA" blockId="6.[151,1437,284,1355]" pageId="6" pageNumber="325">Pronotum elongate, with subequal length and width (1.06 times as wide as long), widest at middle. Lateral margins very weakly rounded, shortly sinuated near middle and base.Anterior margin widely emarginated, base weakly rounded; anterior angles rectangular, weakly projected, posterior ones distinct, right. Disc of pronotum very weakly convex, lateral margins very finely beaded, anterior and base widely beaded. Prothoracic hypomera with very fine, irregular wrinkles.</paragraph>
<paragraph id="8B9E9D27FFDEFFDCFF3E427C5799FC02" blockId="6.[151,1437,284,1355]" pageId="6" pageNumber="325">Elytra strongly elongate (2.24 times as long as wide), widest at middle, 1.35 times as wide and 3.35 as long as pronotum. Lateral margins of elytra evenly weakly rounded, not emarginated near apex and without mucron (but elongated apically), elytral base wider than pronotal base. Striae consist of round, deep, often merged punctures. Elytral intervals weakly convex, very finely and sparsely punctured and covered with distinct granules, which larger at apex and on sides. Epipleura pubescent at apex. Mesoventrite moderately coarsely punctated and finely wrinkled. Metaventrite with fine and sparse punctation.</paragraph>
<paragraph id="8B9E9D27FFDEFFDCFF3E4284576CFBAF" blockId="6.[151,1437,284,1355]" pageId="6" pageNumber="325">Abdominal ventrites with fine, moderately dense punctation; ventrite 1 without pubescence; ventrite 5 completely beaded at apex, surface with longitudinal triangle depression, pubescent with short recumbent setae in middle and suberected long hairs along apex.</paragraph>
<paragraph id="8B9E9D27FFDEFFDCFF3E45315022FAE3" blockId="6.[151,1437,284,1355]" pageId="6" pageNumber="325">Femora smooth on inner side. Protibiae weakly bent, outer side straight; inner side widely weakly bisinuate, with 5 teeth (including apical tooth) and some granules, with dense brush of goldish hairs near apex and some long erected hairs on other surface. Mesotibiae bent, with small teeth and hair brush apically. Metatibiae straight, with dense small granules and some small teeth on inner side, with simple pubescence of recumbent hairs. Protarsi moderately widened, meso- and metatarsi not widened; metatarsomere 1 2 times as long as metatarsomere 2.</paragraph>
<paragraph id="8B9E9D27FFDEFFDCFF3E44645752FAC7" blockId="6.[151,1437,284,1355]" box="[199,600,1292,1319]" pageId="6" pageNumber="325">
Body length 15,2 mm, width
<quantity id="4CD930C2FFDEFFDCFDE94464575EFAC7" box="[528,596,1292,1319]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="325" unit="mm" value="5.0">5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C33BCEACFFDEFFDCFF3E445857DAFAAA" box="[199,720,1328,1355]" pageId="6" pageNumber="325" type="distribution">
<paragraph id="8B9E9D27FFDEFFDCFF3E445857DAFAAA" blockId="6.[151,1437,284,1355]" box="[199,720,1328,1355]" pageId="6" pageNumber="325">
<emphasis id="B9554135FFDEFFDCFF3E4458546BFAAB" bold="true" box="[199,353,1328,1355]" pageId="6" pageNumber="325">Distribution.</emphasis>
<collectingCountry id="F336DDB7FFDEFFDCFE9E445854BFFAAA" box="[359,437,1328,1354]" name="Jordan" pageId="6" pageNumber="325">Jordan</collectingCountry>
, without distinct locality.
</paragraph>
</subSubSection>
</treatment>
</document>