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<document ID-DOI="http://doi.org/10.5281/zenodo.5617005" ID-GBIF-Dataset="440cdbc9-aaf2-4865-8baa-07b1af9b786f" ID-GBIF-Taxon="120609148" ID-Zenodo-Dep="5617005" checkinTime="1462189928338" checkinUser="plazi" docAuthor="Lech Borowiec &amp; Christophe Galkowski" docDate="2016" docId="F76487A93835FFCEFCE8FCD6FB11D5FF" docLanguage="en" docName="borowiec_et_al_2016_ann_zool_redescription_of_tetramorium_atlante.pdf" docOrigin="Annales Zoologici 66 (1)" docStyle="DocumentStyle{}" docTitle="Tetramorium atlante Cagniant 1970, new status" docType="treatment" docVersion="12" lastPageId="7" lastPageNumber="50" masterDocId="0B5DFFD13834FFC9FFA0FFC7FFEFD70D" masterDocTitle="REDESCRIPTION OF TETRAMORIUM ATLANTE CAGNIANT, 1970, NEW STATUS (HYMENOPTERA: FORMICIDAE: MYRMICINAE)" masterLastPageNumber="52" masterPageNumber="43" pageId="1" pageNumber="44" updateTime="1635513642220" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>REDESCRIPTION OF TETRAMORIUM ATLANTE CAGNIANT, 1970, NEW STATUS (HYMENOPTERA: FORMICIDAE: MYRMICINAE)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lech Borowiec</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Christophe Galkowski</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Annales Zoologici</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>66</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>43</mods:start>
<mods:end>52</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3161/00034541ANZ2016.66.1.003</mods:identifier>
<mods:identifier type="GBIF-Dataset">440cdbc9-aaf2-4865-8baa-07b1af9b786f</mods:identifier>
<mods:identifier type="Zenodo-Dep">269540</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5617005" ID-GBIF-Taxon="120609148" ID-Zenodo-Dep="5617005" LSID="urn:lsid:plazi:treatment:F76487A93835FFCEFCE8FCD6FB11D5FF" httpUri="http://treatment.plazi.org/id/F76487A93835FFCEFCE8FCD6FB11D5FF" lastPageId="7" lastPageNumber="50" pageId="1" pageNumber="44">
<subSubSection box="[840,1420,785,808]" pageId="1" pageNumber="44" type="nomenclature">
<paragraph blockId="1.[840,1420,785,839]" box="[840,1420,785,808]" pageId="1" pageNumber="44">
<taxonomicName authority="Cagniant, 1970" authorityName="Cagniant" authorityYear="1970" box="[840,1288,785,808]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="atlante" status="new status">
<emphasis bold="true" box="[840,1109,785,807]" italics="true" pageId="1" pageNumber="44">Tetramorium atlante</emphasis>
<bibRefCitation author="Cagniant, H." box="[1117,1288,786,808]" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de Toulouse" pageId="1" pageNumber="44" pagination="405 - 430" part="105" refString="Cagniant, H. 1970. Deuxieme liste de fourmis d'Algerie recoltees principalement en foret. (1 re partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse (1969), 105: 405 - 430." title="Deuxieme liste de fourmis d'Algerie recoltees principalement en foret. (1 re partie)" type="journal article" year="1970">Cagniant, 1970</bibRefCitation>
</taxonomicName>
<taxonomicNameLabel box="[1295,1420,786,807]" pageId="1" pageNumber="44">new status</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection box="[1065,1194,817,839]" pageId="1" pageNumber="44" type="description">
<paragraph blockId="1.[840,1420,785,839]" box="[1065,1194,817,839]" pageId="1" pageNumber="44">
(
<figureCitation box="[1073,1141,817,839]" captionStart="Figures 1 2" captionStartId="2.[317,385,1882,1899]" captionTargetBox="[317,1269,847,1862]" captionTargetId="figure@2.[317,1270,846,1863]" captionTargetPageId="2" captionText="Figures 1 2. Tetramorium atlante Cagniant, worker: (1) dorsal view; (2) lateral view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/269541/files/figure.png" pageId="1" pageNumber="44">Figs 1</figureCitation>
13)
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="44" type="reference_group">
<paragraph blockId="1.[816,1445,871,940]" pageId="1" pageNumber="44">
<taxonomicName box="[816,1057,872,889]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="caespitum">
<emphasis bold="true" box="[816,1057,872,889]" italics="true" pageId="1" pageNumber="44">Tetramorium caespitum</emphasis>
</taxonomicName>
<taxonomicName authority="Santschi, 1918: 155" authorityName="Santschi" authorityPageNumber="155" authorityYear="1918" pageId="1" pageNumber="44" rank="variety" subSpecies="punicum" variety="atlantis">
st.
<emphasis bold="true" box="[1090,1178,872,889]" italics="true" pageId="1" pageNumber="44">punicum</emphasis>
var.
<emphasis bold="true" box="[1225,1303,871,888]" italics="true" pageId="1" pageNumber="44">atlantis</emphasis>
<bibRefCitation author="Santschi, F." box="[1309,1441,872,889]" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord" pageId="1" pageNumber="44" pagination="153 - 156" part="9" refString="Santschi, F. 1918. Nouveaux Tetramorium africains (Suite et fin). Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord, 9: 153 - 156." title="Nouveaux Tetramorium africains (Suite et fin)" type="journal article" year="1918">Santschi, 1918</bibRefCitation>
: 155
</taxonomicName>
(terra typica: Tunisia), unavailable name.
</paragraph>
<paragraph blockId="1.[816,1445,871,940]" box="[816,1444,923,940]" pageId="1" pageNumber="44">
<taxonomicName box="[816,1052,923,940]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[816,1052,923,940]" italics="true" pageId="1" pageNumber="44">Tetramorium semilaeve</emphasis>
</taxonomicName>
subsp.
<taxonomicName authority="Cagniant, 1970: 430" authorityName="Cagniant" authorityPageNumber="430" authorityYear="1970" box="[1117,1361,923,940]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[1117,1186,923,940]" italics="true" pageId="1" pageNumber="44">atlante</emphasis>
<bibRefCitation author="Cagniant, H." box="[1190,1322,923,940]" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de Toulouse" pageId="1" pageNumber="44" pagination="405 - 430" part="105" refString="Cagniant, H. 1970. Deuxieme liste de fourmis d'Algerie recoltees principalement en foret. (1 re partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse (1969), 105: 405 - 430." title="Deuxieme liste de fourmis d'Algerie recoltees principalement en foret. (1 re partie)" type="journal article" year="1970">Cagniant, 1970</bibRefCitation>
: 430
</taxonomicName>
, 1997: 98.
</paragraph>
</subSubSection>
<subSubSection pageId="1" pageNumber="44" type="materials_examined">
<paragraph blockId="1.[816,1444,980,1902]" pageId="1" pageNumber="44">
<emphasis bold="true" box="[854,1100,980,1002]" pageId="1" pageNumber="44">
<emphasis bold="true" box="[854,1093,980,1002]" italics="true" pageId="1" pageNumber="44">Material examined</emphasis>
.
</emphasis>
Type material:
<materialsCitation ID-GBIF-Occurrence="1265761993" collectionCode="NHMB" collectorName="Dr. F. Santschi" country="Tunisia" location="Kairouan" municipality="Kairouan" pageId="1" pageNumber="44" specimenCode="CASENT0915045" specimenCount="1" typeStatus="type">
syntype work- er on photo (AntWeb resources: Photo by Alexandra Westrich | URL: https://www.antweb.org/specimen/
<specimenCode box="[816,1013,1074,1096]" pageId="1" pageNumber="44">CASENT0915045</specimenCode>
; accessed
<date box="[1133,1277,1074,1096]" pageId="1" pageNumber="44" value="2015-04-01">
<number box="[1133,1146,1074,1095]" pageId="1" pageNumber="44" value="1.0">1</number>
April
<number box="[1219,1277,1074,1096]" pageId="1" pageNumber="44" value="2015.0">2015</number>
</date>
):
<taxonomicName box="[1295,1444,1074,1096]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="species" species="caespitum">T. caespitum</taxonomicName>
L. | st. punicum Sm |
<taxonomicName authority="Sants" authorityName="Sants" box="[1095,1313,1105,1127]" pageId="1" pageNumber="44" rank="variety" variety="atlantis">var. atlantis Sants</taxonomicName>
| Type || Kairouan | (Santschi) ||
<collectingCountry box="[1119,1205,1136,1158]" name="Tunisia" pageId="1" pageNumber="44">Tunisie</collectingCountry>
|
<collectingMunicipality box="[1236,1343,1136,1158]" pageId="1" pageNumber="44">Kairouan</collectingMunicipality>
|
<collectorName pageId="1" pageNumber="44">Dr. F. Santschi</collectorName>
||
<number box="[970,1010,1167,1189]" pageId="1" pageNumber="44" value="135.0">135</number>
|| Sammlung | Dr. F. Santschi | Kairouan ||
<typeStatus box="[983,1040,1199,1220]" pageId="1" pageNumber="44">Type</typeStatus>
|| NATURHIST. | MUSEUM | BASEL || ANTWEB | CASENT | 0
<number box="[1217,1297,1229,1251]" pageId="1" pageNumber="44" value="915045.0">915045</number>
(
<collectionCode box="[1312,1394,1229,1251]" collectionName="Switzerland, Basel, Naturhistorisches Museum" pageId="1" pageNumber="44">NHMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="1.[816,1444,980,1902]" pageId="1" pageNumber="44">
<emphasis bold="true" box="[854,1188,1259,1281]" italics="true" pageId="1" pageNumber="44">Other material examined.</emphasis>
<materialsCitation ID-GBIF-Occurrence="1265761994" collectingDate="2010-07-10" collectionCode="DBET" collectorName="C. Galkowski" country="Morocco" elevation="1600" latitude="33.5409" location="Meknes-Tafilale | Ain Vittel n. Ifrane" longitude="-5.113" pageId="1" pageNumber="44" specimenCode="LBC-MOR00010" specimenCount="19" specimenCount-male="5" specimenCount-queen="4" specimenCount-worker="10">
<specimenCount box="[1200,1292,1260,1281]" pageId="1" pageNumber="44" type="queen">
<number box="[1200,1213,1260,1281]" pageId="1" pageNumber="44" value="4.0">4</number>
gynes
</specimenCount>
,
<specimenCount box="[1308,1402,1260,1282]" pageId="1" pageNumber="44" type="male">
<number box="[1308,1321,1260,1282]" pageId="1" pageNumber="44" value="5.0">5</number>
males
</specimenCount>
,
<specimenCount pageId="1" pageNumber="44" type="worker">
<number box="[1417,1443,1260,1281]" pageId="1" pageNumber="44" value="10.0">10</number>
workers
</specimenCount>
:
<collectingCountry box="[931,1028,1291,1312]" name="Morocco" pageId="1" pageNumber="44">Morocco</collectingCountry>
|
<location LSID="urn:lsid:plazi:treatment:F76487A93835FFCEFCE8FCD6FB11D5FF:7A1260643835FFC8FB8CFACCFC97D24D" country="Morocco" name="Meknes-Tafilale | Ain Vittel n. Ifrane" pageId="1" pageNumber="44">Meknès-Tafilale | Aïn Vittel n. Ifrane</location>
,
<elevation box="[904,990,1322,1344]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="44" unit="m" value="1600.0">
<quantity box="[904,990,1322,1344]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="44" unit="m" value="1600.0">1600 m</quantity>
</elevation>
|
<geoCoordinate box="[1027,1121,1322,1344]" pageId="1" pageNumber="44">
<number box="[1027,1114,1322,1344]" pageId="1" pageNumber="44" value="35.5409">35.5409</number>
°
</geoCoordinate>
/
<geoCoordinate box="[1152,1230,1322,1344]" pageId="1" pageNumber="44">
-
<number box="[1159,1222,1322,1344]" pageId="1" pageNumber="44" value="5.113">5.113</number>
°
</geoCoordinate>
,
<collectingDate box="[1247,1383,1322,1343]" pageId="1" pageNumber="44" value="2010-07-10">
<date box="[1247,1383,1322,1343]" pageId="1" pageNumber="44" value="2010-07-10">
<number box="[1247,1273,1322,1343]" pageId="1" pageNumber="44" value="10.0">10</number>
VII
<number box="[1330,1383,1322,1343]" pageId="1" pageNumber="44" value="2010.0">2010</number>
</date>
</collectingDate>
|
<collectorName pageId="1" pageNumber="44">C. Galkowski</collectorName>
|| Collection L. Borowiec |
<taxonomicName box="[1288,1418,1353,1375]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="44" phylum="Arthropoda" rank="family">Formicidae</taxonomicName>
|
<specimenCode box="[816,996,1384,1405]" pageId="1" pageNumber="44">LBC-MOR00010</specimenCode>
(
<collectionCode box="[1011,1087,1384,1406]" collectionName="DBET" pageId="1" pageNumber="44">DBET</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="6" lastPageNumber="49" pageId="1" pageNumber="44" type="description">
<paragraph blockId="1.[816,1444,980,1902]" lastBlockId="2.[143,772,180,754]" lastPageId="2" lastPageNumber="45" pageId="1" pageNumber="44">
<emphasis bold="true" box="[854,1139,1414,1436]" pageId="1" pageNumber="44">
<emphasis bold="true" box="[854,1029,1414,1436]" italics="true" pageId="1" pageNumber="44">Redescription</emphasis>
. Worker
</emphasis>
(
<figureCitation box="[1154,1254,1415,1437]" captionStart-0="Figures 1 2" captionStart-1="Figures 3 5" captionStartId-0="2.[317,385,1882,1899]" captionStartId-1="3.[143,211,1881,1898]" captionTargetBox-0="[317,1269,847,1862]" captionTargetBox-1="[317,1270,1045,1863]" captionTargetId-0="figure@2.[317,1270,846,1863]" captionTargetId-1="figure@3.[317,1270,1045,1863]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="Figures 1 2. Tetramorium atlante Cagniant, worker: (1) dorsal view; (2) lateral view. Scale bar = 1 mm." captionText-1="Figures 3 5. Tetramorium atlante Cagniant: (3) worker head, scale bar = 0.5 mm; (4) gyne head; (5) gyne mesosoma. Scale bar 4, 5 = 1 mm." httpUri-0="https://zenodo.org/record/269541/files/figure.png" httpUri-1="https://zenodo.org/record/269542/files/figure.png" pageId="1" pageNumber="44">Figs 13</figureCitation>
). Measurements and indices (n=10): CL: 0.77 ± 0.03 (0.7370.844); POC: 0.3 ± 0.02 (0.2680.324); CW: 0.755 ± 0.04 (0.704 0.866); FR: 0.27 ± 0.017 (0.2570.313); FL: 0.271 ± 0.016 (0.2570.313); SL: 0.535 ± 0.02 (0.5060.575); OMD: 0.192 ± 0.015 (0.170.212); EL: 0.146 ± 0.08 (0.1280.291); EH: 0.098 ± 0.01 (0.0780.101); ML: 0.827 ± 0.039 (0.7820.927); SPSP: 0.148 ± 0.013 (0.1120.17); SPL: 0.106 ± 0.011 (0.0890.126); PEL: 0.191 ± 0.014 (0.1790.229); NOL: 0.153 ± 0.011 (0.1340.179); PPL: 0.176 ± 0.007 (0.1620.19); PEH: 0.25 ± 0.013 (0.2350.282); NOH: 0.177 ± 0.016 (0.1650.223); PPH: 0.243 ± 0.011 (0.2290.268); MW: 0.471 ± 0.024 (0.4410.531); PEW: 0.235 ± 0.013 (0.2150.267); PPW: 0.261 ± 0.01 (0.2460.285); CS: 0.763 ± 0.035 (0.7210.855); EYE: 0.16 ± 0.01 (0.138 0.167); CL/CW: 1.021 ± 0.02 (0.9751.047); FR/CS: 0.353 ± 0.009 (0.3410.367); FL/FR: 1.009 ± 0.01 (1.01.043); SL/CS: 0.703 ± 0.016 (0.6720.741); MW/CS: 0.618 ± 0.008 (0.60.632); PEW/PPW: 0.902 ± 0.03 (0.857 0.937); NOH/NOL: 1.163 ± 0.08 (1.0571.336); NOH/ PEL: 0.928 ± 0.05 (0.8521.0); NOL/PEL: 0.801 ± 0.04 (0.7290.872); PEH/NOL: 1.637 ± 0.08 (1.5381.722); PEW/PEH: 0.943 ± 0.03 (0.9140.979); CS/PEW: 3.245 ± 0.07 (3.143.351); CS/PPW: 2.927 ± 0.08 (2.816 3.065); CW/MW: 1.619 ± 0.02 (1.5821.666).
</paragraph>
<paragraph blockId="2.[143,772,180,754]" lastBlockId="3.[143,771,180,969]" lastPageId="3" lastPageNumber="46" pageId="2" pageNumber="45">
Small to medium size, CS 0.763 [0.7210.855]. Whole body brown to dark brown and appendages yellow, the palest specimens never yellow. Head nearly square CL/CW 1.021 [0.9751.047], with almost parallel to slightly rounded sides, straight or slightly concave occipital margin and regularly rounded occipital corners. Eyes small, EYE 0.16 [0.1380.167]. Frons moderately wide, FR/CS 0.353 [0.3410.367], frontal lobes as wide as frons, FL/FR 1.009 [1.01.043]. Scape short, SL/CS 0.703 [0.6720.741], without dorsal carina basally, surface smooth and shiny. Promesonotal dorsum slightly convex, metanotal groove shallow, but distinct. Propodeal teeth short, spiniform, apex of spine located approximately at 2/3 height of mesosoma (
<figureCitation captionStart="Figures 1 2" captionStartId="2.[317,385,1882,1899]" captionTargetBox="[317,1269,847,1862]" captionTargetId="figure@2.[317,1270,846,1863]" captionTargetPageId="2" captionText="Figures 1 2. Tetramorium atlante Cagniant, worker: (1) dorsal view; (2) lateral view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/269541/files/figure.png" pageId="2" pageNumber="45">Fig. 2</figureCitation>
). Dorsal surface of petiole flat, NOH/NOL 1.163 [1.057 1.336], petiole relatively high, PEH/NOL 1.637 [1.5381.722], postpetiole distinctly transverse. General appearance moderately rugose, ground surface finely sculptured. Head dorsum mostly longitudinally rugose and shiny between rugae, rugae extend occipital margin of head, occiput mostly smooth and shiny, sides in anterior half longitudinally rugose and shiny between rugae. In most specimens short band without rugosities runs between frontal rugose area and rugosities along ocular area on each side (
<figureCitation box="[1319,1388,640,662]" captionStart="Figures 3 5" captionStartId="3.[143,211,1881,1898]" captionTargetBox="[317,1270,1045,1863]" captionTargetId="figure@3.[317,1270,1045,1863]" captionTargetPageId="3" captionText="Figures 3 5. Tetramorium atlante Cagniant: (3) worker head, scale bar = 0.5 mm; (4) gyne head; (5) gyne mesosoma. Scale bar 4, 5 = 1 mm." httpUri="https://zenodo.org/record/269542/files/figure.png" pageId="2" pageNumber="45">Fig. 3</figureCitation>
) but smooth area never exceeds 1/7 length of anterior surface of head; in extremely sculptured specimens almost entire frontal surface of head with long rugae with very small smooth patch between interrupted rugae and occiput with fine rugosities. Mesosoma dorsum longitudinally rugose and microreticulate but never reticulate, only occasionally rugae on pronotum partly interrupt- ed with indistinct microreticulation but pronotum nev- er with distinct smooth and shiny areas (
<figureCitation box="[618,684,333,356]" captionStart="Figures 1 2" captionStartId="2.[317,385,1882,1899]" captionTargetBox="[317,1269,847,1862]" captionTargetId="figure@2.[317,1270,846,1863]" captionTargetPageId="2" captionText="Figures 1 2. Tetramorium atlante Cagniant, worker: (1) dorsal view; (2) lateral view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/269541/files/figure.png" pageId="3" pageNumber="46">Fig. 1</figureCitation>
). Sides of pronotum and meso- and metapleuron usually coarsely microreticulate, sometimes reticulation tends to form transverse lines but surface never appears striate or rugose (
<figureCitation box="[310,384,456,478]" captionStart="Figures 1 2" captionStartId="2.[317,385,1882,1899]" captionTargetBox="[317,1269,847,1862]" captionTargetId="figure@2.[317,1270,846,1863]" captionTargetPageId="2" captionText="Figures 1 2. Tetramorium atlante Cagniant, worker: (1) dorsal view; (2) lateral view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/269541/files/figure.png" pageId="3" pageNumber="46">Figs 2</figureCitation>
). Dorsum of petiolar node smooth and shiny with sides carinate, lateral surface microreticulate. Dorsum of postpetiole smooth and shiny, sides microreticulate. First gastral tergite smooth and shiny. Whole dorsum, including head, covered with sparse setae, the longest on pronotum and the shortest on frons. Ventral surface of head with sparse short and 23 moderately long setae not forming a psammophore.
</paragraph>
<caption box="[317,1271,1881,1900]" httpUri="https://zenodo.org/record/269541/files/figure.png" pageId="2" pageNumber="45" targetBox="[317,1269,847,1862]" targetPageId="2">
<paragraph blockId="2.[317,1271,1881,1900]" box="[317,1271,1881,1900]" pageId="2" pageNumber="45">
Figures 12.
<taxonomicName authority="Cagniant" authorityName="Cagniant" box="[432,731,1882,1899]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="45" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[432,641,1882,1899]" italics="true" pageId="2" pageNumber="45">Tetramorium atlante</emphasis>
Cagniant
</taxonomicName>
, worker: (1) dorsal view; (2) lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph blockId="3.[143,771,180,969]" lastBlockId="3.[816,1444,180,969]" pageId="3" pageNumber="46">
<emphasis bold="true" box="[181,245,732,754]" pageId="3" pageNumber="46">Gyne</emphasis>
(
<figureCitation box="[270,385,732,754]" captionStart-0="Figures 3 5" captionStart-1="Figures 6 7" captionStartId-0="3.[143,211,1881,1898]" captionStartId-1="4.[328,396,1883,1900]" captionTargetBox-0="[317,1270,1045,1863]" captionTargetBox-1="[317,1269,687,1866]" captionTargetId-0="figure@3.[317,1270,1045,1863]" captionTargetId-1="figure@4.[317,1270,686,1867]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figures 3 5. Tetramorium atlante Cagniant: (3) worker head, scale bar = 0.5 mm; (4) gyne head; (5) gyne mesosoma. Scale bar 4, 5 = 1 mm." captionText-1="Figures 6 7. Tetramorium atlante Cagniant, gyne: (6) dorsal view; (7) lateral view. Scale bar = 1 mm." httpUri-0="https://zenodo.org/record/269542/files/figure.png" httpUri-1="https://zenodo.org/record/269543/files/figure.png" pageId="3" pageNumber="46">Figs 47</figureCitation>
). Measurements and indicates (n=3): CL: 1.05 ± 0.013 (1.0391.065); POC: 0.388 ± 0.013 (0.374.4); CW: 1.17 ± 0.02 (1.1481.187); FR: 0.409 ± 0.007 (0.40.413); FL: 0.389 ± 0.01 (0.379 0.398); SL: 0.727 ± 0.022 (0.7040.749); OMD: 0.21 ± 0.016 (0.2010.229); EL: 0.287 ± 0.007 (0.2790.291); EH: 0.227 ± 0.02 (0.2120.246); ML: 1.849 ± 0.001 (1.8481.85); SPSP: 0.294 ± 0.02 (0.2680.313); SPL: 0.216 ± 0.006 (0.2120.223); PEL: 0.29 ± 0.02 (0.279 0.313); NOL: 0.214 ± 0.008 (0.2070.223); PPL: 0.315 ± 0.011 (0.3020.324); PEH: 0.458 ± 0.011 (0.4460.469); NOH: 0.287 ± 0.025 (0.2570.302); PPH: 0.436 ± 0.007 (0.4250.446); MW: 1.125 ± 0.01 (1.111.135); PEW: 0.39 ± 0.01 (0.380.4); PPW: 0.488 ± 0.02 (0.4750.508); CS: 1.11 ± 0.014 (1.0941.12); EYE: 0.232 ± 0.012 (0.2240.246); CL/CW: 0.898 ± 0.015 (0.880.907); FR/CS: 0.368 ± 0.01 (0.3570.378); FL/FR: 0.953 ± 0.04 (0.9180.995); SL/CS: 0.655 ± 0.014 (0.6440.671); MW/CS: 1.014 ± 0.09 (0.9951.032); PEW/PPW: 0.801 ± 0.014 (0.7870.815); NOH/NOL: 1.345 ± 0.168 (1.152 1.459); NOH/PEL: 0.989 ± 0.08 (0.9211.082); NOL/PEL: 0.74 ± 0.07 (0.6610.799); PEH/NOL: 2.139 ± 0.03 (2.1032.16); PEW/PEH: 0.853 ± 0.04 (0.830.897); CS/ PEW: 2.845 ± 0.106 (2.7342.946); CS/PPW: 2.278 ± 0.12 (2.1522.357); CW/MW: 0.987 ± 0.02 (0.9671.005); WAIST: 0.792 ± 0.035 (0.7640.83).
</paragraph>
<paragraph blockId="3.[816,1444,180,969]" lastBlockId="5.[143,771,180,846]" lastPageId="5" lastPageNumber="48" pageId="3" pageNumber="46">
Moderate size, CS 1.11 [1.0941.12]. Head and mesosoma dark brown, abdomen brown, appendages yellowish. Head wider than long, CL/CW 0.898 [0.88 0.907], with straight subparallel sides, shallowly emarginate occipital margin and regularly rounded occipital corners. Frons moderately wide, FR/CS 0.368 [0.357 0.378], frontal lobes as wide as frons, FL/FR 0.953 [0.9180.995]. Scape short, SL/CS 0.655 [0.6440.671], without dorsal carina basally, smooth and shiny. Head as wide as scutum, MW/CS 1.014 [0.9951.032]. Propodeal teeth short, triangular. Dorsal crest of petiolar node in frontal view slightly convex. Petiolar node dorsum steeply rounded backward. Petiole and postpetiole relatively narrow, WAIST 0.792 [0.7640.83]. General appearance partly sculptured. Head dorsum, occiput and sides distinctly rugulose, ground surface shiny or indistinctly microreticulate. Frons longitudinally rugulose (
<figureCitation box="[327,397,456,478]" captionStart="Figures 3 5" captionStartId="3.[143,211,1881,1898]" captionTargetBox="[317,1270,1045,1863]" captionTargetId="figure@3.[317,1270,1045,1863]" captionTargetPageId="3" captionText="Figures 3 5. Tetramorium atlante Cagniant: (3) worker head, scale bar = 0.5 mm; (4) gyne head; (5) gyne mesosoma. Scale bar 4, 5 = 1 mm." httpUri="https://zenodo.org/record/269542/files/figure.png" pageId="4" pageNumber="47">Fig. 4</figureCitation>
). Mesosoma flat, pronotal sides visible from above. Anterior margin, sides, and median part of scutum smooth and shiny, lateral to shiny medi- an band punctate and longitudinally striate, striae extending from base to 2/3 length of scutum, in the strongest sculptured specimens median smooth band very narrow and whole base of scutum striate. Scutellum mostly smooth and shiny, only corners or corners and sides with oblique striation and punctate (
<figureCitation box="[1360,1429,272,294]" captionStart="Figures 3 5" captionStartId="3.[143,211,1881,1898]" captionTargetBox="[317,1270,1045,1863]" captionTargetId="figure@3.[317,1270,1045,1863]" captionTargetPageId="3" captionText="Figures 3 5. Tetramorium atlante Cagniant: (3) worker head, scale bar = 0.5 mm; (4) gyne head; (5) gyne mesosoma. Scale bar 4, 5 = 1 mm." httpUri="https://zenodo.org/record/269542/files/figure.png" pageId="4" pageNumber="47">Fig. 5</figureCitation>
). Sides of pronotum ruguloso-reticulate and feebly microreticulate, anepisternum in upper part with fine longitudinal rugae and rest smooth and shiny, katepisternum mostly smooth and shiny but posterior corners and sometimes whole posterior third with fine longitudinal rugae and microreticulation (
<figureCitation box="[1262,1341,456,478]" captionStart="Figures 6 7" captionStartId="4.[328,396,1883,1900]" captionTargetBox="[317,1269,687,1866]" captionTargetId="figure@4.[317,1270,686,1867]" captionTargetPageId="4" captionText="Figures 6 7. Tetramorium atlante Cagniant, gyne: (6) dorsal view; (7) lateral view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/269543/files/figure.png" pageId="4" pageNumber="47">Fig. 7</figureCitation>
). Whole surface of petiolar node distinctly reticulate, posterior surface granulate without or with indistinct transverse rugae. Postpetiole distinctly transverse, sides rounded (
<figureCitation box="[824,892,579,601]" captionStart="Figures 6 7" captionStartId="4.[328,396,1883,1900]" captionTargetBox="[317,1269,687,1866]" captionTargetId="figure@4.[317,1270,686,1867]" captionTargetPageId="4" captionText="Figures 6 7. Tetramorium atlante Cagniant, gyne: (6) dorsal view; (7) lateral view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/269543/files/figure.png" pageId="4" pageNumber="47">Fig. 7</figureCitation>
), dorsum of postpetiole smooth, sides granulate and reticulate. First gastral tergite smooth and shiny. Whole dorsum, including head, covered with short, sparse setae. Ventral surface of head with several short setae, as long as to 1.5 times longer than frontal setae, arising posteriorly to buccal cavity.
</paragraph>
<caption box="[143,1425,1879,1898]" httpUri="https://zenodo.org/record/269542/files/figure.png" pageId="3" pageNumber="46" targetBox="[317,1270,1045,1863]" targetPageId="3">
<paragraph blockId="3.[143,1425,1879,1898]" box="[143,1425,1879,1898]" pageId="3" pageNumber="46">
Figures 35.
<taxonomicName authority="Cagniant" authorityName="Cagniant" box="[259,557,1880,1898]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="46" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[259,468,1880,1898]" italics="true" pageId="3" pageNumber="46">Tetramorium atlante</emphasis>
Cagniant
</taxonomicName>
: (3) worker head, scale bar = 0.5 mm; (4) gyne head; (5) gyne mesosoma. Scale bar 4, 5 = 1 mm.
</paragraph>
</caption>
<caption box="[328,1260,1881,1900]" httpUri="https://zenodo.org/record/269543/files/figure.png" pageId="4" pageNumber="47" targetBox="[317,1269,687,1866]" targetPageId="4">
<paragraph blockId="4.[328,1260,1881,1900]" box="[328,1260,1881,1900]" pageId="4" pageNumber="47">
Figures 67.
<taxonomicName authority="Cagniant" authorityName="Cagniant" box="[443,741,1883,1900]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="atlante">
<taxonomicName box="[443,652,1883,1900]" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[443,652,1883,1900]" italics="true" pageId="4" pageNumber="47">
<taxonomicName box="[443,576,1883,1900]" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="47" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
atlante
</emphasis>
</taxonomicName>
Cagniant
</taxonomicName>
, gyne: (6) dorsal view; (7) lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph blockId="5.[143,771,180,846]" lastBlockId="5.[816,1444,180,846]" pageId="5" pageNumber="48">
<emphasis bold="true" box="[181,241,334,355]" pageId="5" pageNumber="48">Male</emphasis>
(
<figureCitation box="[266,347,333,356]" captionStart="Figures 8 9" captionStartId="5.[327,395,1881,1898]" captionTargetBox="[317,1269,923,1862]" captionTargetId="figure@5.[317,1270,922,1864]" captionTargetPageId="5" captionText="Figures 8 9. Tetramorium atlante Cagniant, male: (8) dorsal view; (9) lateral view. Scale bar = 1 mm." httpUri="https://zenodo.org/record/269544/files/figure.png" pageId="5" pageNumber="48">Figs 8</figureCitation>
13). Measurements and indicates (n=5): CL: 0.694 ± 0.013 (0.6730.715); POC: 0.284 ± 0.01 (0.268.302); CW: 0.836 ± 0.014 (0.810.854); FR: 0.258 ± 0.009 (0.2460.274); FL: 0.292 ± 0.006 (0.285 0.302); SL: 0.311 ± 0.008 (0.3020.324); OMD: 0.076 ± 0.007 (0.0610.084); EL: 0.315 ± 0.008 (0.3020.324); EH: 0.258 ± 0.005 (0.2490.263); ML: 1.81 ± 0.113 (1.6171.935); SPSP: 0.243 ± 0.01 (0.2350.263); SPL: 0.216 ± 0.011 (0.2010.232); PEL: 0.32 ± 0.011 (0.302 0.335); NOL: 0.244 ± 0.019 (0.2120.268); PPL: 0.304 ± 0.018 (0.2790.324); PEH: 0.333 ± 0.01 (0.3130.346); NOH: 0.206 ± 0.015 (0.190.235); PPH: 0.464 ± 0.029 (0.4250.503); MW: 1.108 ± 0.049 (1.0671.2); PEW: 0.36 ± 0.005 (0.3510.366); PPW: 0.518 ± 0.02 (0.492 0.547); CS: 0.765 ± 0.014 (0.7420.785); EYE: 0.374 ± 0.002 (0.3720.379); CL/CW: 0.831 ± 0.004 (0.824 0.837); FR/CS: 0.337 ± 0.01 (0.320.349); FL/FR: 1.134 ± 0.06 (1.0441.228); SL/CS: 0.407 ± 0.009 (0.395 0.421); MW/CS: 1.449 ± 0.06 (1.3891.565); PEW/PPW: 0.696 ± 0.03 (0.6550.744); NOH/NOL: 0.85 ± 0.093 (0.7390.955); NOH/PEL: 0.644 ± 0.05 (0.60.725); NOL/PEL: 0.762 ± 0.044 (0.7020.821); PEH/NOL: 1.372 ± 0.08 (1.251.476); PEW/PEH: 1.084 ± 0.05 (1.014 1.169); CS/PEW: 2.185 ± 0.06 (2.0422.191); CS/PPW: 1.478 ± 0.06 (1.4021.561); CW/MW: 0.691 ± 0.027 (0.6390.719).
</paragraph>
<paragraph blockId="5.[816,1444,180,846]" lastBlockId="6.[143,771,180,877]" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48">
Whole body dark brown, appendages yellowish. Head behind eyes almost trapezoidal, occipital margin slightly convex, occipital corners subangulate. Scutum distinctly wider than head. Propodeum in profile with distinct angulation in position of propodeal teeth. Dorsal crest of petiolar node with obtuse transversal edge, slightly emarginated in frontal view. Head distinctly granulate, dull. Sides of pronotum microreticulate with fine transverse striation. Scutum between sutures in anterior part with oblique striation and microreticulate only along middle runs shiny band, laterally smooth and shiny, rest microreticulate with longitudinal and oblique striation. Scutellum at base and laterally with longitudinal striation, rest smooth and shiny. Anepisternum in anterior half smooth and shiny in posterior half with oblique striation, katepisternum in both dorsal angles distincty striate and microreticulate. Whole surface of petiolar node microgranulate and microreticulate, dull, postpetiole mostly microreticulate only top partly smooth and shiny. Whole surface of propodeum microreticulate, dull. First gastral tergite smooth and shiny. Male genitalia stout (
<figureCitation box="[626,716,425,448]" captionStart="Figures 10 11" captionStartId="6.[143,209,1883,1900]" captionTargetBox="[317,1269,973,1860]" captionTargetId="figure@6.[317,1270,972,1861]" captionTargetPageId="6" captionText="Figures 10 11. Tetramorium atlante Cagniant, male genitalia: (10) dorsal view; (11) ventral view; (12) lateral view; (13) apical view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/269545/files/figure.png" pageId="6" pageNumber="49">Figs 10</figureCitation>
13), in lateral distinctly slightly constricted before apex with obtuse inner angle, top shortly and sparsely pubescent, ventral and dorsal margins of parameres shallowly incised, top of inner margin of paramere before apical denticle straight, without dentiform plate extending beyond the sharp edge of paramere (Fig. 13).
</paragraph>
</subSubSection>
<caption box="[327,1261,1880,1899]" httpUri="https://zenodo.org/record/269544/files/figure.png" pageId="5" pageNumber="48" targetBox="[317,1269,923,1862]" targetPageId="5">
<paragraph blockId="5.[327,1261,1880,1899]" box="[327,1261,1880,1899]" pageId="5" pageNumber="48">
Figures 89.
<taxonomicName authority="Cagniant" authorityName="Cagniant" box="[443,741,1881,1898]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="atlante">
<taxonomicName box="[443,652,1881,1898]" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[443,652,1881,1898]" italics="true" pageId="5" pageNumber="48">
<taxonomicName box="[443,576,1881,1898]" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="48" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
atlante
</emphasis>
</taxonomicName>
Cagniant
</taxonomicName>
, male: (8) dorsal view; (9) lateral view. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49" type="diagnosis">
<paragraph blockId="6.[143,771,180,877]" pageId="6" pageNumber="49">
<emphasis bold="true" box="[181,771,639,662]" pageId="6" pageNumber="49">
<emphasis bold="true" box="[181,463,639,661]" italics="true" pageId="6" pageNumber="49">Differental diagnosis</emphasis>
.
<taxonomicName box="[488,771,640,662]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[488,771,640,662]" italics="true" pageId="6" pageNumber="49">Tetramorium atlante</emphasis>
</taxonomicName>
</emphasis>
and
<taxonomicName box="[200,365,670,692]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[200,365,670,692]" italics="true" pageId="6" pageNumber="49">T. semilaeve</emphasis>
</taxonomicName>
at first glance look similar, and workers are especially difficult to identify. Males and gynes, however, have constant differences in the morphological characters and biometric data (see
<tableCitation box="[670,754,763,785]" captionStart="Table 1" captionStartId="7.[202,252,844,861]" captionTargetBox="[154,1430,896,1896]" captionText="Table 1. Measurements and indices for Tetramorium atlante and T. semilaeve (the most important differences marked with bold)." httpUri="http://table.plazi.org/id/2BB266373833FFCEFF6AFC8BFA9BD450" pageId="6" pageNumber="49" tableUuid="2BB266373833FFCEFF6AFC8BFA9BD450">Table 1</tableCitation>
).
</paragraph>
<paragraph blockId="6.[143,771,180,877]" lastBlockId="6.[816,1444,180,877]" pageId="6" pageNumber="49">
Workers of
<taxonomicName box="[315,438,793,815]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[315,438,793,815]" italics="true" pageId="6" pageNumber="49">T. atlante</emphasis>
</taxonomicName>
differ from workers of
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" italics="true" pageId="6" pageNumber="49">T. semilaeve</emphasis>
</taxonomicName>
(in parentheses characters for
<taxonomicName box="[604,761,824,846]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[604,761,824,846]" italics="true" pageId="6" pageNumber="49">T. semilaeve</emphasis>
</taxonomicName>
) in brown to dark brown body (usually pale yellow to yellowish brown, occasionally brown), head sculpture with longitudinal rugosities, more distinct spread over the frontal surface with very small lateral areas without striation (rugosities less distinct on sides of frontal surface, usually with longitudinal areas without striation). Nevertheless, dark and strongly sculptured workers of
<taxonomicName box="[887,1045,364,386]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[887,1045,364,386]" italics="true" pageId="6" pageNumber="49">T. semilaeve</emphasis>
</taxonomicName>
are extremely similar to
<taxonomicName class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" italics="true" pageId="6" pageNumber="49">T. atlante</emphasis>
</taxonomicName>
and a proper identification requires a nest sample with sexual forms. Workers of
<taxonomicName authority="Forel" authorityName="Forel" box="[1162,1395,425,447]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="depressum">
<emphasis bold="true" box="[1162,1328,425,447]" italics="true" pageId="6" pageNumber="49">T. depressum</emphasis>
Forel
</taxonomicName>
and
<taxonomicName authority="Santschi" authorityName="Santschi" box="[816,1090,456,478]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="punctatum">
<emphasis bold="true" box="[816,984,456,477]" italics="true" pageId="6" pageNumber="49">T. punctatum</emphasis>
Santschi
</taxonomicName>
, two other well-defined species of
<taxonomicName box="[844,999,486,508]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[844,999,486,508]" italics="true" pageId="6" pageNumber="49">T. semilaeve</emphasis>
</taxonomicName>
complex, differ in reduced sculpture of head with at least half frontal surface smooth and shiny.
</paragraph>
<paragraph blockId="6.[816,1444,180,877]" lastBlockId="7.[143,771,180,754]" lastPageId="7" lastPageNumber="50" pageId="6" pageNumber="49">
Gynes of
<taxonomicName box="[964,1089,548,570]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[964,1089,548,570]" italics="true" pageId="6" pageNumber="49">T. atlante</emphasis>
</taxonomicName>
distinctly differ from gynes of
<taxonomicName box="[816,976,578,600]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[816,976,578,600]" italics="true" pageId="6" pageNumber="49">T. semilaeve</emphasis>
</taxonomicName>
(characters for
<taxonomicName box="[1168,1328,578,600]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[1168,1328,578,600]" italics="true" pageId="6" pageNumber="49">T. semilaeve</emphasis>
</taxonomicName>
in parentheses) in larger size with ML 1.8481.850, MW 1.11 1.35, CW 1.1481.187 and SPL 0.2120.223 (1.053 1.813, 1.01.14, 0.9781.161 and 0.1730.201
<emphasis bold="true" box="[1308,1319,670,692]" pageId="6" pageNumber="49">)</emphasis>
, scutum at base and laterally with striation extending from base to at least half length of scutum (scutum completely smooth and shiny or with short striation at base never extending to half length of scutum), scutellum with distinct striation in angles (completely smooth and shiny or with rudiments of striation), anepisternum in posterior half with distinct striation and microreticulation (mostly smooth and shiny), katepisternum along posterior margin with broad area of distinct sculpture (narrow area or reduced to posterior corners), postpetiole less transverse and more sculptured with smooth and shiny area reduced to the middle of top (postpetiole more transverse, less sculpture with whole dorsal surface smooth and shiny).
</paragraph>
<caption box="[143,1444,1881,1900]" httpUri="https://zenodo.org/record/269545/files/figure.png" pageId="6" pageNumber="49" targetBox="[317,1269,973,1860]" targetPageId="6">
<paragraph blockId="6.[143,1444,1881,1900]" box="[143,1444,1881,1900]" pageId="6" pageNumber="49">
Figures 1011.
<taxonomicName authority="Cagniant" authorityName="Cagniant" box="[271,555,1883,1900]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="atlante">
<taxonomicName box="[271,471,1883,1900]" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[271,471,1883,1900]" italics="true" pageId="6" pageNumber="49">
<taxonomicName box="[271,399,1883,1900]" class="Insecta" family="Formicidae" genus="Tetramorium" higherTaxonomySource="CoL" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="49" phylum="Arthropoda" rank="genus">Tetramorium</taxonomicName>
<taxonomicName authority="Cagniant" authorityName="Cagniant" box="[404,555,1883,1900]" pageId="6" pageNumber="49" rank="species" species="atlante">
<taxonomicName box="[404,471,1883,1900]" pageId="6" pageNumber="49" rank="species" species="atlante">atlante</taxonomicName>
Cagniant
</taxonomicName>
</emphasis>
</taxonomicName>
</taxonomicName>
, male genitalia: (10) dorsal view; (11) ventral view; (12) lateral view; (13) apical view. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph blockId="7.[143,771,180,754]" lastBlockId="7.[816,1444,180,754]" pageId="7" pageNumber="50">
Males of
<taxonomicName box="[289,413,425,447]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[289,413,425,447]" italics="true" pageId="7" pageNumber="50">T. atlante</emphasis>
</taxonomicName>
distinctly differ from males of
<taxonomicName box="[143,306,456,478]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[143,306,456,478]" italics="true" pageId="7" pageNumber="50">T. semilaeve</emphasis>
</taxonomicName>
in higher petiole with NOH 0.190.235 and PEH 0.3130.346 (0.1560.179 and 0.2570.313), lower SL/CS ratio 0.3950.421 (0.4360.478), head behind eyes trapezoidal (rounded), scutum between sutures mostly striate and microreticulate (mostly smooth and shiny), scutellum at whole base microreticulate and striate (completely smooth and shiny or in basal corners with fine sculpture), propodeum slightly angulate (round), anepisternum in posterior half with oblique striation (smooth and shiny, at most with few striae close to upper margin), katepisternum in both upper corners with broad microreticulate and striate area (narrow sculptured areas), male genitalia in lateral view distinctly constricted before apex (shallowly constricted) and top of inner margin of paramere without dentiform plate extending beyond the sharp edge of paramere (with dentiform plate).
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="50" type="biology_ecology">
<paragraph blockId="7.[816,1444,180,754]" pageId="7" pageNumber="50">
<emphasis bold="true" box="[854,1063,394,416]" pageId="7" pageNumber="50">
<emphasis bold="true" box="[854,1056,394,416]" italics="true" pageId="7" pageNumber="50">Biological notes</emphasis>
.
</emphasis>
Although locus typicus Kairouan in Tunisia is placed only 70 m a.s.l.
<bibRefCitation author="Cagniant, H." box="[1259,1439,425,447]" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de Toulouse" pageId="7" pageNumber="50" pagination="405 - 430" part="105" refString="Cagniant, H. 1970. Deuxieme liste de fourmis d'Algerie recoltees principalement en foret. (1 re partie). Bulletin de la Societe d'Histoire Naturelle de Toulouse (1969), 105: 405 - 430." title="Deuxieme liste de fourmis d'Algerie recoltees principalement en foret. (1 re partie)" type="journal article" year="1970">Cagniant (1970</bibRefCitation>
,
<bibRefCitation author="Cagniant, H." box="[816,871,456,478]" journalOrPublisher="Annales de la Societe Entomologique de France (n. s.)" pageId="7" pageNumber="50" pagination="89 - 100" part="33" refString="Cagniant, H. 1997. Le genre Tetramorium au Maroc (Hymenoptera: Formicidae): cle et catalogue des especes. Annales de la Societe Entomologique de France (n. s.), 33: 89 - 100." title="Le genre Tetramorium au Maroc (Hymenoptera: Formicidae): cle et catalogue des especes" type="journal article" year="1997">1997</bibRefCitation>
) noted than
<taxonomicName box="[1014,1135,456,478]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[1014,1135,456,478]" italics="true" pageId="7" pageNumber="50">T. atlante</emphasis>
</taxonomicName>
prefers mountainous habitats. Its known localities are mostly between 1600 m (our material) to 2300 m (Haut Atlas). Ants were collected in agricultural habitat such as farmlands, pastures, and in scrubs, luminous forests, roadsides. Preference for mountain habitats additionally differentiates
<taxonomicName box="[870,991,640,662]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="atlante">
<emphasis bold="true" box="[870,991,640,662]" italics="true" pageId="7" pageNumber="50">T. atlante</emphasis>
</taxonomicName>
from
<taxonomicName box="[1055,1211,640,662]" class="Insecta" family="Formicidae" genus="Tetramorium" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="50" phylum="Arthropoda" rank="species" species="semilaeve">
<emphasis bold="true" box="[1055,1211,640,662]" italics="true" pageId="7" pageNumber="50">T. semilaeve</emphasis>
</taxonomicName>
, which, according to
<bibRefCitation author="Borowiec, L. &amp; Galkowski, C. &amp; S. Salata" box="[816,1073,670,693]" journalOrPublisher="ZooKeys" pageId="7" pageNumber="50" pagination="39 - 62" part="512" refString="Borowiec, L., Galkowski, C. and S. Salata. 2015. What is Tetramorium semilaeve Andre, 1883? (Hymenoptera, Formicidae). ZooKeys, 512: 39 - 62." title="What is Tetramorium semilaeve Andre, 1883? (Hymenoptera, Formicidae)" type="journal article" year="2015">
Borowiec
<emphasis bold="true" box="[930,985,670,692]" italics="true" pageId="7" pageNumber="50">et al</emphasis>
. (2015)
</bibRefCitation>
, prefers littoral habitats located usually below 700 m a.s.l. and only occasionally reach to the elevation of just over 1000 m a.s.l.
</paragraph>
</subSubSection>
</treatment>
</document>